Search

Search Results

• 1. [Article] Managing wildlife habitats in forested ecosystems

  Published March 1996. Reprinted August 2006. Please look for up-to-date information in the OSU Extension Catalog: http://extension.oregonstate.edu/catalog

  Citation × Citation Citation Abstract Copy Title: Managing wildlife habitats in forested ecosystems Author: Edge, W. Daniel, Oregon State University. Extension Service Published March 1996. Reprinted August 2006. Please look for up-to-date information in the OSU Extension Catalog: http://extension.oregonstate.edu/catalog Title: Managing wildlife habitats in forested ecosystems Author: Edge, W. Daniel, Oregon State University. Extension Service Published March 1996. Reprinted August 2006. Please look for up-to-date information in the OSU Extension Catalog: http://extension.oregonstate.edu/catalog Close

• 2. [Article] Butterfly and flower community composition among prairie-oak ecosystem habitats in the Willamette Valley, Oregon

  Prairie-oak ecosystems in the Willamette Valley, Oregon have experienced habitat loss and degradation; most of these ecosystems are fragmented into smaller patches. Prairie-oak butterfly species, in the ...

  Citation × Citation Citation Abstract Copy Title: Butterfly and flower community composition among prairie-oak ecosystem habitats in the Willamette Valley, Oregon Author: Powers, Breanna F. Prairie-oak ecosystems in the Willamette Valley, Oregon have experienced habitat loss and degradation; most of these ecosystems are fragmented into smaller patches. Prairie-oak butterfly species, in the Willamette Valley, have decreased dramatically due to loss or degradation of habitat. More research is needed on sustaining the populations of butterflies in prairie-oak ecosystems. The purpose of my study was to examine the role prairie-oak habitats have in structuring butterfly communities in the Willamette Valley. My objectives were to: (1) assess how butterfly community structure and species composition (density, abundance, and species richness) varies along an oak canopy cover gradient in order to understand the role of habitat type in structuring butterfly communities in prairie-oak ecosystems; (2) characterize community composition of nectar resources and environmental variables known to be important for butterfly species. I also investigated temporal and structural relationships between butterfly and flower communities. I found that butterfly abundance and density were greater (p < 0.02) in prairies than in oak savannas or oak woodlands; however, species richness did not differ among habitats (p = 0.54). Ordination of prairie-oak habitats in butterfly species space with a joint plot overlay of environmental variables revealed several strong correlations; butterfly community structure was negatively correlated with litter and oak canopy cover and positively correlated to vegetation cover. The prairie, oak savanna, and oak woodland habitat types differed in nectar species community structure, though this difference was small. Non-native species were some of the most abundant flowers species found in all stands. Butterfly communities were positively related to flower communities. I detected significant seasonal patterns among the habitat types in butterfly and flower communities. Prairies appear to be the most important habitat for native butterfly populations compared to oak savanna and oak woodlands. Flowers were most prevalent in prairies and prairies had the highest percentage of native flower species; the prevalence of non-native flower species was greatest in oak woodlands. My study may represent a short-term community transition as a result of management or it may represent a more permanent community. Further study is needed to understand the effects of different management strategies for habitat restoration and the presence of non-native flower species on butterflies in prairie-oak ecosystems in the Willamette Valley. Title: Butterfly and flower community composition among prairie-oak ecosystem habitats in the Willamette Valley, Oregon Author: Powers, Breanna F. Prairie-oak ecosystems in the Willamette Valley, Oregon have experienced habitat loss and degradation; most of these ecosystems are fragmented into smaller patches. Prairie-oak butterfly species, in the Willamette Valley, have decreased dramatically due to loss or degradation of habitat. More research is needed on sustaining the populations of butterflies in prairie-oak ecosystems. The purpose of my study was to examine the role prairie-oak habitats have in structuring butterfly communities in the Willamette Valley. My objectives were to: (1) assess how butterfly community structure and species composition (density, abundance, and species richness) varies along an oak canopy cover gradient in order to understand the role of habitat type in structuring butterfly communities in prairie-oak ecosystems; (2) characterize community composition of nectar resources and environmental variables known to be important for butterfly species. I also investigated temporal and structural relationships between butterfly and flower communities. I found that butterfly abundance and density were greater (p < 0.02) in prairies than in oak savannas or oak woodlands; however, species richness did not differ among habitats (p = 0.54). Ordination of prairie-oak habitats in butterfly species space with a joint plot overlay of environmental variables revealed several strong correlations; butterfly community structure was negatively correlated with litter and oak canopy cover and positively correlated to vegetation cover. The prairie, oak savanna, and oak woodland habitat types differed in nectar species community structure, though this difference was small. Non-native species were some of the most abundant flowers species found in all stands. Butterfly communities were positively related to flower communities. I detected significant seasonal patterns among the habitat types in butterfly and flower communities. Prairies appear to be the most important habitat for native butterfly populations compared to oak savanna and oak woodlands. Flowers were most prevalent in prairies and prairies had the highest percentage of native flower species; the prevalence of non-native flower species was greatest in oak woodlands. My study may represent a short-term community transition as a result of management or it may represent a more permanent community. Further study is needed to understand the effects of different management strategies for habitat restoration and the presence of non-native flower species on butterflies in prairie-oak ecosystems in the Willamette Valley. Close

• 3. [Article] Independent effects of habitat loss and fragmentation on pollination : tropical forest fragmentation alters hummingbird movements and pollination dynamics

  A growing body of work reveals that animal-mediated pollination is negatively affected by anthropogenic disturbance. Landscape-scale disturbance results in two often inter-related processes: (1) habitat ...

  Citation × Citation Citation Abstract Copy Title: Independent effects of habitat loss and fragmentation on pollination : tropical forest fragmentation alters hummingbird movements and pollination dynamics Author: Hadley, Adam S. A growing body of work reveals that animal-mediated pollination is negatively affected by anthropogenic disturbance. Landscape-scale disturbance results in two often inter-related processes: (1) habitat loss, and (2) disruptions of habitat configuration (i.e. fragmentation). Understanding the relative effects of such processes is critical in designing effective management strategies to limit pollination and pollinator decline. I reviewed existing published work from 1989 to 2009 and found that only six of 303 studies separated the effects of habitat loss from fragmentation. I provide a synthesis of the current landscape, behavioral, and pollination ecology literature in order to present preliminary multiple working hypotheses to explain how these two landscape processes might independently influence pollination dynamics (Chapter 2). Despite the potential importance of independent effects of habitat fragmentation, effects on pollination remain largely untested. Studies designed to disentangle the independent effects of habitat loss and fragmentation are essential for gaining insight into landscape-mediated pollination declines. I also found that the field of landscape pollination ecology could benefit from quantification of the matrix, landscape functional connectivity, and pollinator movement behavior. To test the hypothesis that pollinator movement can be influenced by landscape configuration, I translocated radio-tagged hummingbirds across agricultural and forested landscapes near Las Cruces, Costa Rica (Chapter 3). I found return paths were on average more direct in forested than in agricultural landscapes. In addition, movement paths chosen in agricultural landscapes were more forested than the most direct route suggesting that hummingbirds avoided crossing open areas when possible. To determine if differences in pollinator movement translated to differences in plant reproduction, I tested the relative importance of landscape composition versus configuration on the reproductive success of Heliconia tortuosa, a hummingbird-pollinated forest herb (Chapter 4). I used a stratified random sampling design to select sites across orthogonal gradients in patch size, amount of forest, and elevation. I tested four landscape change hypotheses (i.e., local, landscape composition, landscape fragmentation, and fragmentation threshold). I found that Heliconia reproduction supported both the local site and landscape fragmentation hypotheses. Seed set increased with increasing forest patch size independent of amount of forest in the surrounding landscape. I also found that increasing patch size positively influenced the relative abundance of pollinators. The observed differences in seed set likely resulted from differences in hummingbird movements (Chapter 3) and/or abundance under different landscape configurations. Title: Independent effects of habitat loss and fragmentation on pollination : tropical forest fragmentation alters hummingbird movements and pollination dynamics Author: Hadley, Adam S. A growing body of work reveals that animal-mediated pollination is negatively affected by anthropogenic disturbance. Landscape-scale disturbance results in two often inter-related processes: (1) habitat loss, and (2) disruptions of habitat configuration (i.e. fragmentation). Understanding the relative effects of such processes is critical in designing effective management strategies to limit pollination and pollinator decline. I reviewed existing published work from 1989 to 2009 and found that only six of 303 studies separated the effects of habitat loss from fragmentation. I provide a synthesis of the current landscape, behavioral, and pollination ecology literature in order to present preliminary multiple working hypotheses to explain how these two landscape processes might independently influence pollination dynamics (Chapter 2). Despite the potential importance of independent effects of habitat fragmentation, effects on pollination remain largely untested. Studies designed to disentangle the independent effects of habitat loss and fragmentation are essential for gaining insight into landscape-mediated pollination declines. I also found that the field of landscape pollination ecology could benefit from quantification of the matrix, landscape functional connectivity, and pollinator movement behavior. To test the hypothesis that pollinator movement can be influenced by landscape configuration, I translocated radio-tagged hummingbirds across agricultural and forested landscapes near Las Cruces, Costa Rica (Chapter 3). I found return paths were on average more direct in forested than in agricultural landscapes. In addition, movement paths chosen in agricultural landscapes were more forested than the most direct route suggesting that hummingbirds avoided crossing open areas when possible. To determine if differences in pollinator movement translated to differences in plant reproduction, I tested the relative importance of landscape composition versus configuration on the reproductive success of Heliconia tortuosa, a hummingbird-pollinated forest herb (Chapter 4). I used a stratified random sampling design to select sites across orthogonal gradients in patch size, amount of forest, and elevation. I tested four landscape change hypotheses (i.e., local, landscape composition, landscape fragmentation, and fragmentation threshold). I found that Heliconia reproduction supported both the local site and landscape fragmentation hypotheses. Seed set increased with increasing forest patch size independent of amount of forest in the surrounding landscape. I also found that increasing patch size positively influenced the relative abundance of pollinators. The observed differences in seed set likely resulted from differences in hummingbird movements (Chapter 3) and/or abundance under different landscape configurations. Close

• 4. [Article] Movement and space use by the green hermit (Phaethornis guy) in a fragmented landscape in Costa Rica

  Human activities have altered Earth’s ecosystems. Most biomes have experienced a 20-50% conversion to human use. Loss of habitat has obvious effects on the persistence of species. Fragmentation, however, ...

  Citation × Citation Citation Abstract Copy Title: Movement and space use by the green hermit (Phaethornis guy) in a fragmented landscape in Costa Rica Author: Volpe, Noelia L. Human activities have altered Earth’s ecosystems. Most biomes have experienced a 20-50% conversion to human use. Loss of habitat has obvious effects on the persistence of species. Fragmentation, however, may also negatively affect biodiversity for those species that exhibit behavioral responses to changes in habitat configuration. Such behavioral changes include movement, which is influenced by subdividing and isolating habitats through which animals prefer to travel. Ecosystem services that depend on the ability of animals to move through the landscape could be affected by changes in habitat configuration. Pollination is one such process because access of pollinators to flowers is often the key determinant of plant reproductive success. Yet, relatively little is known about how forest fragmentation may influence the flow of pollen carried by forest-dwelling pollinators, such as hummingbirds. I evaluate how changes in the distribution of pollinators produced by habitat fragmentation in southern Costa Rica affects a pollination web occupied by a generalist tropical pollinator, the green hermit hummingbird (Phaethornis guy). I used radio-telemetry to measure patterns of space use by green hermits living in forested areas with different levels of fragmentation. I first characterize patterns of space use by green hermits at three scales: point, path and home range. I found that green hermits have marked preference for locations in forested areas with high density of Heliconia plants and canopy cover; prefer paths close to streams that minimize crossing large stretches of non-forested matrix; and establish their home ranges in areas with high forest amount. In addition, home range area was negatively related to the amount of forest within it and positively related to fragmentation per se. Average resource density within the home range appeared to be unrelated to the area covered by the green hermits. My observations indicate that while the presence of green hermits in an area is mainly determined by the amount of forest available, the way they move through a landscape will be strongly affected by habitat configuration. In this context, pollen transfer by green hermits will be limited by the presence of open, non-forested areas owing to matrix avoidance. In landscapes with small discontinuous patches, the total area covered by an individual green hermit is likely to be small so I hypothesize that the genetic variability of pollen it transports may be diminished. I also assess the effectiveness of a tool commonly used to study the effect of habitat fragmentation on animal movement: translocation experiments. Such experiments are purported to have important advantages of being time efficient and standardizing across individuals the ‘motivation’ to move. Yet, we lack tests of whether movement behavior of translocated birds reflects natural behavior of unmanipulated birds. I compared the routine movement behavior of green hermits to that of experimentally translocated individuals. Behaviors documented during translocation experiments reflected those observed during routine movements. Both translocated and non-translocated birds showed similar levels of preference for mature tropical forest at the point level and avoided moving across non-forested matrix while selecting streams as movement corridors at the path level. Movement rates (distance covered per unit time) were generally higher during translocation experiments. However, the negative influence of forest cover on movement rates was proportionately similar in translocation and routine movement treatments. My results show that movement behavior of birds during translocation experiments is similar to natural movement behavior of birds. Therefore, translocation experiments may be reliable tools to address effects of landscape structure on animal movement. Title: Movement and space use by the green hermit (Phaethornis guy) in a fragmented landscape in Costa Rica Author: Volpe, Noelia L. Human activities have altered Earth’s ecosystems. Most biomes have experienced a 20-50% conversion to human use. Loss of habitat has obvious effects on the persistence of species. Fragmentation, however, may also negatively affect biodiversity for those species that exhibit behavioral responses to changes in habitat configuration. Such behavioral changes include movement, which is influenced by subdividing and isolating habitats through which animals prefer to travel. Ecosystem services that depend on the ability of animals to move through the landscape could be affected by changes in habitat configuration. Pollination is one such process because access of pollinators to flowers is often the key determinant of plant reproductive success. Yet, relatively little is known about how forest fragmentation may influence the flow of pollen carried by forest-dwelling pollinators, such as hummingbirds. I evaluate how changes in the distribution of pollinators produced by habitat fragmentation in southern Costa Rica affects a pollination web occupied by a generalist tropical pollinator, the green hermit hummingbird (Phaethornis guy). I used radio-telemetry to measure patterns of space use by green hermits living in forested areas with different levels of fragmentation. I first characterize patterns of space use by green hermits at three scales: point, path and home range. I found that green hermits have marked preference for locations in forested areas with high density of Heliconia plants and canopy cover; prefer paths close to streams that minimize crossing large stretches of non-forested matrix; and establish their home ranges in areas with high forest amount. In addition, home range area was negatively related to the amount of forest within it and positively related to fragmentation per se. Average resource density within the home range appeared to be unrelated to the area covered by the green hermits. My observations indicate that while the presence of green hermits in an area is mainly determined by the amount of forest available, the way they move through a landscape will be strongly affected by habitat configuration. In this context, pollen transfer by green hermits will be limited by the presence of open, non-forested areas owing to matrix avoidance. In landscapes with small discontinuous patches, the total area covered by an individual green hermit is likely to be small so I hypothesize that the genetic variability of pollen it transports may be diminished. I also assess the effectiveness of a tool commonly used to study the effect of habitat fragmentation on animal movement: translocation experiments. Such experiments are purported to have important advantages of being time efficient and standardizing across individuals the ‘motivation’ to move. Yet, we lack tests of whether movement behavior of translocated birds reflects natural behavior of unmanipulated birds. I compared the routine movement behavior of green hermits to that of experimentally translocated individuals. Behaviors documented during translocation experiments reflected those observed during routine movements. Both translocated and non-translocated birds showed similar levels of preference for mature tropical forest at the point level and avoided moving across non-forested matrix while selecting streams as movement corridors at the path level. Movement rates (distance covered per unit time) were generally higher during translocation experiments. However, the negative influence of forest cover on movement rates was proportionately similar in translocation and routine movement treatments. My results show that movement behavior of birds during translocation experiments is similar to natural movement behavior of birds. Therefore, translocation experiments may be reliable tools to address effects of landscape structure on animal movement. Close

• 5. [Article] Interactions between ecosystem nitrogen and bedrock control long-term calcium sources in Oregon Coast Range forests

  Ecosystem nitrogen (N) supply strongly influences the availability and cycling of other essential nutrients in temperate forests, especially calcium (Ca). Short-term additions of N that exceed ecosystem ...

  Citation × Citation Citation Abstract Copy Title: Interactions between ecosystem nitrogen and bedrock control long-term calcium sources in Oregon Coast Range forests Author: Hynicka, Justin D. Ecosystem nitrogen (N) supply strongly influences the availability and cycling of other essential nutrients in temperate forests, especially calcium (Ca). Short-term additions of N that exceed ecosystem demands often increase dissolved nitrate fluxes and decrease soil pH, which can stimulate soil Ca loss. However, the long-term effects of high N supply on ecosystem Ca availability are more difficult to determine, and may depend on the Ca content of bedrock and mineral soils. To address this, we examined major and trace element concentrations and ⁸⁷Sr/⁸⁶Sr ratios that trace Ca sources in precipitation, foliage, soil pools, and bedrock at 24 forested sites in the Oregon Coast Range having a wide, natural range of soil N (0.16 - 0.97 % N, 0-10 cm) on contrasting basaltic and sedimentary bedrock. Using a suite of 17 site properties, we also evaluated whether soil N variation across sites was related to the five major state-factors of soil and ecosystem development: climate, organisms, topography, parent material, and time. We found that as soil N increased across sites, its ¹⁵N/¹⁴N ratio declined towards atmospheric values, suggesting that soil N variation reflects a biotic legacy of symbiotic N fixation inputs. In contrast, soil N variation was unrelated to 17 other metrics of soil forming factors that represented climate (mean annual precipitation, mean annual temperature, and distance from the coast), topography (slope, soil depth, and abundance of coarse rock fragments), parent material (within bedrock type bulk and 1 M HNO₃ leachable rock Ca chemistry), and proxies of soil age (Hurst's redness rating, effective cation exchange capacity, Ca in non-exchangeable soil residues, chemical index of alteration, weathering index of Parker, Ca in coarse soil fragments, and soil Ca loss relative to bedrock). These analyses highlight symbiotic N-fixing red alder as a keystone organismal state-factor that produces a wide range of soil N accumulation in coastal Oregon forests. Strontium isotopes (⁸⁷Sr/⁸⁶Sr) and other geochemical analyses indicate that long-term Ca sources in foliage and exchangeable soil pools in Oregon Coast Range forests depend on an interactive effect between N availability and bedrock. Basaltic rocks contained nearly 20-times more Ca than sedimentary rocks across our sites, and this difference was reflected in Sr-isotope partitioning of base cation sources. Atmospheric sources dominated plant and soil pools in forests overlying Ca-poor sedimentary rock, regardless of variation in soil N, indicating extremely limited capacity of weathering to support forest Ca demands. In contrast, forests overlying basaltic rock obtained as much as 80% of Ca from rock weathering in low N sites, yet relied to a greater extent on atmospheric Ca as soil N increased, with less than 10% of Ca from rock weathering at sites with the highest soil N. Surprisingly, differences in fresh rock Ca content and base cation sources between sedimentary and basaltic sites was not reflected in ecosystem Ca availability, and instead increasing soil N caused similar declines in foliar and exchangeable Ca across both rock types. This illustrates that nutrient pool sizes do not necessarily reflect long-term nutrient supply, and highlights how coupled biogeochemical cycles within ecosystems can regulate nutrient loss and supply to biota. Broadly, our results highlight how interactions between biological and geologic factors can influence base cation sources in forest ecosystems. The sustainability of base cation supplies to forests may therefore depend greatly on variation in bedrock weathering at low N sites, yet converge to depend on atmospheric inputs in sites that receive high N loading from biological fixation or anthropogenic deposition. Title: Interactions between ecosystem nitrogen and bedrock control long-term calcium sources in Oregon Coast Range forests Author: Hynicka, Justin D. Ecosystem nitrogen (N) supply strongly influences the availability and cycling of other essential nutrients in temperate forests, especially calcium (Ca). Short-term additions of N that exceed ecosystem demands often increase dissolved nitrate fluxes and decrease soil pH, which can stimulate soil Ca loss. However, the long-term effects of high N supply on ecosystem Ca availability are more difficult to determine, and may depend on the Ca content of bedrock and mineral soils. To address this, we examined major and trace element concentrations and ⁸⁷Sr/⁸⁶Sr ratios that trace Ca sources in precipitation, foliage, soil pools, and bedrock at 24 forested sites in the Oregon Coast Range having a wide, natural range of soil N (0.16 - 0.97 % N, 0-10 cm) on contrasting basaltic and sedimentary bedrock. Using a suite of 17 site properties, we also evaluated whether soil N variation across sites was related to the five major state-factors of soil and ecosystem development: climate, organisms, topography, parent material, and time. We found that as soil N increased across sites, its ¹⁵N/¹⁴N ratio declined towards atmospheric values, suggesting that soil N variation reflects a biotic legacy of symbiotic N fixation inputs. In contrast, soil N variation was unrelated to 17 other metrics of soil forming factors that represented climate (mean annual precipitation, mean annual temperature, and distance from the coast), topography (slope, soil depth, and abundance of coarse rock fragments), parent material (within bedrock type bulk and 1 M HNO₃ leachable rock Ca chemistry), and proxies of soil age (Hurst's redness rating, effective cation exchange capacity, Ca in non-exchangeable soil residues, chemical index of alteration, weathering index of Parker, Ca in coarse soil fragments, and soil Ca loss relative to bedrock). These analyses highlight symbiotic N-fixing red alder as a keystone organismal state-factor that produces a wide range of soil N accumulation in coastal Oregon forests. Strontium isotopes (⁸⁷Sr/⁸⁶Sr) and other geochemical analyses indicate that long-term Ca sources in foliage and exchangeable soil pools in Oregon Coast Range forests depend on an interactive effect between N availability and bedrock. Basaltic rocks contained nearly 20-times more Ca than sedimentary rocks across our sites, and this difference was reflected in Sr-isotope partitioning of base cation sources. Atmospheric sources dominated plant and soil pools in forests overlying Ca-poor sedimentary rock, regardless of variation in soil N, indicating extremely limited capacity of weathering to support forest Ca demands. In contrast, forests overlying basaltic rock obtained as much as 80% of Ca from rock weathering in low N sites, yet relied to a greater extent on atmospheric Ca as soil N increased, with less than 10% of Ca from rock weathering at sites with the highest soil N. Surprisingly, differences in fresh rock Ca content and base cation sources between sedimentary and basaltic sites was not reflected in ecosystem Ca availability, and instead increasing soil N caused similar declines in foliar and exchangeable Ca across both rock types. This illustrates that nutrient pool sizes do not necessarily reflect long-term nutrient supply, and highlights how coupled biogeochemical cycles within ecosystems can regulate nutrient loss and supply to biota. Broadly, our results highlight how interactions between biological and geologic factors can influence base cation sources in forest ecosystems. The sustainability of base cation supplies to forests may therefore depend greatly on variation in bedrock weathering at low N sites, yet converge to depend on atmospheric inputs in sites that receive high N loading from biological fixation or anthropogenic deposition. Close

• 6. [Article] Predicting global change effects on forest biomass and composition in south-central Siberia

  Multiple global changes such as timber harvesting in areas not previously disturbed by cutting and climate change will undoubtedly affect the composition and spatial distribution of boreal forests, which ...

  Citation × Citation Citation Abstract Copy Title: Predicting global change effects on forest biomass and composition in south-central Siberia Author: Gustafson, Eric J., Shvidenko, Anatoly Z., Sturtevant, Brian R., Scheller, Robert M. Year: 2010 Multiple global changes such as timber harvesting in areas not previously disturbed by cutting and climate change will undoubtedly affect the composition and spatial distribution of boreal forests, which will, in turn, affect the ability of these forests to retain carbon and maintain biodiversity. To predict future states of the boreal forest reliably, it is necessary to understand the complex interactions among forest regenerative processes (succession), natural disturbances (e.g., fire, wind, and insects), and anthropogenic disturbances (e.g., timber harvest). We used a landscape succession and disturbance model (LANDIS-II) to study the relative effects of climate change, timber harvesting, and insect outbreaks on forest composition, biomass (carbon), and landscape pattern in south-central Siberia. We found that most response variables were more strongly influenced by timber harvest and insect outbreaks than by the direct effects of climate change. Direct climate effects generally increased tree productivity and modified probability of establishment, but indirect effects on the fire regime generally counteracted the direct effects of climate on forest composition. Harvest and insects significantly changed forest composition, reduced living aboveground biomass, and increased forest fragmentation. We concluded that: (1) Global change is likely to significantly change forest composition of south-central Siberian landscapes, with some changes taking ecosystems outside the historic range of variability. (2) The direct effects of climate change in the study area are not as significant as the exploitation of virgin forest by timber harvest and the potential increased outbreaks of the Siberian silk moth. (3) Novel disturbance by timber harvest and insect outbreaks may greatly reduce the aboveground living biomass of Siberian forests and may significantly alter ecosystem dynamics and wildlife populations by increasing forest fragmentation. Title: Predicting global change effects on forest biomass and composition in south-central Siberia Author: Gustafson, Eric J., Shvidenko, Anatoly Z., Sturtevant, Brian R., Scheller, Robert M. Year: 2010 Multiple global changes such as timber harvesting in areas not previously disturbed by cutting and climate change will undoubtedly affect the composition and spatial distribution of boreal forests, which will, in turn, affect the ability of these forests to retain carbon and maintain biodiversity. To predict future states of the boreal forest reliably, it is necessary to understand the complex interactions among forest regenerative processes (succession), natural disturbances (e.g., fire, wind, and insects), and anthropogenic disturbances (e.g., timber harvest). We used a landscape succession and disturbance model (LANDIS-II) to study the relative effects of climate change, timber harvesting, and insect outbreaks on forest composition, biomass (carbon), and landscape pattern in south-central Siberia. We found that most response variables were more strongly influenced by timber harvest and insect outbreaks than by the direct effects of climate change. Direct climate effects generally increased tree productivity and modified probability of establishment, but indirect effects on the fire regime generally counteracted the direct effects of climate on forest composition. Harvest and insects significantly changed forest composition, reduced living aboveground biomass, and increased forest fragmentation. We concluded that: (1) Global change is likely to significantly change forest composition of south-central Siberian landscapes, with some changes taking ecosystems outside the historic range of variability. (2) The direct effects of climate change in the study area are not as significant as the exploitation of virgin forest by timber harvest and the potential increased outbreaks of the Siberian silk moth. (3) Novel disturbance by timber harvest and insect outbreaks may greatly reduce the aboveground living biomass of Siberian forests and may significantly alter ecosystem dynamics and wildlife populations by increasing forest fragmentation. Close

• 7. [Article] Managing for Ecosystem Services through Governance Networks: An Analysis of Oregon Senate Bill 513

  Presented at The Oregon Water Conference, May 24-25, 2011, Corvallis, OR.

  Citation × Citation Citation Abstract Copy Title: Managing for Ecosystem Services through Governance Networks: An Analysis of Oregon Senate Bill 513 Author: Paulsen, Harmony Presented at The Oregon Water Conference, May 24-25, 2011, Corvallis, OR. Title: Managing for Ecosystem Services through Governance Networks: An Analysis of Oregon Senate Bill 513 Author: Paulsen, Harmony Presented at The Oregon Water Conference, May 24-25, 2011, Corvallis, OR. Close

• 8. [Article] The Rise of the Mesopredator

  Apex predators have experienced catastrophic declines throughout the world as a result of human persecution and habitat loss. These collapses in top predator populations are commonly associated with dramatic ...

  Citation × Citation Citation Abstract Copy Title: The Rise of the Mesopredator Author: Stoner, Chantal J., Brashares, Justin S., Bean, William T., Prugh, Laura R., Epps, Clinton W., Laliberte, Andrea S., Ripple, William J. Apex predators have experienced catastrophic declines throughout the world as a result of human persecution and habitat loss. These collapses in top predator populations are commonly associated with dramatic increases in the abundance of smaller predators. Known as “mesopredator release,” this trophic interaction has been recorded across a range of communities and ecosystems. Mesopredator outbreaks often lead to declining prey populations, sometimes destabilizing communities and driving local extinctions.We present an overview of mesopredator release and illustrate how its underlying concepts can be used to improve predator management in an increasingly fragmented world. We also examine shifts in North American carnivore ranges during the past 200 years and show that 60% of mesopredator ranges have expanded, whereas all apex predator ranges have contracted. The need to understand how best to predict and manage mesopredator release is urgent—mesopredator outbreaks are causing high ecological, economic, and social costs around the world. Title: The Rise of the Mesopredator Author: Stoner, Chantal J., Brashares, Justin S., Bean, William T., Prugh, Laura R., Epps, Clinton W., Laliberte, Andrea S., Ripple, William J. Apex predators have experienced catastrophic declines throughout the world as a result of human persecution and habitat loss. These collapses in top predator populations are commonly associated with dramatic increases in the abundance of smaller predators. Known as “mesopredator release,” this trophic interaction has been recorded across a range of communities and ecosystems. Mesopredator outbreaks often lead to declining prey populations, sometimes destabilizing communities and driving local extinctions.We present an overview of mesopredator release and illustrate how its underlying concepts can be used to improve predator management in an increasingly fragmented world. We also examine shifts in North American carnivore ranges during the past 200 years and show that 60% of mesopredator ranges have expanded, whereas all apex predator ranges have contracted. The need to understand how best to predict and manage mesopredator release is urgent—mesopredator outbreaks are causing high ecological, economic, and social costs around the world. Close

• 9. [Article] Community profiles of ammonia oxidizers across high-elevation forest-to-meadow transects

  In recent years considerable interest has been shown in the diversity of ammonia-oxidizing bacteria in soil communities. The majority of the research has been carried out in Northern Europe where soils ...

  Citation × Citation Citation Abstract Copy Title: Community profiles of ammonia oxidizers across high-elevation forest-to-meadow transects Author: Mintie, Ann In recent years considerable interest has been shown in the diversity of ammonia-oxidizing bacteria in soil communities. The majority of the research has been carried out in Northern Europe where soils have received high atmospheric inputs of nitrogen over the past two centuries. In contrast, although much work has been conducted on nitrogen cycling processes in nitrogen limited forest ecosystems in western North America, no studies have examined the characteristics of ammonia-oxidizing communities in those environments. I was interested in measuring nitrification potential along a high-elevation temperate meadow-to-forest gradient, and characterizing the ammonia-oxidizing communities along that gradient using both molecular and culturing methods. Two experimental sites (Lookout and Carpenter) were chosen in the H.J. Andrews Experimental Forest, located in the western Cascade Range of Oregon, at elevations of approximately 1500 meters. Although nitrification potential rates (NPRs) between sites were not significantly different (P=0.544), variation was observed both within and between sites for specific vegetation types. NPRs were significantly lower in forest (F) soil samples than in meadow (M) soil samples, averaging 5 and 2% of meadow NPRs at Lookout and Carpenter, respectively. In meadow soil samples, most probable number (MPN) population densities of ammonia-oxidizers ranged from 0.6 to 2.6 x 10⁴ cells gram⁻¹ of oven dry soil and 0.9 x 10³ to 1.1 x 10⁵ cells g⁻¹ OD soil at Lookout and Carpenter, respectively. In forest soil samples, population densities ranged from undetectable to 1.1 x 10⁴ cells g⁻¹ OD soil, and 0.9 x 10² to 2.3 x 10³ cells g⁻¹ OD soil at Lookout and Carpenter, respectively. Microbial community DNA was amplified using primers to the ammonia monooxygenase subunit A. Terminal restriction fragments polymorphism analysis with three different restriction enzymes (CfoI, TaqI, and AluI) revealed community profiles dominated by Nitrosospira species. One fragment from CfoI (66 bp) and one fragment from AluI (392-bp) were prominent in 47 soil samples from both sites, and represented between 32 to 100% of the Genescan fragment analyses of PCR products. A full length fragment from AluI digests (491-bp), and three fragments from CfoI (68, 100, and 135- bp) were found sporadically in fewer soil sample T-RFLPs, and within those samples represented smaller percentages of total peak areas. The CfoI 135-bp fragment length was associated primarily with M and meadow/forest (M/F) soils where it was observed in approximately 58 and 100% of the respective transect locations. Eight isolates recovered from soil samples were analyzed using the same molecular methods as the field samples. The T-RFLP patterns of the isolates corresponded with many of those found in the community fingerprints. Four unique amoA sequences were identified among these isolates, including one that possessed the dominant T-RFLP amoA fingerprint in soil samples. This sequence shared 99.8% similarity with Nitrosospira sp. Ka4, a cluster 4 ammonia oxidizer isolated in Norway. Sequence analysis phylogenetically associated the other three isolates (with unique amoA sequences) near Nitrosospira sp. Nsp 1 and Nitrosospira briensis, both cluster 3 ammonia oxidizers. Cloning and sequencing of soil DNA confirmed that ammonia oxidizers with these amoA sequences were present in the soil samples. Two additional amoA sequences were identified in clones that were 95% similar and paraphylogenetically positioned between representatives of clusters 3 and 4. So far, these sequences have not been found in any of the isolates analyzed. Title: Community profiles of ammonia oxidizers across high-elevation forest-to-meadow transects Author: Mintie, Ann In recent years considerable interest has been shown in the diversity of ammonia-oxidizing bacteria in soil communities. The majority of the research has been carried out in Northern Europe where soils have received high atmospheric inputs of nitrogen over the past two centuries. In contrast, although much work has been conducted on nitrogen cycling processes in nitrogen limited forest ecosystems in western North America, no studies have examined the characteristics of ammonia-oxidizing communities in those environments. I was interested in measuring nitrification potential along a high-elevation temperate meadow-to-forest gradient, and characterizing the ammonia-oxidizing communities along that gradient using both molecular and culturing methods. Two experimental sites (Lookout and Carpenter) were chosen in the H.J. Andrews Experimental Forest, located in the western Cascade Range of Oregon, at elevations of approximately 1500 meters. Although nitrification potential rates (NPRs) between sites were not significantly different (P=0.544), variation was observed both within and between sites for specific vegetation types. NPRs were significantly lower in forest (F) soil samples than in meadow (M) soil samples, averaging 5 and 2% of meadow NPRs at Lookout and Carpenter, respectively. In meadow soil samples, most probable number (MPN) population densities of ammonia-oxidizers ranged from 0.6 to 2.6 x 10⁴ cells gram⁻¹ of oven dry soil and 0.9 x 10³ to 1.1 x 10⁵ cells g⁻¹ OD soil at Lookout and Carpenter, respectively. In forest soil samples, population densities ranged from undetectable to 1.1 x 10⁴ cells g⁻¹ OD soil, and 0.9 x 10² to 2.3 x 10³ cells g⁻¹ OD soil at Lookout and Carpenter, respectively. Microbial community DNA was amplified using primers to the ammonia monooxygenase subunit A. Terminal restriction fragments polymorphism analysis with three different restriction enzymes (CfoI, TaqI, and AluI) revealed community profiles dominated by Nitrosospira species. One fragment from CfoI (66 bp) and one fragment from AluI (392-bp) were prominent in 47 soil samples from both sites, and represented between 32 to 100% of the Genescan fragment analyses of PCR products. A full length fragment from AluI digests (491-bp), and three fragments from CfoI (68, 100, and 135- bp) were found sporadically in fewer soil sample T-RFLPs, and within those samples represented smaller percentages of total peak areas. The CfoI 135-bp fragment length was associated primarily with M and meadow/forest (M/F) soils where it was observed in approximately 58 and 100% of the respective transect locations. Eight isolates recovered from soil samples were analyzed using the same molecular methods as the field samples. The T-RFLP patterns of the isolates corresponded with many of those found in the community fingerprints. Four unique amoA sequences were identified among these isolates, including one that possessed the dominant T-RFLP amoA fingerprint in soil samples. This sequence shared 99.8% similarity with Nitrosospira sp. Ka4, a cluster 4 ammonia oxidizer isolated in Norway. Sequence analysis phylogenetically associated the other three isolates (with unique amoA sequences) near Nitrosospira sp. Nsp 1 and Nitrosospira briensis, both cluster 3 ammonia oxidizers. Cloning and sequencing of soil DNA confirmed that ammonia oxidizers with these amoA sequences were present in the soil samples. Two additional amoA sequences were identified in clones that were 95% similar and paraphylogenetically positioned between representatives of clusters 3 and 4. So far, these sequences have not been found in any of the isolates analyzed. Close

• 10. [Article] Human effects on ecological connectivity in aquatic ecosystems: Integrating scientific approaches to support management and mitigation

  This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/science-of-the-t...

  Citation × Citation Citation Abstract Copy Title: Human effects on ecological connectivity in aquatic ecosystems: Integrating scientific approaches to support management and mitigation Author: Riginos, Cynthia, Nagelkerken, Ivan, Finn, Debra S., Hughes, Jane M., Harrod, Chris, Crook, David A., Jennings, Simon, Gillanders, Bronwyn M., Hansen, Michael M., Fry, Brian, Kilada, Raouf W., Erős, Tibor, Allendorf, Frederick W., Hermoso, Virgilio, Page, Timothy J., Hadwen, Wade L., Lowe, Winsor H. This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/science-of-the-total-environment/ Title: Human effects on ecological connectivity in aquatic ecosystems: Integrating scientific approaches to support management and mitigation Author: Riginos, Cynthia, Nagelkerken, Ivan, Finn, Debra S., Hughes, Jane M., Harrod, Chris, Crook, David A., Jennings, Simon, Gillanders, Bronwyn M., Hansen, Michael M., Fry, Brian, Kilada, Raouf W., Erős, Tibor, Allendorf, Frederick W., Hermoso, Virgilio, Page, Timothy J., Hadwen, Wade L., Lowe, Winsor H. This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/science-of-the-total-environment/ Close

• 11. [Article] Range-wide abundance and fluctuating asymmetry patterns of sagebrush-obligate passerine birds

  North American sagebrush (Artemisia spp.) ecosystems are suffering from reductions in habitat extent and quality. Only about 50% of sagebrush remains from pre-settlement conditions, and much of the remaining ...

  Citation × Citation Citation Abstract Copy Title: Range-wide abundance and fluctuating asymmetry patterns of sagebrush-obligate passerine birds Author: Krissman, Cortnie M. North American sagebrush (Artemisia spp.) ecosystems are suffering from reductions in habitat extent and quality. Only about 50% of sagebrush remains from pre-settlement conditions, and much of the remaining habitat is fragmented or degraded by invasive species, fire suppression and overgrazing. Sagebrush-obligate species are experiencing population declines as a result of these ecosystem changes. To effectively conserve this ecosystem, it is essential to understand patterns of abundance and stress of the inhabitants at the landscape-level. Abundance of species across their geographic range is not uniform. Instead, abundance often decreases towards the periphery of the range where resources and habitat conditions become less suitable. In addition, stress in populations closer to the periphery of the range may be expressed in condition-dependent traits where suboptimal environmental conditions occur. Fluctuating asymmetry, random deviations from perfect symmetry in bilateral body parts, may reveal increased stress in these populations. I examined patterns of abundance and fluctuating asymmetry of Brewer’s Sparrow (Spizella breweri), Sage Sparrow (Amphisipiza belli), and Sage Thrasher (Oreoscoptes montanus), in the western United States, to determine how these responses vary with proximity to range periphery of sagebrush. Using negative binomial regression and Akaike’s Information Criterion, I investigated associations between species abundance from North American Breeding Bird Survey count data and several local- and landscapelevel variables derived from digital maps depicting the distribution of sagebrush throughout the United States. Abundance for these three sagebrushobligate passerine birds was greatest in mid-elevation (1,200-2,300 m) areas and increased with sagebrush cover. I found little support for abundance declining as a function of proximity to range periphery. Using mixed model regression, I assessed the relationship between fluctuating asymmetry estimates in the tarsi of juveniles and the proximity of specimens to the periphery of sagebrush distribution. I predicted higher levels of fluctuating asymmetry in individuals nearer the range periphery. However, fluctuating asymmetry decreased with proximity to the range periphery for Sage Sparrow, and, although present Sage Thrasher, fluctuating asymmetry did not differ significantly across the geographic range for this species. Fluctuating asymmetry could not be estimated with confidence for Brewer’s Sparrow. While this study revealed stress in these species, a more logistically complex study to evaluate fluctuating asymmetry patterns across the landscape is necessary for determining areas of conservation priority. Loss of areas of high percent sagebrush cover due to habitat fragmentation and degradation will result in continued declines in abundance of sagebrush-obligate passerine birds. Knowing high abundances of sagebrush-obligate passerine birds occur in locations with high sagebrush cover at mid-elevations will aid land managers and conservation biologists in designing effective conservation strategies for these species. Title: Range-wide abundance and fluctuating asymmetry patterns of sagebrush-obligate passerine birds Author: Krissman, Cortnie M. North American sagebrush (Artemisia spp.) ecosystems are suffering from reductions in habitat extent and quality. Only about 50% of sagebrush remains from pre-settlement conditions, and much of the remaining habitat is fragmented or degraded by invasive species, fire suppression and overgrazing. Sagebrush-obligate species are experiencing population declines as a result of these ecosystem changes. To effectively conserve this ecosystem, it is essential to understand patterns of abundance and stress of the inhabitants at the landscape-level. Abundance of species across their geographic range is not uniform. Instead, abundance often decreases towards the periphery of the range where resources and habitat conditions become less suitable. In addition, stress in populations closer to the periphery of the range may be expressed in condition-dependent traits where suboptimal environmental conditions occur. Fluctuating asymmetry, random deviations from perfect symmetry in bilateral body parts, may reveal increased stress in these populations. I examined patterns of abundance and fluctuating asymmetry of Brewer’s Sparrow (Spizella breweri), Sage Sparrow (Amphisipiza belli), and Sage Thrasher (Oreoscoptes montanus), in the western United States, to determine how these responses vary with proximity to range periphery of sagebrush. Using negative binomial regression and Akaike’s Information Criterion, I investigated associations between species abundance from North American Breeding Bird Survey count data and several local- and landscapelevel variables derived from digital maps depicting the distribution of sagebrush throughout the United States. Abundance for these three sagebrushobligate passerine birds was greatest in mid-elevation (1,200-2,300 m) areas and increased with sagebrush cover. I found little support for abundance declining as a function of proximity to range periphery. Using mixed model regression, I assessed the relationship between fluctuating asymmetry estimates in the tarsi of juveniles and the proximity of specimens to the periphery of sagebrush distribution. I predicted higher levels of fluctuating asymmetry in individuals nearer the range periphery. However, fluctuating asymmetry decreased with proximity to the range periphery for Sage Sparrow, and, although present Sage Thrasher, fluctuating asymmetry did not differ significantly across the geographic range for this species. Fluctuating asymmetry could not be estimated with confidence for Brewer’s Sparrow. While this study revealed stress in these species, a more logistically complex study to evaluate fluctuating asymmetry patterns across the landscape is necessary for determining areas of conservation priority. Loss of areas of high percent sagebrush cover due to habitat fragmentation and degradation will result in continued declines in abundance of sagebrush-obligate passerine birds. Knowing high abundances of sagebrush-obligate passerine birds occur in locations with high sagebrush cover at mid-elevations will aid land managers and conservation biologists in designing effective conservation strategies for these species. Close

• 12. [Article] Soil community dynamics in sagebrush and cheatgrass-invaded ecosystems of the northern Great Basin

  Sagebrush steppe ecosystems in the Great Basin have become increasingly threatened by the proliferation of cheatgrass (Bromus tectorum L.), an invasive annual grass. Diverse sagebrush and perennial bunchgrass ...

  Citation × Citation Citation Abstract Copy Title: Soil community dynamics in sagebrush and cheatgrass-invaded ecosystems of the northern Great Basin Author: DeCrappeo, Nicole M. Sagebrush steppe ecosystems in the Great Basin have become increasingly threatened by the proliferation of cheatgrass (Bromus tectorum L.), an invasive annual grass. Diverse sagebrush and perennial bunchgrass landscapes can be converted to homogenous cheatgrass grasslands mainly through the effects of fire. Although the consequences of this conversion are well understood in the context of plant community dynamics, information on changes to soil communities has not been well documented. I characterized soil surface, microbial, and nematode community dynamics in sagebrush steppe and cheatgrass-invaded areas across the northern Great Basin. I also examined how restoration treatments, such as seeding with a low impact rangeland drill and applying herbicide or sugar to plots, affected soil communities. Soil community functional diversity and structure were alike at sites where soil pH and percent bare ground were similar. Rangeland drill seeding and associated human trampling decreased biological soil crust cover at sites with high proportions of cyanobacteria. Herbicide treatments had little effect on soil communities, but addition of sugar to plots increased carbohydrate utilization and fungal biomass of cheatgrass- invaded soils. In studying paired intact and cheatgrass-invaded sagebrush plots, I found that microbial functional diversity and community composition were different in sagebrush, bunchgrass, cheatgrass, and interspace soils. Fungal biomass and species richness were highest under sagebrush and decreased under cheatgrass. To examine how soil community shifts might affect ecosystem processes, I investigated the contribution of fungi to inorganic nitrogen (N) mineralization in sagebrush and cheatgrass rhizospheres. Results from a ¹⁵N pool dilution experiment modified with the fungal protein synthesis inhibitor cycloheximide showed that gross and net N cycling rates did not differ between control sagebrush and cheatgrass soils and that fungi were important for gross NH₄⁺ production and consumption in both soil types. However, net nitrification increased in sagebrush soils after 24 h, suggesting that when organic matter decomposition by fungi ceased bacteria became carbon limited and could no longer assimilate NH₄⁺. These studies demonstrate that cheatgrass invasion into sagebrush steppe ecosystems can bring about significant changes to soil communities and that these changes may have repercussions for ecosystem functioning in the northern Great Basin. Title: Soil community dynamics in sagebrush and cheatgrass-invaded ecosystems of the northern Great Basin Author: DeCrappeo, Nicole M. Sagebrush steppe ecosystems in the Great Basin have become increasingly threatened by the proliferation of cheatgrass (Bromus tectorum L.), an invasive annual grass. Diverse sagebrush and perennial bunchgrass landscapes can be converted to homogenous cheatgrass grasslands mainly through the effects of fire. Although the consequences of this conversion are well understood in the context of plant community dynamics, information on changes to soil communities has not been well documented. I characterized soil surface, microbial, and nematode community dynamics in sagebrush steppe and cheatgrass-invaded areas across the northern Great Basin. I also examined how restoration treatments, such as seeding with a low impact rangeland drill and applying herbicide or sugar to plots, affected soil communities. Soil community functional diversity and structure were alike at sites where soil pH and percent bare ground were similar. Rangeland drill seeding and associated human trampling decreased biological soil crust cover at sites with high proportions of cyanobacteria. Herbicide treatments had little effect on soil communities, but addition of sugar to plots increased carbohydrate utilization and fungal biomass of cheatgrass- invaded soils. In studying paired intact and cheatgrass-invaded sagebrush plots, I found that microbial functional diversity and community composition were different in sagebrush, bunchgrass, cheatgrass, and interspace soils. Fungal biomass and species richness were highest under sagebrush and decreased under cheatgrass. To examine how soil community shifts might affect ecosystem processes, I investigated the contribution of fungi to inorganic nitrogen (N) mineralization in sagebrush and cheatgrass rhizospheres. Results from a ¹⁵N pool dilution experiment modified with the fungal protein synthesis inhibitor cycloheximide showed that gross and net N cycling rates did not differ between control sagebrush and cheatgrass soils and that fungi were important for gross NH₄⁺ production and consumption in both soil types. However, net nitrification increased in sagebrush soils after 24 h, suggesting that when organic matter decomposition by fungi ceased bacteria became carbon limited and could no longer assimilate NH₄⁺. These studies demonstrate that cheatgrass invasion into sagebrush steppe ecosystems can bring about significant changes to soil communities and that these changes may have repercussions for ecosystem functioning in the northern Great Basin. Close

• 13. [Article] Genesis of some soils in the central western Cascades of Oregon

  Soils representative of several landscape units in the H. J. Andrews Experimental Forest, Western Cascade Range, were sampled, analyzed, and tentatively classified. Genetic inferences were drawn relating ...

  Citation × Citation Citation Abstract Copy Title: Genesis of some soils in the central western Cascades of Oregon Author: Brown, R. B. Soils representative of several landscape units in the H. J. Andrews Experimental Forest, Western Cascade Range, were sampled, analyzed, and tentatively classified. Genetic inferences were drawn relating soils to landscape position and other factors of soil formation. Descriptive information and nutrient capital data were provided to support ecosystem modelling efforts by the Coniferous Forest Biome study group of the U. S./International Biological Program (IBP). To meet the "nutrient capital" requirements of IBP, and to gain insight particularly into the effects of coarse fragments on soil genesis, a volumetric approach was used. Soil organic matter, total N, extractable P, exchangeable cations, free Fe oxides, and cation exchange capacity were expressed in terms of weight or equivalents per unit volume of "whole soil," defined as organic and mineral fine earth components plus pore space plus coarse fragments. The various entities, in grams or equivalents per liter of whole soil, were observed as to their variation with depth. Additional calculations showed levels of the various entities per surface meter ³ of whole soil. Soil temperature data from several sites within the Andrews Forest showed the mesic-frigid soil temperature regime boundary to fall at about the 600 m (2, 000 ft) elevation on south slopes and at about the 450 m (1, 500 ft) elevation on north slopes. The frigidcryic boundary apparently was above the 1, 500 m (4,900 ft) elevation in the Andrews Forest. A sequence of three fluvial and two colluvial soils ranging in elevation from 440 to 460 m was studied in conjunction with concurrent IBP investigations into the geomorphic history of the area The soil on a floodplain adjacent to Lookout Creek, in the sandy-skeletal, mixed, mesic family of Fluventic Hapludolls, was between 500 and 7, 000 yrs in age. The adjacent stream terrace soil, in the loamyskeletal, mixed, mesic family of Fluventic Dystrochrepts, was > 7, 000 yrs old as evidenced Mazama pumice erposi tE-; on or near the surface of the terrace. Volumetric analysis suggested that the floodplain soil had a mollic epipedon largely by virtue of its high content of coarse fragments. The coarse fragments caused a concentration of soil organic matter and recycled cations into a smaller volume of fine earth as compared with the terrace soil, which was lower in coarse fragments. An alluvial-colluvial fan emanated from an adjacent slope and lapped onto the terrace. The soil in this fan was a member of the Fluventic Eutrochrepts, loamy-skeletal, mixed, mesic. It was high in base status and moderately high in clay content, apparently because the southeast-facing source area for parent material here had experienced only shallow weathering and minimal leaching. Across Lookout Creek from these landscape units was a remnant of a high colluvial terrace emanating from a northwest-facing watershed. At the crest of this fan remnant the soil was a member of the loamy-skeletal, mixed, mesic family of Fluventic Dystrochrepts with a distinct layer of Mazama pumice at the 75 to 85 cm depth. This terrace is cut by the watershed stream, which has deposited a comparatively well sorted fan. Soils are in the coarseloamy, mixed, mesic family of Fluventic Dystrochrepts, Eight landscape units in longitudinal and transverse crosssections of upper McRae Creek valley, ranging in elevation from 800 to 1, 200 m, were chosen to study upland soil genesis. Proceeding up the valley, stage of profile development appeared to decrease, indicating a series of depositional events. Soils varied from Eutric Glossoboralfs, fine, mixed on the lowermost surface to Fluventic Dystrochrepts, fine-loamy, mixed, frigid on the next higher surface, to Fluventic Dystrochrepts, loamy-skeletal, mixed, frigid on the next higher surface, to Typic Haplumbrepts, loamy-skeletal, mixed, frigid on the backslope at the valley headwall. The two lowermost soils contrasted markedly with the two uppermost soils, being lower in content of organic matter and N, and higher in base status and clay content. The upper two soils, typical of upper valley bottom and sideslope soils in the region, were extremely low in exchangeable bases and base saturation as measured at pH 7. Compared with the two lower soils, however, these upper soils had relatively high soil: water pH values and relatively small drops in pH from soil:water to soil :KCI measurement. This may be an indication that the upper soils were higher in amorphous content. Greater pH-dependent-CEC would have caused the upper soils to exhibit unrealistically high CEO s--and thus low base saturations--when measured at pH 7. A topoclimosequence of soils on north, east (saddle), and south-facing landscape units with a single parent rock lithology was studied in the transverse valley transect. All three soils were placed tentatively in the Andic Dystrochrepts. The north-facing soil was in a medial - skeletal, frigid family, was the deepest to bedrock ( > 1 1/2 m), aria had the freshest coarse fragments of the three soils. The saddle and south-facing soils were in medial-skeletal, frigid and medial, frigid families, respectively. They were shallow ( <1 m) to saprolite bedrock, with well weathered coarse fragments in the regolith, demonstrating shallower, but apparently more intense weathering on the more exposed sites. These more exposed soils were darker in color than the north-facing soil. Soil organic matter levels were not strikingly different among the three soils. Soil N levels were significantly higher in the south-facing soil than in the east and north-facing soils. Levels of exchangeable bases, while low, were not as low in these three soils as in the upper valley bottom and backslope soils. Saprolite horizons had higher base saturations than overlying horizons. Title: Genesis of some soils in the central western Cascades of Oregon Author: Brown, R. B. Soils representative of several landscape units in the H. J. Andrews Experimental Forest, Western Cascade Range, were sampled, analyzed, and tentatively classified. Genetic inferences were drawn relating soils to landscape position and other factors of soil formation. Descriptive information and nutrient capital data were provided to support ecosystem modelling efforts by the Coniferous Forest Biome study group of the U. S./International Biological Program (IBP). To meet the "nutrient capital" requirements of IBP, and to gain insight particularly into the effects of coarse fragments on soil genesis, a volumetric approach was used. Soil organic matter, total N, extractable P, exchangeable cations, free Fe oxides, and cation exchange capacity were expressed in terms of weight or equivalents per unit volume of "whole soil," defined as organic and mineral fine earth components plus pore space plus coarse fragments. The various entities, in grams or equivalents per liter of whole soil, were observed as to their variation with depth. Additional calculations showed levels of the various entities per surface meter ³ of whole soil. Soil temperature data from several sites within the Andrews Forest showed the mesic-frigid soil temperature regime boundary to fall at about the 600 m (2, 000 ft) elevation on south slopes and at about the 450 m (1, 500 ft) elevation on north slopes. The frigidcryic boundary apparently was above the 1, 500 m (4,900 ft) elevation in the Andrews Forest. A sequence of three fluvial and two colluvial soils ranging in elevation from 440 to 460 m was studied in conjunction with concurrent IBP investigations into the geomorphic history of the area The soil on a floodplain adjacent to Lookout Creek, in the sandy-skeletal, mixed, mesic family of Fluventic Hapludolls, was between 500 and 7, 000 yrs in age. The adjacent stream terrace soil, in the loamyskeletal, mixed, mesic family of Fluventic Dystrochrepts, was > 7, 000 yrs old as evidenced Mazama pumice erposi tE-; on or near the surface of the terrace. Volumetric analysis suggested that the floodplain soil had a mollic epipedon largely by virtue of its high content of coarse fragments. The coarse fragments caused a concentration of soil organic matter and recycled cations into a smaller volume of fine earth as compared with the terrace soil, which was lower in coarse fragments. An alluvial-colluvial fan emanated from an adjacent slope and lapped onto the terrace. The soil in this fan was a member of the Fluventic Eutrochrepts, loamy-skeletal, mixed, mesic. It was high in base status and moderately high in clay content, apparently because the southeast-facing source area for parent material here had experienced only shallow weathering and minimal leaching. Across Lookout Creek from these landscape units was a remnant of a high colluvial terrace emanating from a northwest-facing watershed. At the crest of this fan remnant the soil was a member of the loamy-skeletal, mixed, mesic family of Fluventic Dystrochrepts with a distinct layer of Mazama pumice at the 75 to 85 cm depth. This terrace is cut by the watershed stream, which has deposited a comparatively well sorted fan. Soils are in the coarseloamy, mixed, mesic family of Fluventic Dystrochrepts, Eight landscape units in longitudinal and transverse crosssections of upper McRae Creek valley, ranging in elevation from 800 to 1, 200 m, were chosen to study upland soil genesis. Proceeding up the valley, stage of profile development appeared to decrease, indicating a series of depositional events. Soils varied from Eutric Glossoboralfs, fine, mixed on the lowermost surface to Fluventic Dystrochrepts, fine-loamy, mixed, frigid on the next higher surface, to Fluventic Dystrochrepts, loamy-skeletal, mixed, frigid on the next higher surface, to Typic Haplumbrepts, loamy-skeletal, mixed, frigid on the backslope at the valley headwall. The two lowermost soils contrasted markedly with the two uppermost soils, being lower in content of organic matter and N, and higher in base status and clay content. The upper two soils, typical of upper valley bottom and sideslope soils in the region, were extremely low in exchangeable bases and base saturation as measured at pH 7. Compared with the two lower soils, however, these upper soils had relatively high soil: water pH values and relatively small drops in pH from soil:water to soil :KCI measurement. This may be an indication that the upper soils were higher in amorphous content. Greater pH-dependent-CEC would have caused the upper soils to exhibit unrealistically high CEO s--and thus low base saturations--when measured at pH 7. A topoclimosequence of soils on north, east (saddle), and south-facing landscape units with a single parent rock lithology was studied in the transverse valley transect. All three soils were placed tentatively in the Andic Dystrochrepts. The north-facing soil was in a medial - skeletal, frigid family, was the deepest to bedrock ( > 1 1/2 m), aria had the freshest coarse fragments of the three soils. The saddle and south-facing soils were in medial-skeletal, frigid and medial, frigid families, respectively. They were shallow ( <1 m) to saprolite bedrock, with well weathered coarse fragments in the regolith, demonstrating shallower, but apparently more intense weathering on the more exposed sites. These more exposed soils were darker in color than the north-facing soil. Soil organic matter levels were not strikingly different among the three soils. Soil N levels were significantly higher in the south-facing soil than in the east and north-facing soils. Levels of exchangeable bases, while low, were not as low in these three soils as in the upper valley bottom and backslope soils. Saprolite horizons had higher base saturations than overlying horizons. Close

• 14. [Article] Beyond smoke and mirrors: a synthesis of fire policy and science

  Fire performs many beneficial ecosystem functions in dry forests and rangelands across much of North America. In the last century, however, the role of fire has been dramatically altered by numerous anthropogenic ...

  Citation × Citation Citation Abstract Copy Title: Beyond smoke and mirrors: a synthesis of fire policy and science Author: Deacon Williams, Cindy, Williams, Jack E., Franklin, Jerry F., Dellasala, Dominick A. Fire performs many beneficial ecosystem functions in dry forests and rangelands across much of North America. In the last century, however, the role of fire has been dramatically altered by numerous anthropogenic factors acting as root causes of the current fire crisis, including widespread logging, road building, fire suppression, habitat fragmentation, urban development, livestock grazing, and, more recently, climate change. The intensity and extent of fires in the western United States, specifically, have dramatically increased over the past several decades. Such shifts in fire behavior have triggered sweeping policy changes that were intended to prevent or contain fires but that pose significant risks to the integrity of ecosystems and the role fire historically played in shaping them. Here, we provide a social and ecological context for summarizing this special issue on fires, including general guidelines and principles for managers concerned about balancing the risks of inaction against the risks of action over extensive areas. Fundamental to our understanding of fire is the notion that it is extremely variable, has multiple causes, and requires ecological solutions that are sensitive to spatial scale and context. Therefore, forest managers must recognize that different forest types have different fire regimes and require fundamentally different fire- management policies. Furthermore, to restore or maintain ecological integrity, including the role of fire, treatments need to be tailored to site-specific conditions with an adaptive approach. We provide a conceptual framework for prioritizing fuel treatments and restoration activities in the wildlands-urban intermix versus those in wildland areas farther from human settlement. In general, the science of conservation biology has much to offer in helping to shape wildfire policy direction; however, conservation biologists must become more engaged to better ensure that policy decisions are based on sound science and that ecological risks are incorporated. Title: Beyond smoke and mirrors: a synthesis of fire policy and science Author: Deacon Williams, Cindy, Williams, Jack E., Franklin, Jerry F., Dellasala, Dominick A. Fire performs many beneficial ecosystem functions in dry forests and rangelands across much of North America. In the last century, however, the role of fire has been dramatically altered by numerous anthropogenic factors acting as root causes of the current fire crisis, including widespread logging, road building, fire suppression, habitat fragmentation, urban development, livestock grazing, and, more recently, climate change. The intensity and extent of fires in the western United States, specifically, have dramatically increased over the past several decades. Such shifts in fire behavior have triggered sweeping policy changes that were intended to prevent or contain fires but that pose significant risks to the integrity of ecosystems and the role fire historically played in shaping them. Here, we provide a social and ecological context for summarizing this special issue on fires, including general guidelines and principles for managers concerned about balancing the risks of inaction against the risks of action over extensive areas. Fundamental to our understanding of fire is the notion that it is extremely variable, has multiple causes, and requires ecological solutions that are sensitive to spatial scale and context. Therefore, forest managers must recognize that different forest types have different fire regimes and require fundamentally different fire- management policies. Furthermore, to restore or maintain ecological integrity, including the role of fire, treatments need to be tailored to site-specific conditions with an adaptive approach. We provide a conceptual framework for prioritizing fuel treatments and restoration activities in the wildlands-urban intermix versus those in wildland areas farther from human settlement. In general, the science of conservation biology has much to offer in helping to shape wildfire policy direction; however, conservation biologists must become more engaged to better ensure that policy decisions are based on sound science and that ecological risks are incorporated. Close

• 15. [Article] Towards a cohesive, holistic view of top predation: a definition, synthesis and perspective

  This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by the Authors and the Nordic Society Oikos. It is published by John Wiley & Sons, ...

  Citation × Citation Citation Abstract Copy Title: Towards a cohesive, holistic view of top predation: a definition, synthesis and perspective Author: Schmitz, Oswald J., et al., Krebs, Charles J., Ripple, William J., Sergio, Fabrizio This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by the Authors and the Nordic Society Oikos. It is published by John Wiley & Sons, Inc., and can be found at: http://onlinelibrary.wiley.com/journal/10.1111/%28ISSN%291600-0706 Title: Towards a cohesive, holistic view of top predation: a definition, synthesis and perspective Author: Schmitz, Oswald J., et al., Krebs, Charles J., Ripple, William J., Sergio, Fabrizio This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by the Authors and the Nordic Society Oikos. It is published by John Wiley & Sons, Inc., and can be found at: http://onlinelibrary.wiley.com/journal/10.1111/%28ISSN%291600-0706 Close

• 16. [Article] The influence of herbivore generated inputs on nutrient cycling and soil processes in a lower montane tropical rain forest of Puerto Rico

  The role of insect herbivores in the nutrient cycling dynamics of forest ecosystems remains poorly understood. Although past research in herbivory has focused primarily on the deleterious effects that ...

  Citation × Citation Citation Abstract Copy Title: The influence of herbivore generated inputs on nutrient cycling and soil processes in a lower montane tropical rain forest of Puerto Rico Author: Fonte, Steven J. The role of insect herbivores in the nutrient cycling dynamics of forest ecosystems remains poorly understood. Although past research in herbivory has focused primarily on the deleterious effects that insects can have on tree growth and mortality, the overall effects of herbivory are more complex. Herbivores have the potential to alter key ecosystem processes in a number of ways. One is by altering the flow of nutrients from the canopy to the forest floor. The studies presented here examine the effects that frass (insect excreta), greenfall (green leaf fragments) and herbivore modified throughfall, have on decomposition processes and soil nutrient dynamics. Both studies were carried out in a lower montane tropical rain forest near the El Verde Field Station, Luquillo Long-Term Ecological Research (LTER) site, Puerto Rico. The first study presented here tested the hypotheses that green leaves are of higher quality and decompose more rapidly than senesced leaves. Green and senesced leaves of four common native tree species (Dacryodes excelsa, Manilkara bidentata, Guarea guidonia and Cecropia schreberiana), were collected and analyzed for C, N, and complex C compounds. Litterbags containing green and senescent leaves of each species were placed in the field for up to 16 weeks in order to compare rates of litter decomposition. Green leaves contained significantly higher (p < 0.05) concentrations of N and lower lignin:N ratios than senescent leaves for all four species. Decomposition rates were significantly higher (p < 0.05) for green leaves compared to senescent leaves for all four species. These results demonstrate that insect herbivores may influence key ecosystem processes via the production of greenfall. The relevance of this study extends to green leaf deposition resulting from multiple sources, including high wind and rain events, as well as disturbance by larger canopy organisms. The second study tested a direct link between herbivory and soil processes. By altering levels of herbivory on a common understory plant (Piper glabrescens), using a prevalent folivore (Lamponius portoricensis), this study tested for effects of herbivory on nutrient inputs to the soil, as well as rates of litter decomposition. Enclosures were constructed around P. glabrescens individuals in the field and assigned to one of four treatments: herbivore exclusion, control, low herbivory and high herbivory. Total leaf area loss and greenfall production were measured for each plant using a sub-sample of randomly chosen leaves. Litterbags and ion exchange resin bags, placed under each plant, recorded rates of litter decomposition and the flow of NO3, NH4 and PO4 to the forest floor during this 76-day experiment. The treatments were effective in establishing a wide range of herbivory. Both the total leaf area removed and greenfall deposition demonstrated significant positive correlations (p <0.05) with NO3 transfer to the forest floor, but not with NH4 or PO4. Although decomposition rates showed no significant correlation with total leaf area losses or the greenfall component, a significant correlation was found between decay rates and frass related inputs (defined as the total leaf area removed minus the portion removed as greenfall). This study clearly demonstrates the ability of insect herbivores to alter nutrient cycling in tropical forest ecosystems and is the first study to demonstrate a direct link between herbivory and decomposition processes. These experiments provide clear evidence that insect herbivores can alter nutrient cycling and soil processes in forest systems. Such findings elucidate the need to more fully consider the effects of herbivores in both ecosystem models and management issues of tropical forest ecosystems. Title: The influence of herbivore generated inputs on nutrient cycling and soil processes in a lower montane tropical rain forest of Puerto Rico Author: Fonte, Steven J. The role of insect herbivores in the nutrient cycling dynamics of forest ecosystems remains poorly understood. Although past research in herbivory has focused primarily on the deleterious effects that insects can have on tree growth and mortality, the overall effects of herbivory are more complex. Herbivores have the potential to alter key ecosystem processes in a number of ways. One is by altering the flow of nutrients from the canopy to the forest floor. The studies presented here examine the effects that frass (insect excreta), greenfall (green leaf fragments) and herbivore modified throughfall, have on decomposition processes and soil nutrient dynamics. Both studies were carried out in a lower montane tropical rain forest near the El Verde Field Station, Luquillo Long-Term Ecological Research (LTER) site, Puerto Rico. The first study presented here tested the hypotheses that green leaves are of higher quality and decompose more rapidly than senesced leaves. Green and senesced leaves of four common native tree species (Dacryodes excelsa, Manilkara bidentata, Guarea guidonia and Cecropia schreberiana), were collected and analyzed for C, N, and complex C compounds. Litterbags containing green and senescent leaves of each species were placed in the field for up to 16 weeks in order to compare rates of litter decomposition. Green leaves contained significantly higher (p < 0.05) concentrations of N and lower lignin:N ratios than senescent leaves for all four species. Decomposition rates were significantly higher (p < 0.05) for green leaves compared to senescent leaves for all four species. These results demonstrate that insect herbivores may influence key ecosystem processes via the production of greenfall. The relevance of this study extends to green leaf deposition resulting from multiple sources, including high wind and rain events, as well as disturbance by larger canopy organisms. The second study tested a direct link between herbivory and soil processes. By altering levels of herbivory on a common understory plant (Piper glabrescens), using a prevalent folivore (Lamponius portoricensis), this study tested for effects of herbivory on nutrient inputs to the soil, as well as rates of litter decomposition. Enclosures were constructed around P. glabrescens individuals in the field and assigned to one of four treatments: herbivore exclusion, control, low herbivory and high herbivory. Total leaf area loss and greenfall production were measured for each plant using a sub-sample of randomly chosen leaves. Litterbags and ion exchange resin bags, placed under each plant, recorded rates of litter decomposition and the flow of NO3, NH4 and PO4 to the forest floor during this 76-day experiment. The treatments were effective in establishing a wide range of herbivory. Both the total leaf area removed and greenfall deposition demonstrated significant positive correlations (p <0.05) with NO3 transfer to the forest floor, but not with NH4 or PO4. Although decomposition rates showed no significant correlation with total leaf area losses or the greenfall component, a significant correlation was found between decay rates and frass related inputs (defined as the total leaf area removed minus the portion removed as greenfall). This study clearly demonstrates the ability of insect herbivores to alter nutrient cycling in tropical forest ecosystems and is the first study to demonstrate a direct link between herbivory and decomposition processes. These experiments provide clear evidence that insect herbivores can alter nutrient cycling and soil processes in forest systems. Such findings elucidate the need to more fully consider the effects of herbivores in both ecosystem models and management issues of tropical forest ecosystems. Close

• 17. [Article] Green Roofs and Urban Biodiversity: Their Role as Invertebrate Habitat and the Effect of Design on Beetle Community

  With over half the world's population now living in cities, urban areas represent one of earth's few ecosystems that are increasing in extent, and are sites of altered biogeochemical cycles, habitat fragmentation, ...

  Citation × Citation Citation Abstract Copy Title: Green Roofs and Urban Biodiversity: Their Role as Invertebrate Habitat and the Effect of Design on Beetle Community Author: Gonsalves, Sydney Marie Year: 2016 With over half the world's population now living in cities, urban areas represent one of earth's few ecosystems that are increasing in extent, and are sites of altered biogeochemical cycles, habitat fragmentation, and changes in biodiversity. However, urban green spaces, including green roofs, can also provide important pools of biodiversity and contribute to regional gamma diversity, while novel species assemblages can enhance some ecosystem services. Green roofs may also mitigate species loss in urban areas and have been shown to support a surprising diversity of invertebrates, including rare and endangered species. In the first part of this study I reviewed the literature on urban invertebrate communities and diversity to better understand the role of green roofs in providing habitat in the context of the larger urban mosaic. My review concluded that, while other factors such as surrounding land use and connectivity are also important to specific invertebrate taxa, local habitat variables contribute substantially to the structure and diversity of urban invertebrate communities. The importance of local habitat variables in urban green spaces and strong support for the habitat complexity hypothesis in a number of other ecosystems has led to proposals that "biodiverse" roofs--those intentionally designed with varied substrate depth, greater plant diversity, or added elements such as logs or stones--would support greater invertebrate diversity, but there is currently limited peer reviewed data to support this. In order to address the habitat complexity hypothesis in the context of green roofs, in the second part of this study I surveyed three roofs designed primarily for stormwater management, three biodiverse roofs, and five ground-level green spaces, from March until September of 2014 in the Portland metropolitan area. Beetles (Coleoptera) were sampled bi-weekly as representatives of total species diversity. Biodiverse roofs had greater richness, abundance, and diversity of beetle species compared to stormwater roofs, but were not more diverse than ground sites. Both biodiverse roofs and ground sites had approximately 20% native beetle species while stormwater roofs had only 5%. Functional diversity was also higher on biodiverse roofs with an average of seven trophic groups represented, while stormwater roofs averaged only three. Ground sites, biodiverse roofs, and stormwater roofs each grouped distinctively in terms of beetle community composition and biodiverse roof communities were found to be positively correlated with roof age, percent plant cover, average plant height, and plant species richness. These results support the findings of previous studies on the importance of local variables in structuring urban invertebrate communities and suggest that biodiverse design can reliably increase greenroof diversity, with the caution that they remain no replacement for ground level conservation. Title: Green Roofs and Urban Biodiversity: Their Role as Invertebrate Habitat and the Effect of Design on Beetle Community Author: Gonsalves, Sydney Marie Year: 2016 With over half the world's population now living in cities, urban areas represent one of earth's few ecosystems that are increasing in extent, and are sites of altered biogeochemical cycles, habitat fragmentation, and changes in biodiversity. However, urban green spaces, including green roofs, can also provide important pools of biodiversity and contribute to regional gamma diversity, while novel species assemblages can enhance some ecosystem services. Green roofs may also mitigate species loss in urban areas and have been shown to support a surprising diversity of invertebrates, including rare and endangered species. In the first part of this study I reviewed the literature on urban invertebrate communities and diversity to better understand the role of green roofs in providing habitat in the context of the larger urban mosaic. My review concluded that, while other factors such as surrounding land use and connectivity are also important to specific invertebrate taxa, local habitat variables contribute substantially to the structure and diversity of urban invertebrate communities. The importance of local habitat variables in urban green spaces and strong support for the habitat complexity hypothesis in a number of other ecosystems has led to proposals that "biodiverse" roofs--those intentionally designed with varied substrate depth, greater plant diversity, or added elements such as logs or stones--would support greater invertebrate diversity, but there is currently limited peer reviewed data to support this. In order to address the habitat complexity hypothesis in the context of green roofs, in the second part of this study I surveyed three roofs designed primarily for stormwater management, three biodiverse roofs, and five ground-level green spaces, from March until September of 2014 in the Portland metropolitan area. Beetles (Coleoptera) were sampled bi-weekly as representatives of total species diversity. Biodiverse roofs had greater richness, abundance, and diversity of beetle species compared to stormwater roofs, but were not more diverse than ground sites. Both biodiverse roofs and ground sites had approximately 20% native beetle species while stormwater roofs had only 5%. Functional diversity was also higher on biodiverse roofs with an average of seven trophic groups represented, while stormwater roofs averaged only three. Ground sites, biodiverse roofs, and stormwater roofs each grouped distinctively in terms of beetle community composition and biodiverse roof communities were found to be positively correlated with roof age, percent plant cover, average plant height, and plant species richness. These results support the findings of previous studies on the importance of local variables in structuring urban invertebrate communities and suggest that biodiverse design can reliably increase greenroof diversity, with the caution that they remain no replacement for ground level conservation. Close

• 18. [Article] Spatial heterogeneity, meta-analysis and spatial dependence : three essays on environmental valuation and spatial land management decisions

  This dissertation consists of three essays that collectively address the importance of accounting for spatial heterogeneity and spatial dependence of environmental assets and natural resources in policy ...

  Citation × Citation Citation Abstract Copy Title: Spatial heterogeneity, meta-analysis and spatial dependence : three essays on environmental valuation and spatial land management decisions Author: Bu, Meidan This dissertation consists of three essays that collectively address the importance of accounting for spatial heterogeneity and spatial dependence of environmental assets and natural resources in policy making. The first essay examines the value of different wetland ecosystem services using an expanded meta-dataset of valuation studies in North America. The purpose of this essay is to explore the sources of variation in wetland values from valuation methodology, ecosystem functions, and site-specific geographic, demographic and socio-economic characteristics. This essay used panel random effect models stratified "by study", "by state" and "by wetland site" to address the issues of correlation between wetland value estimates. Results indicated that wetland welfare measures reported by the same study and in the same wetland site are correlated. By comparing regression results and conclusions from this study with two internationally scoped meta-analysis studies, this essay found that the wetland valuation literature is not robust to regional characteristics, and wetland welfare estimates are sensitive to geographic extent. The second essay extends the first essay to investigate spatial spillover effects of wetland welfare estimates in North America. The primary purpose of the essay is to explore whether wetland values are correlated across space and what determines the correlation. The goal is accomplished by incorporating spatial econometric methods into the meta-analysis framework. The essay constructed three spatial weight matrices based on threshold distances, the ecological similarity and the economic similarity of wetland sites in the metadata. Results indicate that spatial proximity is an important predictor of wetland values regardless of wetland type and function. In part this is captured through shared contexts including resource availability and market characteristics. However, similar ecological and economic profiles are insufficient to capture shared values when transferring value estimates across spatial areas. Results from this study also raised questions on whether ecological functions and economic markets are sufficient for improving international transfers when studies are beyond geographic thresholds. The third essay incorporates spatial econometric models into a stochastic optimization framework to explore the consequences of ignoring the spatial linkage of management units in land management outcomes. The integrated framework combines an economic decision model, a spatial fire simulation model, a spotted owl population prediction model to maximize the expected post-fire and post-treatment NSO population under a budget constraint. Results from this essay inform us that ignoring spatial habitat connections leads to an underestimate of the fuel treatment disturbance and an over-estimate of the expected post-fire and post-treatment NSO population. However, the negative externality of habitat conversions depends on the degree of habitat connections. Additionally, the amount of total treatment area depends on the degree of habitat connections. The optimization outcome suggests less fuel treatment for a landscape with a higher degree of habitat connections. Moreover, the optimization outcome informs us that the spatial configuration of fuel treatment matters for the NSO population protection. Ignoring the spatial habitat connections leads to a fragmented treatment pattern and fails to protect the core NSO habitat from treatment disturbances. Title: Spatial heterogeneity, meta-analysis and spatial dependence : three essays on environmental valuation and spatial land management decisions Author: Bu, Meidan This dissertation consists of three essays that collectively address the importance of accounting for spatial heterogeneity and spatial dependence of environmental assets and natural resources in policy making. The first essay examines the value of different wetland ecosystem services using an expanded meta-dataset of valuation studies in North America. The purpose of this essay is to explore the sources of variation in wetland values from valuation methodology, ecosystem functions, and site-specific geographic, demographic and socio-economic characteristics. This essay used panel random effect models stratified "by study", "by state" and "by wetland site" to address the issues of correlation between wetland value estimates. Results indicated that wetland welfare measures reported by the same study and in the same wetland site are correlated. By comparing regression results and conclusions from this study with two internationally scoped meta-analysis studies, this essay found that the wetland valuation literature is not robust to regional characteristics, and wetland welfare estimates are sensitive to geographic extent. The second essay extends the first essay to investigate spatial spillover effects of wetland welfare estimates in North America. The primary purpose of the essay is to explore whether wetland values are correlated across space and what determines the correlation. The goal is accomplished by incorporating spatial econometric methods into the meta-analysis framework. The essay constructed three spatial weight matrices based on threshold distances, the ecological similarity and the economic similarity of wetland sites in the metadata. Results indicate that spatial proximity is an important predictor of wetland values regardless of wetland type and function. In part this is captured through shared contexts including resource availability and market characteristics. However, similar ecological and economic profiles are insufficient to capture shared values when transferring value estimates across spatial areas. Results from this study also raised questions on whether ecological functions and economic markets are sufficient for improving international transfers when studies are beyond geographic thresholds. The third essay incorporates spatial econometric models into a stochastic optimization framework to explore the consequences of ignoring the spatial linkage of management units in land management outcomes. The integrated framework combines an economic decision model, a spatial fire simulation model, a spotted owl population prediction model to maximize the expected post-fire and post-treatment NSO population under a budget constraint. Results from this essay inform us that ignoring spatial habitat connections leads to an underestimate of the fuel treatment disturbance and an over-estimate of the expected post-fire and post-treatment NSO population. However, the negative externality of habitat conversions depends on the degree of habitat connections. Additionally, the amount of total treatment area depends on the degree of habitat connections. The optimization outcome suggests less fuel treatment for a landscape with a higher degree of habitat connections. Moreover, the optimization outcome informs us that the spatial configuration of fuel treatment matters for the NSO population protection. Ignoring the spatial habitat connections leads to a fragmented treatment pattern and fails to protect the core NSO habitat from treatment disturbances. Close

• 19. [Article] Effects of spatially dispersed green-tree retention on ectomycorrhiza diversity

  The Demonstration for Ecosystem Management Options (DEMO) study originated out of the changing management priorities associated with federal forest lands in the Pacific Northwest which included an objective ...

  Citation × Citation Citation Abstract Copy Title: Effects of spatially dispersed green-tree retention on ectomycorrhiza diversity Author: Kolaczkowski, Oralia The Demonstration for Ecosystem Management Options (DEMO) study originated out of the changing management priorities associated with federal forest lands in the Pacific Northwest which included an objective to maintain mature and old-growth forest characteristics in managed stands. The DEMO project examines the effects that different levels and patterns of green tree retention have on various aspects of biodiversity including the diversity of ectomycorrhizal fungi (EMF). Ectomycorrhizal fungi play an important functional role in forest ecosystems. Ectomycorrhizal fungi are essential for nutrient uptake by the host tree, they have been shown to play a critical role in forest regeneration success, and EMF provide an important food source (in the form of sporocarps) for many small mammals. As part of the DEMO study, this thesis documents EMF diversity in the control unit (100% retention) and the spatially dispersed 40°/a basal area retention treatment (40%D). The measures of diversity studied included EMF constancy, abundance, and species richness. Constancy and abundance measures were analyzed only for the common ectomycorrhizae (EM) types. Morphological techniques for identification of the EMF were employed together with molecular techniques to achieve the finest level of identification possible. Characters used to assess morphology include mantle structures, emanating hyphae, rhizomorphs, root morphology, chemical reactions, and mantle color. Molecular techniques were used for identification purposes and for checking EM morphotype groups. Molecular techniques used included DNA extraction of the internal transcribed spacer region, polymerase chain reaction amplification, restriction fragment length polymorphism, and DNA sequencing. Significant responses to the 40%D treatment were determined by testing both pre and post EMF diversity within the treatment and by comparing changes in diversity between the control and the 40%D treatment. Overall, within the 40%D treatment, the majority of the common EMF species did not decrease significantly. A moderately significant difference was detected between the pre-treatment and post-treatment mean number of EM types per soil core (measure of species richness). This decrease in richness of about 50% in the post-treatment2 samples seems to indicate a delayed response to the treatment since the post-treatment1 samples were not different from the pre-treatment conditions. Since no dominant types were lost, the decline in richness occurred in the less common EM types. This research demonstrates that the 40%D treatment can maintain a relatively high legacy of EMF. However, there is a loss of some rare EM types and a reduction in abundance and constancy in some of the more common EM types. These results lend support for the consideration of40% dispersed green-tree retention as a silvicultural option when management goals include the maintenance of EMF diversity. Timber harvest via implementation of ecosystem-based forest management practices will require the continuing study of complex below-ground systems for the development of potential silvicultural treatments during further iterations of adaptive management. Title: Effects of spatially dispersed green-tree retention on ectomycorrhiza diversity Author: Kolaczkowski, Oralia The Demonstration for Ecosystem Management Options (DEMO) study originated out of the changing management priorities associated with federal forest lands in the Pacific Northwest which included an objective to maintain mature and old-growth forest characteristics in managed stands. The DEMO project examines the effects that different levels and patterns of green tree retention have on various aspects of biodiversity including the diversity of ectomycorrhizal fungi (EMF). Ectomycorrhizal fungi play an important functional role in forest ecosystems. Ectomycorrhizal fungi are essential for nutrient uptake by the host tree, they have been shown to play a critical role in forest regeneration success, and EMF provide an important food source (in the form of sporocarps) for many small mammals. As part of the DEMO study, this thesis documents EMF diversity in the control unit (100% retention) and the spatially dispersed 40°/a basal area retention treatment (40%D). The measures of diversity studied included EMF constancy, abundance, and species richness. Constancy and abundance measures were analyzed only for the common ectomycorrhizae (EM) types. Morphological techniques for identification of the EMF were employed together with molecular techniques to achieve the finest level of identification possible. Characters used to assess morphology include mantle structures, emanating hyphae, rhizomorphs, root morphology, chemical reactions, and mantle color. Molecular techniques were used for identification purposes and for checking EM morphotype groups. Molecular techniques used included DNA extraction of the internal transcribed spacer region, polymerase chain reaction amplification, restriction fragment length polymorphism, and DNA sequencing. Significant responses to the 40%D treatment were determined by testing both pre and post EMF diversity within the treatment and by comparing changes in diversity between the control and the 40%D treatment. Overall, within the 40%D treatment, the majority of the common EMF species did not decrease significantly. A moderately significant difference was detected between the pre-treatment and post-treatment mean number of EM types per soil core (measure of species richness). This decrease in richness of about 50% in the post-treatment2 samples seems to indicate a delayed response to the treatment since the post-treatment1 samples were not different from the pre-treatment conditions. Since no dominant types were lost, the decline in richness occurred in the less common EM types. This research demonstrates that the 40%D treatment can maintain a relatively high legacy of EMF. However, there is a loss of some rare EM types and a reduction in abundance and constancy in some of the more common EM types. These results lend support for the consideration of40% dispersed green-tree retention as a silvicultural option when management goals include the maintenance of EMF diversity. Timber harvest via implementation of ecosystem-based forest management practices will require the continuing study of complex below-ground systems for the development of potential silvicultural treatments during further iterations of adaptive management. Close

• 20. [Image] Integrated scientific assessment for ecosystem management in the interior Columbia Basin and portions of the Klamath and Great Basins

  	Abstract Quigley, Thomas M.; Haynes, Richard W.; Graham, Russell T., tech. eds. 1996. Integrated scientific assessment for ecosystem management in the interior Columbia basin and portions of the Klamath ...
  	Citation × Citation Citation Abstract Copy Title: Integrated scientific assessment for ecosystem management in the interior Columbia Basin and portions of the Klamath and Great Basins Year: 1996, 2005, 2000 Abstract Quigley, Thomas M.; Haynes, Richard W.; Graham, Russell T., tech. eds. 1996. Integrated scientific assessment for ecosystem management in the interior Columbia basin and portions of the Klamath and Great Basins. Gen. Tech. Rep. PNW-GTR-382. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 303 p. (Quigley, Thomas M., tech. ed. The Interior Columbia Basin Ecosystem Management Project: Scientific Assessment.) The Integrated Scientific Assessment for Ecosystem Management for the Interior Columbia Basin links landscape, aquatic, terrestrial, social, and economic characterizations to describe biophysical and social systems. Integration was achieved through a framework built around six goals for ecosystem management and three different views of the future. These goals are: maintain evolutionary and ecological processes; manage for multiple ecological domains and evolutionary timeframes; maintain viable populations of native and desired non-native species; encourage social and economic resiliency; manage for places with definable values; and, manage to maintain a variety of ecosystem goods, services, and conditions that society wants. Ratings of relative ecological integrity and socioeconomic resiliency were used to make broad statements about ecosystem conditions in the Basin. Currently in the Basin high integrity and resiliency are found on 16 and 20 percent of the area, respectively. Low integrity and resiliency are found on 60 and 68 percent of the area. Different approaches to management can alter the risks to the assets of people living in the Basin and to the ecosystem itself. Continuation of current management leads to increasing risks while management approaches focusing on reserves or restoration result in trends that mostly stabilize or reduce risks. Even where ecological integrity is projected to improve with the application of active management, population increases and the pressures of expanding demands on resources may cause increasing trends in risk. Keywords: Ecosystem assessment, management and goals; ecological integrity; socio-economic resiliency; risk management Title: Integrated scientific assessment for ecosystem management in the interior Columbia Basin and portions of the Klamath and Great Basins Year: 1996, 2005, 2000 Abstract Quigley, Thomas M.; Haynes, Richard W.; Graham, Russell T., tech. eds. 1996. Integrated scientific assessment for ecosystem management in the interior Columbia basin and portions of the Klamath and Great Basins. Gen. Tech. Rep. PNW-GTR-382. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 303 p. (Quigley, Thomas M., tech. ed. The Interior Columbia Basin Ecosystem Management Project: Scientific Assessment.) The Integrated Scientific Assessment for Ecosystem Management for the Interior Columbia Basin links landscape, aquatic, terrestrial, social, and economic characterizations to describe biophysical and social systems. Integration was achieved through a framework built around six goals for ecosystem management and three different views of the future. These goals are: maintain evolutionary and ecological processes; manage for multiple ecological domains and evolutionary timeframes; maintain viable populations of native and desired non-native species; encourage social and economic resiliency; manage for places with definable values; and, manage to maintain a variety of ecosystem goods, services, and conditions that society wants. Ratings of relative ecological integrity and socioeconomic resiliency were used to make broad statements about ecosystem conditions in the Basin. Currently in the Basin high integrity and resiliency are found on 16 and 20 percent of the area, respectively. Low integrity and resiliency are found on 60 and 68 percent of the area. Different approaches to management can alter the risks to the assets of people living in the Basin and to the ecosystem itself. Continuation of current management leads to increasing risks while management approaches focusing on reserves or restoration result in trends that mostly stabilize or reduce risks. Even where ecological integrity is projected to improve with the application of active management, population increases and the pressures of expanding demands on resources may cause increasing trends in risk. Keywords: Ecosystem assessment, management and goals; ecological integrity; socio-economic resiliency; risk management Close

• 21. [Article] Landscape features affecting genetic diversity and structure in East African ungulate species

  Habitat loss and fragmentation is a crisis affecting wildlife worldwide. In Tanzania, East Africa, a dramatic and recent (<80 years) expansion in human settlement and agriculture threatens to reduce gene ...

  Citation × Citation Citation Abstract Copy Title: Landscape features affecting genetic diversity and structure in East African ungulate species Author: Crowhurst, Rachel Selena Habitat loss and fragmentation is a crisis affecting wildlife worldwide. In Tanzania, East Africa, a dramatic and recent (<80 years) expansion in human settlement and agriculture threatens to reduce gene flow among protected areas for many species of large mammals. Wildlife linkages can mitigate population isolation, but linkage designs lacking empirical justification may be controversial and ineffective. Connectivity conservation requires an understanding of how biogeographic factors shaped gene flow prior to habitat loss or fragmentation, however the history of interaction among populations is rarely known. The goal of my study was to provide context for connectivity conservation in central and southern Tanzania by identifying landscape features that have shaped gene flow for three ungulate species with different dispersal capabilities. I investigated historical patterns of connectivity for Maasai giraffe (Giraffa camelopardalis tippelskirchi), impala (Aepyceros melampus), and eland (Tragelaphus oryx) by estimating genetic structure among four to eight protected areas per species. Genetic structure changes very slowly among large populations and thus is likely to reflect historical processes instead of recent anthropogenic influences. I collected noninvasive DNA samples and generated microsatellite genotypes at 8 to 15 loci per species, then estimated genetic diversity metrics (allelic richness, AR, and expected heterozygosity, H[subscript E]) for each population (defined by reserve). I also calculated genetic distance (F[subscript ST] and Nei's unbiased genetic distance, D[subscript hat]) and an estimate of gene flow (Nm) between all population pairs for each species. To elucidate the possible causes of genetic structure between these populations, I tested for isolation by distance and isolation by resistance based on a suite of biogeographic factors hypothesized to affect gene flow. To do this, I created GIS-based resistance surfaces that assigned different costs of movement to landscape features. I created one or more resistance surfaces for each hypothesis of landscape effect. I used circuit theory to estimate the cumulative resistance between each pair of reserves for each weighting scheme, and then performed Mantel tests to calculate the correlation between these resistances and the observed population pairwise genetic distances (D[subscript hat]). I chose the optimal resistance model for each species as the model that was most highly correlated with observed genetic patterns. To verify that the correlation of resistance models with genetic distance was not an artefact of geographic distance, I performed partial Mantel tests to calculate correlation while controlling for the effect of geographic distance. Finally, I compared historical gene flow patterns to the distribution of contemporary human activity to predict areas that are at risk of a loss of connectivity. Indices of genetic diversity were moderate for all three species and comparable to previously reported values for other savannah ungulates. Diversity (both H[subscript E] and A[subscript R]) was highest in eland and lowest in giraffe for these populations, and was not consistently correlated with reserve size as has been reported for other species in East Africa. Although patterns in genetic distance were broadly similar across all three species there were also striking differences in connectivity, highlighting the importance of cross-species comparisons in connectivity conservation. At this scale, resistance models based on slope strongly predicted population structure for all three species; distance to water was also highly correlated with genetic distance in eland. For all three species, the greatest genetic distances occurred between populations separated by the Eastern Arc Mountains, suggesting that the topography of this area has long acted as a barrier to gene flow, but this effect is present in varying degrees for each species. I observed high levels of historical gene flow between centrally located populations (Ruaha National Park and Rungwa Game Reserve) and those in the southwest (Katavi National Park and Rukwa Game Reserve). Although human settlement in this area has been low relative to other areas, the connection between the Katavi/Rukwa and Ruaha ecosystems may be threatened by increased human activity and warrants conservation. High levels of historical gene flow were also seen between reserves in the northeast (Tarangire National Park, Swagaswaga Game Reserve) and the central and southwest populations. These connections appear highly threatened due to current land use practices, and may have already suffered a loss of gene flow. Field surveys in the lands surrounding the northeastern reserves are needed to quantify current levels of connectivity and determine whether corridors could be established to maintain or restore gene flow with other reserves. Title: Landscape features affecting genetic diversity and structure in East African ungulate species Author: Crowhurst, Rachel Selena Habitat loss and fragmentation is a crisis affecting wildlife worldwide. In Tanzania, East Africa, a dramatic and recent (<80 years) expansion in human settlement and agriculture threatens to reduce gene flow among protected areas for many species of large mammals. Wildlife linkages can mitigate population isolation, but linkage designs lacking empirical justification may be controversial and ineffective. Connectivity conservation requires an understanding of how biogeographic factors shaped gene flow prior to habitat loss or fragmentation, however the history of interaction among populations is rarely known. The goal of my study was to provide context for connectivity conservation in central and southern Tanzania by identifying landscape features that have shaped gene flow for three ungulate species with different dispersal capabilities. I investigated historical patterns of connectivity for Maasai giraffe (Giraffa camelopardalis tippelskirchi), impala (Aepyceros melampus), and eland (Tragelaphus oryx) by estimating genetic structure among four to eight protected areas per species. Genetic structure changes very slowly among large populations and thus is likely to reflect historical processes instead of recent anthropogenic influences. I collected noninvasive DNA samples and generated microsatellite genotypes at 8 to 15 loci per species, then estimated genetic diversity metrics (allelic richness, AR, and expected heterozygosity, H[subscript E]) for each population (defined by reserve). I also calculated genetic distance (F[subscript ST] and Nei's unbiased genetic distance, D[subscript hat]) and an estimate of gene flow (Nm) between all population pairs for each species. To elucidate the possible causes of genetic structure between these populations, I tested for isolation by distance and isolation by resistance based on a suite of biogeographic factors hypothesized to affect gene flow. To do this, I created GIS-based resistance surfaces that assigned different costs of movement to landscape features. I created one or more resistance surfaces for each hypothesis of landscape effect. I used circuit theory to estimate the cumulative resistance between each pair of reserves for each weighting scheme, and then performed Mantel tests to calculate the correlation between these resistances and the observed population pairwise genetic distances (D[subscript hat]). I chose the optimal resistance model for each species as the model that was most highly correlated with observed genetic patterns. To verify that the correlation of resistance models with genetic distance was not an artefact of geographic distance, I performed partial Mantel tests to calculate correlation while controlling for the effect of geographic distance. Finally, I compared historical gene flow patterns to the distribution of contemporary human activity to predict areas that are at risk of a loss of connectivity. Indices of genetic diversity were moderate for all three species and comparable to previously reported values for other savannah ungulates. Diversity (both H[subscript E] and A[subscript R]) was highest in eland and lowest in giraffe for these populations, and was not consistently correlated with reserve size as has been reported for other species in East Africa. Although patterns in genetic distance were broadly similar across all three species there were also striking differences in connectivity, highlighting the importance of cross-species comparisons in connectivity conservation. At this scale, resistance models based on slope strongly predicted population structure for all three species; distance to water was also highly correlated with genetic distance in eland. For all three species, the greatest genetic distances occurred between populations separated by the Eastern Arc Mountains, suggesting that the topography of this area has long acted as a barrier to gene flow, but this effect is present in varying degrees for each species. I observed high levels of historical gene flow between centrally located populations (Ruaha National Park and Rungwa Game Reserve) and those in the southwest (Katavi National Park and Rukwa Game Reserve). Although human settlement in this area has been low relative to other areas, the connection between the Katavi/Rukwa and Ruaha ecosystems may be threatened by increased human activity and warrants conservation. High levels of historical gene flow were also seen between reserves in the northeast (Tarangire National Park, Swagaswaga Game Reserve) and the central and southwest populations. These connections appear highly threatened due to current land use practices, and may have already suffered a loss of gene flow. Field surveys in the lands surrounding the northeastern reserves are needed to quantify current levels of connectivity and determine whether corridors could be established to maintain or restore gene flow with other reserves. Close

• 22. [Article] Broad-scale patterns of invertebrate richness and community composition in temporary rivers: effects of flow intermittence

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by John Wiley ...

  Citation × Citation Citation Abstract Copy Title: Broad-scale patterns of invertebrate richness and community composition in temporary rivers: effects of flow intermittence Author: Larned, S. T., Bogan, M. T., Wood, P. J., Santos, A. N., Meyer, E. I., Datry, T., Fritz, K. M. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by John Wiley & Sons, Ltd. and can be found at: http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1600-0587. Title: Broad-scale patterns of invertebrate richness and community composition in temporary rivers: effects of flow intermittence Author: Larned, S. T., Bogan, M. T., Wood, P. J., Santos, A. N., Meyer, E. I., Datry, T., Fritz, K. M. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by John Wiley & Sons, Ltd. and can be found at: http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1600-0587. Close

• 23. [Article] Intercomparison of MODIS albedo retrievals and in situ measurements across the global FLUXNET network

  This is the publisher’s final pdf. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/remote-sensing-of-environment/#description. To the best of our ...

  Citation × Citation Citation Abstract Copy Title: Intercomparison of MODIS albedo retrievals and in situ measurements across the global FLUXNET network Author: Yang, Xiaoyuan, Vannan, Suresh K. Santhana, Law, Beverly E., Richardson, Andrew D., Roman, Miguel O., Moors, Eddy, Matteucci, Giorgio, Migliavacca, Mirco, Seufert, Guenther, Ciais, Philippe, Cook, Robert B., Schaaf, Crystal B., Pan, Jerry Yun, Cescatti, Alessandro, Marcolla, Barbara This is the publisher’s final pdf. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/remote-sensing-of-environment/#description. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. Title: Intercomparison of MODIS albedo retrievals and in situ measurements across the global FLUXNET network Author: Yang, Xiaoyuan, Vannan, Suresh K. Santhana, Law, Beverly E., Richardson, Andrew D., Roman, Miguel O., Moors, Eddy, Matteucci, Giorgio, Migliavacca, Mirco, Seufert, Guenther, Ciais, Philippe, Cook, Robert B., Schaaf, Crystal B., Pan, Jerry Yun, Cescatti, Alessandro, Marcolla, Barbara This is the publisher’s final pdf. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/remote-sensing-of-environment/#description. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. Close

• 24. [Article] The Icelandic Pelagic Sector and its Development under an ITQ Management System

  Suggested Bibliographic Reference: Challenging New Frontiers in the Global Seafood Sector: Proceedings of the Eighteenth Biennial Conference of the International Institute of Fisheries Economics and Trade, ...

  Citation × Citation Citation Abstract Copy Title: The Icelandic Pelagic Sector and its Development under an ITQ Management System Author: Saevaldsson, Hordur, Gunnlaugsson, Stefan B. Year: 2016 Suggested Bibliographic Reference: Challenging New Frontiers in the Global Seafood Sector: Proceedings of the Eighteenth Biennial Conference of the International Institute of Fisheries Economics and Trade, July 11-15, 2016. Compiled by Stefani J. Evers and Ann L. Shriver. International Institute of Fisheries Economics and Trade (IIFET), Corvallis, 2016. Title: The Icelandic Pelagic Sector and its Development under an ITQ Management System Author: Saevaldsson, Hordur, Gunnlaugsson, Stefan B. Year: 2016 Suggested Bibliographic Reference: Challenging New Frontiers in the Global Seafood Sector: Proceedings of the Eighteenth Biennial Conference of the International Institute of Fisheries Economics and Trade, July 11-15, 2016. Compiled by Stefani J. Evers and Ann L. Shriver. International Institute of Fisheries Economics and Trade (IIFET), Corvallis, 2016. Close

• 25. [Article] Analyzing the transmission of wildfire exposure on a fire-prone landscape in Oregon, USA

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Elsevier ...

  Citation × Citation Citation Abstract Copy Title: Analyzing the transmission of wildfire exposure on a fire-prone landscape in Oregon, USA Author: Ager, Alan A., Vaillant, Nicole M., Finney, Mark A., Vance-Borland, Ken, Day, Michelle A. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/forest-ecology-and-management. Title: Analyzing the transmission of wildfire exposure on a fire-prone landscape in Oregon, USA Author: Ager, Alan A., Vaillant, Nicole M., Finney, Mark A., Vance-Borland, Ken, Day, Michelle A. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/forest-ecology-and-management. Close

• 26. [Article] Sex-biased dispersal in a rare butterfly and the implications for its conservation

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer ...

  Citation × Citation Citation Abstract Copy Title: Sex-biased dispersal in a rare butterfly and the implications for its conservation Author: Bennett, Victoria J., Smith, Winston P., Pack, Shalynn M., Betts, Matthew G. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer and can be found at: http://www.springer.com/?SGWID=0-102-0-0-0. Title: Sex-biased dispersal in a rare butterfly and the implications for its conservation Author: Bennett, Victoria J., Smith, Winston P., Pack, Shalynn M., Betts, Matthew G. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer and can be found at: http://www.springer.com/?SGWID=0-102-0-0-0. Close

• 27. [Article] How sweet is the land? Modeling caloric landscapes for tropical hummingbirds

  Pollination is a critical ecosystem function for sustaining biodiversity. However, pollinators and the services they provide are threatened by landscape-altering anthropogenic activities across the globe. ...

  Citation × Citation Citation Abstract Copy Title: How sweet is the land? Modeling caloric landscapes for tropical hummingbirds Author: Birkett, Christina Nozomi Pollination is a critical ecosystem function for sustaining biodiversity. However, pollinators and the services they provide are threatened by landscape-altering anthropogenic activities across the globe. Habitat loss and fragmentation, introduction of invasive species, chemical use, and urbanization have been shown to impact pollination. Pollinator foraging behavior is thought to be largely a function of available floral rewards, therefore, understanding the role of resource distributions in pollinator abundance and behavior within disturbed landscapes is a key piece of information for conservation. Fine scale information on floral resource distribution across disturbed landscapes is lacking in most systems. Here we demonstrate how existing presence-only species distribution modeling techniques (i.e., Maximum entropy modeling [MaxEnt]) can be combined with widely available environmental information to create resource landscapes for both pollinator communities and specific pollinators of interest. This model is the first of its kind, making possible simultaneous visualization of fine-scale resource configuration and quantity across abroad spatial extent. We tested this method to build caloric landscapes using tropical hummingbird-plant system in Costa Rica. We found that our MaxEnt models performed well on independent data for all 13 flower species we examined. Our landscape-scale caloric map showed that available calories within each 35m² pixel ranged from 0 to greater than 30,000 across our study region. Our model provides the possibility of predicting pollinator movement and abundance based upon resource supply. As its parameters are flexible, it is broad in its potential applications. The flexibility in calibration to desired resource landscapes permits applying the model to other pollinator-plant systems. We hope that this model will complement the current ecologist’s toolbox, aiding in ensuring the continuation and health of pollinator systems. Title: How sweet is the land? Modeling caloric landscapes for tropical hummingbirds Author: Birkett, Christina Nozomi Pollination is a critical ecosystem function for sustaining biodiversity. However, pollinators and the services they provide are threatened by landscape-altering anthropogenic activities across the globe. Habitat loss and fragmentation, introduction of invasive species, chemical use, and urbanization have been shown to impact pollination. Pollinator foraging behavior is thought to be largely a function of available floral rewards, therefore, understanding the role of resource distributions in pollinator abundance and behavior within disturbed landscapes is a key piece of information for conservation. Fine scale information on floral resource distribution across disturbed landscapes is lacking in most systems. Here we demonstrate how existing presence-only species distribution modeling techniques (i.e., Maximum entropy modeling [MaxEnt]) can be combined with widely available environmental information to create resource landscapes for both pollinator communities and specific pollinators of interest. This model is the first of its kind, making possible simultaneous visualization of fine-scale resource configuration and quantity across abroad spatial extent. We tested this method to build caloric landscapes using tropical hummingbird-plant system in Costa Rica. We found that our MaxEnt models performed well on independent data for all 13 flower species we examined. Our landscape-scale caloric map showed that available calories within each 35m² pixel ranged from 0 to greater than 30,000 across our study region. Our model provides the possibility of predicting pollinator movement and abundance based upon resource supply. As its parameters are flexible, it is broad in its potential applications. The flexibility in calibration to desired resource landscapes permits applying the model to other pollinator-plant systems. We hope that this model will complement the current ecologist’s toolbox, aiding in ensuring the continuation and health of pollinator systems. Close

• 28. [Article] Microbial community structure and dynamics in restored subtropical seagrass sediments

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Inter-Research ...

  Citation × Citation Citation Abstract Copy Title: Microbial community structure and dynamics in restored subtropical seagrass sediments Author: Fourqurean, James W., Bourque, Amanda S., Vega-Thurber, Rebecca To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Inter-Research and can be found at: http://www.int-res.com/journals/ame/ame-home/. Title: Microbial community structure and dynamics in restored subtropical seagrass sediments Author: Fourqurean, James W., Bourque, Amanda S., Vega-Thurber, Rebecca To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Inter-Research and can be found at: http://www.int-res.com/journals/ame/ame-home/. Close

• 29. [Article] Comparing Himalayan blackberry (Rubus armeniacus) management techniques in upland prairie communities of the W.L. Finley National Wildlife Refuge

  Dedicated to the preservation and promotion of many of the nation’s most threatened and endangered species, the W.L. Finley National Wildlife Refuge is highly invested in the management of some of the ...

  Citation × Citation Citation Abstract Copy Title: Comparing Himalayan blackberry (Rubus armeniacus) management techniques in upland prairie communities of the W.L. Finley National Wildlife Refuge Author: Ensley, Jona L. Dedicated to the preservation and promotion of many of the nation’s most threatened and endangered species, the W.L. Finley National Wildlife Refuge is highly invested in the management of some of the last remaining upland prairies once prevalent throughout Oregon's Willamette Valley. More than a century of land fragmentation, fire suppression, and cultivation has shifted species composition and physical structure of the native prairies toward woodland ecosystems, reducing habitat quality and quantity. This transition is greatly driven by the encroachment and introduction of the non-native, woody invader Himalayan blackberry (Rubus armeniacus). This study focused on the three most common Himalayan blackberry management techniques used within the refuge's upland prairies: late summer mowing, late summer burning, and a combination of both. The goal was to analyze the efficacy of each technique in meeting the refuge objectives of controlling Himalayan blackberry populations and promoting native prairie physiognomy after one application using a complete random block design. Findings were presented to refuge managers to facilitate improved management of these fragile communities. Mowing was found to have the greatest reduction in blackberry stem and plant density and resulted in the lowest resprout density post-treatment. This treatment also met the refuge objectives of reducing woody cover and increasing graminoid cover, but did little to increase herbaceous cover when compared to the control group. Burning produced no significant increase in herbaceous cover or reduction in woody cover and provided favorable conditions for Himalayan blackberry seedling germination, contributing to a larger blackberry problem in years to come. In promoting herbaceous habitat for upland-dependent species, mowing with subsequent burning was the most successful technique. Though not as effective in reducing blackberry vigor as mowing alone, this treatment showed the most potential of the three in managing for all objectives. Title: Comparing Himalayan blackberry (Rubus armeniacus) management techniques in upland prairie communities of the W.L. Finley National Wildlife Refuge Author: Ensley, Jona L. Dedicated to the preservation and promotion of many of the nation’s most threatened and endangered species, the W.L. Finley National Wildlife Refuge is highly invested in the management of some of the last remaining upland prairies once prevalent throughout Oregon's Willamette Valley. More than a century of land fragmentation, fire suppression, and cultivation has shifted species composition and physical structure of the native prairies toward woodland ecosystems, reducing habitat quality and quantity. This transition is greatly driven by the encroachment and introduction of the non-native, woody invader Himalayan blackberry (Rubus armeniacus). This study focused on the three most common Himalayan blackberry management techniques used within the refuge's upland prairies: late summer mowing, late summer burning, and a combination of both. The goal was to analyze the efficacy of each technique in meeting the refuge objectives of controlling Himalayan blackberry populations and promoting native prairie physiognomy after one application using a complete random block design. Findings were presented to refuge managers to facilitate improved management of these fragile communities. Mowing was found to have the greatest reduction in blackberry stem and plant density and resulted in the lowest resprout density post-treatment. This treatment also met the refuge objectives of reducing woody cover and increasing graminoid cover, but did little to increase herbaceous cover when compared to the control group. Burning produced no significant increase in herbaceous cover or reduction in woody cover and provided favorable conditions for Himalayan blackberry seedling germination, contributing to a larger blackberry problem in years to come. In promoting herbaceous habitat for upland-dependent species, mowing with subsequent burning was the most successful technique. Though not as effective in reducing blackberry vigor as mowing alone, this treatment showed the most potential of the three in managing for all objectives. Close

• 30. [Article] Assessing the Current State of Transboundary Cooperation in the International Columbia River Basin

  The Columbia River Treaty (CRT) is often used as an example of how treaties can work to normalize, regulate, and improve the cooperative management of shared water resources between basin countries. Few ...

  Citation × Citation Citation Abstract Copy Title: Assessing the Current State of Transboundary Cooperation in the International Columbia River Basin Author: Timboe, Ingrid A. The Columbia River Treaty (CRT) is often used as an example of how treaties can work to normalize, regulate, and improve the cooperative management of shared water resources between basin countries. Few would argue that, in its limited mandate to regulate hydropower generation and flood risk management for downstream communities, the CRT has been anything but a success. However, there are many aspects of water resource management, including water quality, aquatic species, and indigenous peoples' rights and responsibilities that are left unaddressed by the Treaty. These issues are of increasing importance to managers and stakeholders in river basins around the world. This paper asks, in the absence of a formal agreement governing these issues, are the United States and Canada coordinating the broader management of the Columbia River and its international tributaries? If so, around which water resource issues do they collaborate and at what level? If not, do these gaps in co-management negatively affect the basin? In order to assess the current state of transboundary water cooperation between the United States and Canada within the basin, 50 transboundary initiatives were identified and categorized into management categories including: ecosystem function, fish passage & habitat, climate change, invasive species or toxics management, and energy. Interviews were then conducted with 34 natural resource managers throughout the international basin on their work with these initiatives, specifically addressing areas where they are, and are not, collaborating at this time. The interviews revealed that, while there is a great deal of collaborative work being done, it appears largely fragmented and restricted to sub-basin activities. International, basin-wide cooperation, outside of treaty obligations under the CRT, appears limited at this time. This thesis contends that there is a need for these sub-basin activities to be networked with one another, either through the Treaty, or through an inclusive, legitimate, and transparent river basin organization. Title: Assessing the Current State of Transboundary Cooperation in the International Columbia River Basin Author: Timboe, Ingrid A. The Columbia River Treaty (CRT) is often used as an example of how treaties can work to normalize, regulate, and improve the cooperative management of shared water resources between basin countries. Few would argue that, in its limited mandate to regulate hydropower generation and flood risk management for downstream communities, the CRT has been anything but a success. However, there are many aspects of water resource management, including water quality, aquatic species, and indigenous peoples' rights and responsibilities that are left unaddressed by the Treaty. These issues are of increasing importance to managers and stakeholders in river basins around the world. This paper asks, in the absence of a formal agreement governing these issues, are the United States and Canada coordinating the broader management of the Columbia River and its international tributaries? If so, around which water resource issues do they collaborate and at what level? If not, do these gaps in co-management negatively affect the basin? In order to assess the current state of transboundary water cooperation between the United States and Canada within the basin, 50 transboundary initiatives were identified and categorized into management categories including: ecosystem function, fish passage & habitat, climate change, invasive species or toxics management, and energy. Interviews were then conducted with 34 natural resource managers throughout the international basin on their work with these initiatives, specifically addressing areas where they are, and are not, collaborating at this time. The interviews revealed that, while there is a great deal of collaborative work being done, it appears largely fragmented and restricted to sub-basin activities. International, basin-wide cooperation, outside of treaty obligations under the CRT, appears limited at this time. This thesis contends that there is a need for these sub-basin activities to be networked with one another, either through the Treaty, or through an inclusive, legitimate, and transparent river basin organization. Close

• 31. [Article] Host selection and wood channelization by beetles in conifer logs in western Oregon

  The host preferences and wood channelization rates of scolytid and cerambycid beetles were studied at the H. J. Andrews Experimental Forest (Lane County, Oregon) during 1986 and 1987. Attack density and ...

  Citation × Citation Citation Abstract Copy Title: Host selection and wood channelization by beetles in conifer logs in western Oregon Author: Zhong, He The host preferences and wood channelization rates of scolytid and cerambycid beetles were studied at the H. J. Andrews Experimental Forest (Lane County, Oregon) during 1986 and 1987. Attack density and gallery volumes were measured in fall-felled experimental logs of four major conifer species: Douglas-fir (Pseudotsuga mensiesii (Mirb.) Franco), western hemlock (Tsuga heterophylla (Raf.)Sarg.), Pacific silver fir (Abies amabalis (Dougl.)Forbes), and western redcedar (Thu'a plicata Donn). The ambrosia beetles (Coleoptera: Scolytidae), Trypodendron lineatum (Olivier) and Gnathotrichus spp., were significantly more abundant in Douglas-fir and western hemlock than in silver fir or western redcedar, but Xyleborinus saxeseni (Ratzeburg) showed no discrimination among the four tree species. The average emergence of ambrosia beetle brood per gallery was 1.8 in spring-felled Douglas-fir logs and 2.2 in western hemlock. The data indicate a declining ambrosia beetle population emerging from these logs. About 0.16% of the sapwood volume of Douglas-fir and about 0.18% of the sapwood volume of western hemlock were removed by Trypodendron. About 0.01% of the sapwood volume of Douglas-fir and about 0.06% of the sapwood volume of western hemlock were removed by Gnathotrichus. The bark beetles (Coleoptera: Scolytidae) Dendroctonus pseudotsugae Hopkins and Dryocoetes autographus Ratzeburg excavated about 7.6% of Douglas-fir phloem surface area. Pseudohylesinus sericeus (Mannerheim) removed 6.8% of Pacific silver fir phloem surface area. Long-horned beetles (Coleoptera: Cerambycidae) excavated an additional 2.3% of Pacific silver fir phloem surface area. These data indicate that each tree species has a distinct boring insect species pool with differences in channelization rates through time. Colonization and fragmentation of fallen trees by beetles are important to the initiation of the decomposition process in forest ecosystems. Title: Host selection and wood channelization by beetles in conifer logs in western Oregon Author: Zhong, He The host preferences and wood channelization rates of scolytid and cerambycid beetles were studied at the H. J. Andrews Experimental Forest (Lane County, Oregon) during 1986 and 1987. Attack density and gallery volumes were measured in fall-felled experimental logs of four major conifer species: Douglas-fir (Pseudotsuga mensiesii (Mirb.) Franco), western hemlock (Tsuga heterophylla (Raf.)Sarg.), Pacific silver fir (Abies amabalis (Dougl.)Forbes), and western redcedar (Thu'a plicata Donn). The ambrosia beetles (Coleoptera: Scolytidae), Trypodendron lineatum (Olivier) and Gnathotrichus spp., were significantly more abundant in Douglas-fir and western hemlock than in silver fir or western redcedar, but Xyleborinus saxeseni (Ratzeburg) showed no discrimination among the four tree species. The average emergence of ambrosia beetle brood per gallery was 1.8 in spring-felled Douglas-fir logs and 2.2 in western hemlock. The data indicate a declining ambrosia beetle population emerging from these logs. About 0.16% of the sapwood volume of Douglas-fir and about 0.18% of the sapwood volume of western hemlock were removed by Trypodendron. About 0.01% of the sapwood volume of Douglas-fir and about 0.06% of the sapwood volume of western hemlock were removed by Gnathotrichus. The bark beetles (Coleoptera: Scolytidae) Dendroctonus pseudotsugae Hopkins and Dryocoetes autographus Ratzeburg excavated about 7.6% of Douglas-fir phloem surface area. Pseudohylesinus sericeus (Mannerheim) removed 6.8% of Pacific silver fir phloem surface area. Long-horned beetles (Coleoptera: Cerambycidae) excavated an additional 2.3% of Pacific silver fir phloem surface area. These data indicate that each tree species has a distinct boring insect species pool with differences in channelization rates through time. Colonization and fragmentation of fallen trees by beetles are important to the initiation of the decomposition process in forest ecosystems. Close

• 32. [Article] How sweet is the land? Modeling caloric landscapes for tropical hummingbirds

  Powerpoint presentation for the defense of this thesis.

  Citation × Citation Citation Abstract Copy Title: How sweet is the land? Modeling caloric landscapes for tropical hummingbirds Author: Birkett, Christina Nozomi Powerpoint presentation for the defense of this thesis. Title: How sweet is the land? Modeling caloric landscapes for tropical hummingbirds Author: Birkett, Christina Nozomi Powerpoint presentation for the defense of this thesis. Close

• 33. [Article] Policy Patterns across Riverscapes : Riparian Land Standards in the Oregon Coast Range

  Land management policies are ideas about nature projected onto the landscape. Culminations of social, economic, and scientific influences, these policies create standards affecting the function of ecological ...

  Citation × Citation Citation Abstract Copy Title: Policy Patterns across Riverscapes : Riparian Land Standards in the Oregon Coast Range Author: Boisjolie, Brett A. Land management policies are ideas about nature projected onto the landscape. Culminations of social, economic, and scientific influences, these policies create standards affecting the function of ecological systems. In the case of riparian lands in the Oregon Coast Range, policy requirements vary considerably across federal, state, and private land ownerships. Protective measures, such as the adoption of fixed buffer widths for maintaining vegetation, are intended to preserve natural processes important to instream habitat and water quality. However, the "policy landscape" of variable management standards can result in a fragmented approach to policy protection. This fragmentation complicates recovery efforts for threatened anadromous fish species such as coho salmon (Oncorhynchus kisutch), which require connectivity in suitable habitat segments throughout the river network. Taking a riverscape perspective, this study evaluates how aquatic protections vary across the river and its valley. To explore the patterns created from variable standard, the study first delineates categories of riparian management standards in the Oregon Coast Range. Using a geographic information system (GIS), these standards are mapped across the river networks of this region. The extent of each policy category is quantified in stream kilometers in order to evaluate policy efforts within and across the riverscapes of the Oregon Coast Range. A model of intrinsic potential for coho salmon habitat is used to identify the policy standards in stream reaches where flow, valley constraint, and channel gradient are appropriate for coho salmon habitat. A second model of the stream network (hydrography) is developed to evaluate how differences in stream delineation can influence policy patterns. Results indicate that riparian lands of the Oregon Coast Range are protected by variable standards for land management, including buffer widths that range from 0 to ~152m. In streams important to coho salmon, a lower proportion of the stream network was found to be protected by policy standards compared to the watershed and ESU scale. A lack of stream data central to policy application (fish use, stream flow duration, mean annual flow, water use) complicated our ability to attribute stream segments with specific land management requirements. The choice of hydrographic data was also found to influence the proportional extent of policies within riverscapes. This framework for assessing the spatial distribution of policies can provide insights for future studies of riverscape systems. Title: Policy Patterns across Riverscapes : Riparian Land Standards in the Oregon Coast Range Author: Boisjolie, Brett A. Land management policies are ideas about nature projected onto the landscape. Culminations of social, economic, and scientific influences, these policies create standards affecting the function of ecological systems. In the case of riparian lands in the Oregon Coast Range, policy requirements vary considerably across federal, state, and private land ownerships. Protective measures, such as the adoption of fixed buffer widths for maintaining vegetation, are intended to preserve natural processes important to instream habitat and water quality. However, the "policy landscape" of variable management standards can result in a fragmented approach to policy protection. This fragmentation complicates recovery efforts for threatened anadromous fish species such as coho salmon (Oncorhynchus kisutch), which require connectivity in suitable habitat segments throughout the river network. Taking a riverscape perspective, this study evaluates how aquatic protections vary across the river and its valley. To explore the patterns created from variable standard, the study first delineates categories of riparian management standards in the Oregon Coast Range. Using a geographic information system (GIS), these standards are mapped across the river networks of this region. The extent of each policy category is quantified in stream kilometers in order to evaluate policy efforts within and across the riverscapes of the Oregon Coast Range. A model of intrinsic potential for coho salmon habitat is used to identify the policy standards in stream reaches where flow, valley constraint, and channel gradient are appropriate for coho salmon habitat. A second model of the stream network (hydrography) is developed to evaluate how differences in stream delineation can influence policy patterns. Results indicate that riparian lands of the Oregon Coast Range are protected by variable standards for land management, including buffer widths that range from 0 to ~152m. In streams important to coho salmon, a lower proportion of the stream network was found to be protected by policy standards compared to the watershed and ESU scale. A lack of stream data central to policy application (fish use, stream flow duration, mean annual flow, water use) complicated our ability to attribute stream segments with specific land management requirements. The choice of hydrographic data was also found to influence the proportional extent of policies within riverscapes. This framework for assessing the spatial distribution of policies can provide insights for future studies of riverscape systems. Close

• 34. [Article] Carbon storage in a Pacific Northwest conifer forest ecosystem : a chronosequence approach

  As concern over global warming intensifies, sequestration and storage of atmospheric CO2 has become an important scientific and policy issue. Confusion persists, however, over interpretation of forest ...

  Citation × Citation Citation Abstract Copy Title: Carbon storage in a Pacific Northwest conifer forest ecosystem : a chronosequence approach Author: Janisch, Jack E. As concern over global warming intensifies, sequestration and storage of atmospheric CO2 has become an important scientific and policy issue. Confusion persists, however, over interpretation of forest carbon (C) source-sink dynamics, in part because conclusions drawn depend on temporal and spatial scales of analysis (e.g. day-week scale vs. successional-scale), type of disturbance, and methodology (e.g. massbased vs. flux-based). There is a need to resolve this confusion given that strategies for mitigating anthropogenic CO2 emissions are based on estimates of forest C fluxes during various stages of succession, over which C fluxes and stores may change. Empirical study of changes in forest C stores can help to resolve this confusion by clarifying the C sources-sink dynamics of forests in space and time. To better understand the impacts of disturbance on C source-sink dynamics, changes in C stores of an evergreen-dominated forest on the Wind River Ranger District in Southwestern Washington, U.S.A., were investigated along a 500-year chronosequence of 36 stands. Principle objectives were to evaluate 1) decomposition rates (k) of logs, stumps, and below-ground coarse roots, 2) net primary productivity (NPP) of dominant tree species' boles at the stand level, and 3) successional changes in net ecosystem productivity (NEP) for live trees and coarse woody debris (CWD), here called NEPW. In the case of decomposition, log and stump k values did not differ significantly within the two principle species studied, indicating substitution of log k values for stump k values in models of forest C budgets may be valid when stump decomposition data is lacking. Decomposition rates between species differed, with Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) decomposing more slowly (k = 0.013 yr-1) relative to western hemlock (Tsuga heterophylla (Rafi) Sarg. (k = 0.036 yr-1). This difference in k between species was observed for both above-ground stumps and logs as well as below-ground coarse roots. Given our mean k estimates and adjusting for regenerating stand age, these stands are losing C at a rate of 0.16-0.83 Mg C ha-1 yr-1 (assuming all CWD is P. menziesii) to 0.13-1.68 Mg C ha-1 yr-1 (assuming all CWD is T. heterophylla) from stumps, logs, and snags. Including coarse roots increases these losses to 0.28-1.25 Mg C ha-1 yr-1 and 0.30-2.53 Mg C ha-1 yr-1, respectively. Based on these findings, if fragmentation of these decomposing C pools is ignored, and fragmented fractions have oxidized to CO2, stands thought to be net C sinks could in reality be net C sources to the atmosphere. Net primary production in tree boles (NPPb) of regenerating stands (so called second-growth) ranged between 0.15-5.28 Mg C ha-1 yr-1. NPPb of 500-year old stands ranged between 1.3-3.9 Mg C ha-1 yr-1, similar to NPPb of boles in 20-25 year old secondgrowth. Mean radial increment widths from old-growth stands indicated that NPPb of these stands (neglecting mortality) can increase, decrease, or remain relatively constant. Based on 5-year increments for the previous fifteen years, the majority of old-growth stands sampled showed small increases in radial growth over time. Timing of the transition from negative to positive of NEPW ranged between 0 and 57 years after disturbance and depended strongly on live-tree growth rates as well as the fate of CWD and harvested wood. Estimated maximum and minimum NEPW were 3.9 Mg C ha-1 yr-1 and 14.1 Mg C ha-1 yr-1, respectively. Maximum mean C stores of 393 Mg C ha-1 were reached approximately 200 years after disturbance. At a rotation age of 80 years, regenerating stands stored approximately 50% as much C in woody biomass as a 500-year old primary forest, indicating conversion of older forests to plantations released C to the atmosphere. Given the high biomass of mature and old-growth stands relative to younger regenerating stands in the forest studied, landscape C stores in live wood would appear to be maximized in stands of older age classes. Title: Carbon storage in a Pacific Northwest conifer forest ecosystem : a chronosequence approach Author: Janisch, Jack E. As concern over global warming intensifies, sequestration and storage of atmospheric CO2 has become an important scientific and policy issue. Confusion persists, however, over interpretation of forest carbon (C) source-sink dynamics, in part because conclusions drawn depend on temporal and spatial scales of analysis (e.g. day-week scale vs. successional-scale), type of disturbance, and methodology (e.g. massbased vs. flux-based). There is a need to resolve this confusion given that strategies for mitigating anthropogenic CO2 emissions are based on estimates of forest C fluxes during various stages of succession, over which C fluxes and stores may change. Empirical study of changes in forest C stores can help to resolve this confusion by clarifying the C sources-sink dynamics of forests in space and time. To better understand the impacts of disturbance on C source-sink dynamics, changes in C stores of an evergreen-dominated forest on the Wind River Ranger District in Southwestern Washington, U.S.A., were investigated along a 500-year chronosequence of 36 stands. Principle objectives were to evaluate 1) decomposition rates (k) of logs, stumps, and below-ground coarse roots, 2) net primary productivity (NPP) of dominant tree species' boles at the stand level, and 3) successional changes in net ecosystem productivity (NEP) for live trees and coarse woody debris (CWD), here called NEPW. In the case of decomposition, log and stump k values did not differ significantly within the two principle species studied, indicating substitution of log k values for stump k values in models of forest C budgets may be valid when stump decomposition data is lacking. Decomposition rates between species differed, with Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) decomposing more slowly (k = 0.013 yr-1) relative to western hemlock (Tsuga heterophylla (Rafi) Sarg. (k = 0.036 yr-1). This difference in k between species was observed for both above-ground stumps and logs as well as below-ground coarse roots. Given our mean k estimates and adjusting for regenerating stand age, these stands are losing C at a rate of 0.16-0.83 Mg C ha-1 yr-1 (assuming all CWD is P. menziesii) to 0.13-1.68 Mg C ha-1 yr-1 (assuming all CWD is T. heterophylla) from stumps, logs, and snags. Including coarse roots increases these losses to 0.28-1.25 Mg C ha-1 yr-1 and 0.30-2.53 Mg C ha-1 yr-1, respectively. Based on these findings, if fragmentation of these decomposing C pools is ignored, and fragmented fractions have oxidized to CO2, stands thought to be net C sinks could in reality be net C sources to the atmosphere. Net primary production in tree boles (NPPb) of regenerating stands (so called second-growth) ranged between 0.15-5.28 Mg C ha-1 yr-1. NPPb of 500-year old stands ranged between 1.3-3.9 Mg C ha-1 yr-1, similar to NPPb of boles in 20-25 year old secondgrowth. Mean radial increment widths from old-growth stands indicated that NPPb of these stands (neglecting mortality) can increase, decrease, or remain relatively constant. Based on 5-year increments for the previous fifteen years, the majority of old-growth stands sampled showed small increases in radial growth over time. Timing of the transition from negative to positive of NEPW ranged between 0 and 57 years after disturbance and depended strongly on live-tree growth rates as well as the fate of CWD and harvested wood. Estimated maximum and minimum NEPW were 3.9 Mg C ha-1 yr-1 and 14.1 Mg C ha-1 yr-1, respectively. Maximum mean C stores of 393 Mg C ha-1 were reached approximately 200 years after disturbance. At a rotation age of 80 years, regenerating stands stored approximately 50% as much C in woody biomass as a 500-year old primary forest, indicating conversion of older forests to plantations released C to the atmosphere. Given the high biomass of mature and old-growth stands relative to younger regenerating stands in the forest studied, landscape C stores in live wood would appear to be maximized in stands of older age classes. Close

• 35. [Article] Drought, dispersal, and community dynamics in arid-land streams

  Understanding the mechanisms that regulate local species diversity and community structure is a perennial goal of ecology. Local community structure can be viewed as the result of numerous local and regional ...

  Citation × Citation Citation Abstract Copy Title: Drought, dispersal, and community dynamics in arid-land streams Author: Bogan, Michael T. Understanding the mechanisms that regulate local species diversity and community structure is a perennial goal of ecology. Local community structure can be viewed as the result of numerous local and regional processes; these processes act as filters that reduce the regional species pool down to the observed local community. In stream ecosystems, the natural flow regime (including the timing, magnitude, and duration of high and low flow events) is widely recognized as a primary regulator of local diversity and community composition. This is especially true in aridland streams, where low- and zero-flow events can occur frequently and for extended periods of time (months to years). Additionally, wetted habitat patches in arid-land stream networks are often fragmented within and among stream networks. Thus dispersal between isolated aquatic patches may also play a large role in regulating local communities. In my dissertation, I explored the roles that drought, dispersal, and local habitat factors play in structuring arid-land stream communities. I examined the impact of flow permanence and seasonal variation in flow and other abiotic factors on aquatic communities at both fine spatial scales over a long time period (8 years; Chapter 2) and at a broad spatial scale over a shorter time period (1-2 years; Chapter 4). Additionally, I quantified aquatic invertebrate aerial dispersal over moderate spatial scales (≤ 0.5 km) by conducting a colonization experiment using artificial stream pools placed along and inland from two arid-land streams (Chapter 4). Finally, I examined the roles of spatial isolation, microhabitat type, and local abiotic and biotic factors in structuring aquatic communities in freshwater oases scattered across one of the most arid regions of North America, the southern Sonoran Desert (Chapter 5). In Chapter 2, I found that severe drought caused an unprecedented drying event in isolated perennial stream pools, and that several additional drying events occurred over the following four years. This transition to intermittent flow caused the extirpation of several large, long-lived species with low dispersal abilities (including the top predator) and drove the local community into an alternative state. In the colonization experiment described in Chapter 3, I found that several arid-land stream invertebrate taxa disperse widely and frequently. The widespread dispersers identified by this experiment included several of the earliest colonist taxa observed following the severe drought described in Chapter 2. Other taxa, though, only dispersed overland after receiving an environmental cue (rainfall) or preferentially dispersed along stream corridors. In Chapter 4, where I examined invertebrate community structure across a large network of well-connected intermittent and perennial reaches, I found low diversity in intermittent reaches, regardless of their connectivity to diverse upstream perennial reaches. These species-poor, intermittent communities were composed of a unique suite of species with lifehistory adaptations that conferred desiccation resistance, including extended egg and larval diapause stages. The short flow duration of intermittent reaches (<100 days) likely precluded upstream perennial taxa from establishing populations in downstream intermittent reaches before drying occurred, while the relative predictability of flow timing (Dec-Apr) likely allowed for a small number of species to develop appropriate life-history traits (e.g., diapause stage, rapid development time) to exploit these temporally-fleeting habitats. In Chapter 5, I found over 220 species of aquatic animals (including ≥ 5 undescribed species) in the 19 desert oases that were sampled across the southern Sonoran Desert. Local community composition in these oases was strongly driven by microhabitat type. Additionally, native aquatic species richness and abundance in these oases were significantly reduced by the introduction of tilapia, an exotic fish species. The threats to arid-land streams presented by increased drought severity, anthropogenic water withdrawals, and local habitat degradation (e.g., introduced species, unmanaged recreational use) are grave across the southwestern US and northwestern Mexico. I hope that in addition to furthering our understanding of ecological processes in arid-land streams, this dissertation makes a small contribution towards the efforts to preserve these habitats. Title: Drought, dispersal, and community dynamics in arid-land streams Author: Bogan, Michael T. Understanding the mechanisms that regulate local species diversity and community structure is a perennial goal of ecology. Local community structure can be viewed as the result of numerous local and regional processes; these processes act as filters that reduce the regional species pool down to the observed local community. In stream ecosystems, the natural flow regime (including the timing, magnitude, and duration of high and low flow events) is widely recognized as a primary regulator of local diversity and community composition. This is especially true in aridland streams, where low- and zero-flow events can occur frequently and for extended periods of time (months to years). Additionally, wetted habitat patches in arid-land stream networks are often fragmented within and among stream networks. Thus dispersal between isolated aquatic patches may also play a large role in regulating local communities. In my dissertation, I explored the roles that drought, dispersal, and local habitat factors play in structuring arid-land stream communities. I examined the impact of flow permanence and seasonal variation in flow and other abiotic factors on aquatic communities at both fine spatial scales over a long time period (8 years; Chapter 2) and at a broad spatial scale over a shorter time period (1-2 years; Chapter 4). Additionally, I quantified aquatic invertebrate aerial dispersal over moderate spatial scales (≤ 0.5 km) by conducting a colonization experiment using artificial stream pools placed along and inland from two arid-land streams (Chapter 4). Finally, I examined the roles of spatial isolation, microhabitat type, and local abiotic and biotic factors in structuring aquatic communities in freshwater oases scattered across one of the most arid regions of North America, the southern Sonoran Desert (Chapter 5). In Chapter 2, I found that severe drought caused an unprecedented drying event in isolated perennial stream pools, and that several additional drying events occurred over the following four years. This transition to intermittent flow caused the extirpation of several large, long-lived species with low dispersal abilities (including the top predator) and drove the local community into an alternative state. In the colonization experiment described in Chapter 3, I found that several arid-land stream invertebrate taxa disperse widely and frequently. The widespread dispersers identified by this experiment included several of the earliest colonist taxa observed following the severe drought described in Chapter 2. Other taxa, though, only dispersed overland after receiving an environmental cue (rainfall) or preferentially dispersed along stream corridors. In Chapter 4, where I examined invertebrate community structure across a large network of well-connected intermittent and perennial reaches, I found low diversity in intermittent reaches, regardless of their connectivity to diverse upstream perennial reaches. These species-poor, intermittent communities were composed of a unique suite of species with lifehistory adaptations that conferred desiccation resistance, including extended egg and larval diapause stages. The short flow duration of intermittent reaches (<100 days) likely precluded upstream perennial taxa from establishing populations in downstream intermittent reaches before drying occurred, while the relative predictability of flow timing (Dec-Apr) likely allowed for a small number of species to develop appropriate life-history traits (e.g., diapause stage, rapid development time) to exploit these temporally-fleeting habitats. In Chapter 5, I found over 220 species of aquatic animals (including ≥ 5 undescribed species) in the 19 desert oases that were sampled across the southern Sonoran Desert. Local community composition in these oases was strongly driven by microhabitat type. Additionally, native aquatic species richness and abundance in these oases were significantly reduced by the introduction of tilapia, an exotic fish species. The threats to arid-land streams presented by increased drought severity, anthropogenic water withdrawals, and local habitat degradation (e.g., introduced species, unmanaged recreational use) are grave across the southwestern US and northwestern Mexico. I hope that in addition to furthering our understanding of ecological processes in arid-land streams, this dissertation makes a small contribution towards the efforts to preserve these habitats. Close

• 36. [Image] Botrychium summit: 16 March 1993

  	Cover title; Includes draft: Species conservation strategy: pumice grape fern, 1992, botrychium pumicola cov. in underw, Deschutes National Forest sensitive plant program
  	Citation × Citation Citation Abstract Copy Title: Botrychium summit: 16 March 1993 Author: Christine O. Hopkins Year: 1940, 2009, 2010 Cover title; Includes draft: Species conservation strategy: pumice grape fern, 1992, botrychium pumicola cov. in underw, Deschutes National Forest sensitive plant program Title: Botrychium summit: 16 March 1993 Author: Christine O. Hopkins Year: 1940, 2009, 2010 Cover title; Includes draft: Species conservation strategy: pumice grape fern, 1992, botrychium pumicola cov. in underw, Deschutes National Forest sensitive plant program Close

• 37. [Article] Terrestrial amphibian distribution, habitat associations and downed wood temperature profiles in managed headwater forests with riparian buffers in the Oregon Coast Range

  Western forests have become increasingly fragmented landscapes dominated by young stands. Given that western Oregon forests largely consist of headwater systems, there is a need to better understand how ...

  Citation × Citation Citation Abstract Copy Title: Terrestrial amphibian distribution, habitat associations and downed wood temperature profiles in managed headwater forests with riparian buffers in the Oregon Coast Range Author: Kluber, Matthew R. Western forests have become increasingly fragmented landscapes dominated by young stands. Given that western Oregon forests largely consist of headwater systems, there is a need to better understand how headwater forest taxa and their habitats are impacted by forest management practices. Several amphibian species associated with forested headwater systems have emerged as management concerns. Forest management strategies, such as harvests that remove only part of the canopy and retention of riparian buffer strips, may help ameliorate some of the negative effects on amphibians in managed forests. Pre-existing site conditions, such as legacy downed wood, also may play a role in buffering the impacts of silvicultural practices on terrestrial amphibians. Downed wood is an important habitat component for many amphibians, because the cool, moist microclimates of downed wood can provide refugia for terrestrial amphibians during warmer summer months. However, downed wood habitat suitability is another emerging concern as the rate of input and size of downed wood declines in managed forests. As part of the USDI Bureau of Land Managements Density Management Study, we investigated how untreated streamside buffers modify impacts of upland thinning on headwater forest terrestrial amphibians and their habitat at three sites in the Oregon Coast Range. To further assess habitat associations of these animals, we conducted a field experiment to address amphibian cover use, including downed wood, moss and coarse and fine substrates. In addition, we examined how temperature profiles inside small- and large-diameter downed wood and soil temperatures differed from ambient air temperatures. Temperatures of wood and soil were monitored at different slope positions (near streams and upslope) and overstory regimes (thinned and unthinned stands) to assess potential habitat suitability and buffering capabilities against seasonal temperature extremes for plethodontid salamanders. Our results suggest that pre-existing site conditions (e.g., amount of rocky or fine substrate) play an important role in determining the response of terrestrial amphibians to upland forest thinning. However, retention of stream buffers is important in maintaining unaltered stream and riparian conditions. Moderate thinning and preservation of vital habitat in riparian and nearby upland areas by way of variable-width buffers (15 m minimum width) may be sufficient in maintaining suitable habitat and microclimatic conditions vital to amphibian assemblages in managed headwater forests. Additionally, logs of a wide size range and soils may provide sufficient protection against thermal extremes harmful to plethodontid salamanders in thinned stands with limited overstory. However, this alone cannot support plethodontid salamanders. These salamanders require exposed areas (e.g., leaf litter, soil surface, rock faces) where much of their foraging and well as courtship occurs. Partial retention of the canopy through moderate thinning coupled with variable-width riparain buffers that may increase in width when suitable terrestrial habitat is encountered, may provide sufficient microhabitat, microclimate, and protection in maintaining terrestrial amphibian assemblages in managed headwater forests. Title: Terrestrial amphibian distribution, habitat associations and downed wood temperature profiles in managed headwater forests with riparian buffers in the Oregon Coast Range Author: Kluber, Matthew R. Western forests have become increasingly fragmented landscapes dominated by young stands. Given that western Oregon forests largely consist of headwater systems, there is a need to better understand how headwater forest taxa and their habitats are impacted by forest management practices. Several amphibian species associated with forested headwater systems have emerged as management concerns. Forest management strategies, such as harvests that remove only part of the canopy and retention of riparian buffer strips, may help ameliorate some of the negative effects on amphibians in managed forests. Pre-existing site conditions, such as legacy downed wood, also may play a role in buffering the impacts of silvicultural practices on terrestrial amphibians. Downed wood is an important habitat component for many amphibians, because the cool, moist microclimates of downed wood can provide refugia for terrestrial amphibians during warmer summer months. However, downed wood habitat suitability is another emerging concern as the rate of input and size of downed wood declines in managed forests. As part of the USDI Bureau of Land Managements Density Management Study, we investigated how untreated streamside buffers modify impacts of upland thinning on headwater forest terrestrial amphibians and their habitat at three sites in the Oregon Coast Range. To further assess habitat associations of these animals, we conducted a field experiment to address amphibian cover use, including downed wood, moss and coarse and fine substrates. In addition, we examined how temperature profiles inside small- and large-diameter downed wood and soil temperatures differed from ambient air temperatures. Temperatures of wood and soil were monitored at different slope positions (near streams and upslope) and overstory regimes (thinned and unthinned stands) to assess potential habitat suitability and buffering capabilities against seasonal temperature extremes for plethodontid salamanders. Our results suggest that pre-existing site conditions (e.g., amount of rocky or fine substrate) play an important role in determining the response of terrestrial amphibians to upland forest thinning. However, retention of stream buffers is important in maintaining unaltered stream and riparian conditions. Moderate thinning and preservation of vital habitat in riparian and nearby upland areas by way of variable-width buffers (15 m minimum width) may be sufficient in maintaining suitable habitat and microclimatic conditions vital to amphibian assemblages in managed headwater forests. Additionally, logs of a wide size range and soils may provide sufficient protection against thermal extremes harmful to plethodontid salamanders in thinned stands with limited overstory. However, this alone cannot support plethodontid salamanders. These salamanders require exposed areas (e.g., leaf litter, soil surface, rock faces) where much of their foraging and well as courtship occurs. Partial retention of the canopy through moderate thinning coupled with variable-width riparain buffers that may increase in width when suitable terrestrial habitat is encountered, may provide sufficient microhabitat, microclimate, and protection in maintaining terrestrial amphibian assemblages in managed headwater forests. Close

• 38. [Image] An assessment of ecosystem components in the interior Columbia Basin and portions of the Klamath and Great Basins [volume 1]

  	Quigley, Thomas M.; Arbelbide, Sylvia J., tech. eds. 1997. An assessment of ecosystem components in the interior Columbia basin and portions of the Klamath and Great Basins: volume 1. Gen. Tech. Rep. PNW-GTR-405. ...
  	Citation × Citation Citation Abstract Copy Title: An assessment of ecosystem components in the interior Columbia Basin and portions of the Klamath and Great Basins [volume 1] Author: Quigley, Thomas Milton; Arbelbide, S. J. (Sylvia J.) Year: 1997, 2008, 2005 Quigley, Thomas M.; Arbelbide, Sylvia J., tech. eds. 1997. An assessment of ecosystem components in the interior Columbia basin and portions of the Klamath and Great Basins: volume 1. Gen. Tech. Rep. PNW-GTR-405. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 4 vol. (Quigley, Thomas M., tech. ed.; The Interior Columbia Basin Ecosystem Management Project: Scientific Assessment). The Assessment of Ecosystem Components in the Interior Columbia Basin and Portions of the Klamath and Great Basins provides detailed information about current conditions and trends for the biophysical and social systems within the Basin. This information can be used by land managers to develop broad land management goals and priorities and provides the context for decisions specific to smaller geographic areas. The Assessment area covers about 8 percent of the U.S. land area, 24 percent of the Nation's National Forest System lands, 10 percent of the Nation's BLM-administered lands, and contains about 1.2 percent of the Nation's population. This results in a population density that is less than one-sixth of the U.S. average. The area has experienced recent, rapid population growth and generally has a robust, diverse economy. As compared to historic conditions, the terrestrial, aquatic, forest, and rangeland systems have undergone dramatic changes. Forested landscapes are more susceptible to fire, insect, and disease than under historic conditions. Rangelands are highly susceptible to noxious weed invasion. The disturbance regimes that operate on forest and rangeland have changed substantially, with lethal fires dominating many areas where non-lethal fires were the norm historically. Terrestrial habitats that have experienced the greatest decline include the native grassland, native shrubland, and old forest structures. There are areas within the Assessment area that have higher diversity than others. Aquatic systems are now more fragmented and isolated than historically and the introduction of non-native fish species has complicated current status of native fishes. Core habitat and population centers do remain as building blocks for restoration. Social and economic conditions within the Assessment area vary considerably, depending to a great extent on population, diversity of employment opportunities, and changing demographics. Those counties with the higher population densities and greater diversity of employment opportunities are generally more resilient to economic downturns. This Assessment provides a rich information base, including over 170 mapped themes with associated models and databases, from which future decisions can benefit. Keywords: Columbia basin, biophysical systems, social systems, ecosystem. Title: An assessment of ecosystem components in the interior Columbia Basin and portions of the Klamath and Great Basins [volume 1] Author: Quigley, Thomas Milton; Arbelbide, S. J. (Sylvia J.) Year: 1997, 2008, 2005 Quigley, Thomas M.; Arbelbide, Sylvia J., tech. eds. 1997. An assessment of ecosystem components in the interior Columbia basin and portions of the Klamath and Great Basins: volume 1. Gen. Tech. Rep. PNW-GTR-405. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 4 vol. (Quigley, Thomas M., tech. ed.; The Interior Columbia Basin Ecosystem Management Project: Scientific Assessment). The Assessment of Ecosystem Components in the Interior Columbia Basin and Portions of the Klamath and Great Basins provides detailed information about current conditions and trends for the biophysical and social systems within the Basin. This information can be used by land managers to develop broad land management goals and priorities and provides the context for decisions specific to smaller geographic areas. The Assessment area covers about 8 percent of the U.S. land area, 24 percent of the Nation's National Forest System lands, 10 percent of the Nation's BLM-administered lands, and contains about 1.2 percent of the Nation's population. This results in a population density that is less than one-sixth of the U.S. average. The area has experienced recent, rapid population growth and generally has a robust, diverse economy. As compared to historic conditions, the terrestrial, aquatic, forest, and rangeland systems have undergone dramatic changes. Forested landscapes are more susceptible to fire, insect, and disease than under historic conditions. Rangelands are highly susceptible to noxious weed invasion. The disturbance regimes that operate on forest and rangeland have changed substantially, with lethal fires dominating many areas where non-lethal fires were the norm historically. Terrestrial habitats that have experienced the greatest decline include the native grassland, native shrubland, and old forest structures. There are areas within the Assessment area that have higher diversity than others. Aquatic systems are now more fragmented and isolated than historically and the introduction of non-native fish species has complicated current status of native fishes. Core habitat and population centers do remain as building blocks for restoration. Social and economic conditions within the Assessment area vary considerably, depending to a great extent on population, diversity of employment opportunities, and changing demographics. Those counties with the higher population densities and greater diversity of employment opportunities are generally more resilient to economic downturns. This Assessment provides a rich information base, including over 170 mapped themes with associated models and databases, from which future decisions can benefit. Keywords: Columbia basin, biophysical systems, social systems, ecosystem. Close

• 39. [Article] Road Ecology Course and Seminar Series

  The American road network is a major economic investment that is a major organizing force for human activity. The road system has profoundly altered ecological processes and, as a result, it is also an ...

  Citation × Citation Citation Abstract Copy Title: Road Ecology Course and Seminar Series Author: Sytsma, Mark D. Year: 2008 The American road network is a major economic investment that is a major organizing force for human activity. The road system has profoundly altered ecological processes and, as a result, it is also an important organizing force for ecosystems. Understanding the ecological consequences of road system design and use is critical to effective engineering and management of road systems to minimize impacts. I propose a new course within Environmental Science and Resources at PSU that will expose upper division undergraduate and graduate students to the fundamental concepts of road ecology through discussion, guest speakers, and field trips. I propose a student-led discussion course focusing on the book Road Ecology, edited by Forman et al. This book provides an introduction to the basic concepts underlying road impacts on the environment. Students will be required to read the book; discussion of each chapter will be led by a student who will make a presentation on results of a search of the primary literature on the subject of each chapter. The classroom discussion will be supplemented with guest speakers that will bring real- world examples to the classroom. Potential topics for guest speakers include: impacts of highway runoff and roadside spraying on water quality, impacts of roads on erosional processes, impacts of culverts on anadromous fish passage, mitigation of habitat fragmentation and road impacts on animal movements, and role of maintenance activities in facilitating weed invasions. Invited speakers will also be requested to provide a departmental seminar to communicate road ecology principles to a wider audience of undergraduate and graduate students in Environmental Science and Resources. Field trips to observe road impacts and mitigation measures will add to students understanding of road ecology. We will schedule field trips to observe impacts as well as efforts to mitigate environmental effects of roads. Students performance will be evaluated by participation in class discussions and field trips, and a term paper. Undergraduates will be required to write a literature review on an aspect of road ecology of their choosing. Graduate students will be required to write a proposal to investigate some aspect of road ecology. Following completion of the course students will have a broad understanding of the environmental impacts of road systems on ecological processes and methods of mitigating impacts. Graduate students will have a research proposal that may be appropriate for submittal to OTREC for funding. Title: Road Ecology Course and Seminar Series Author: Sytsma, Mark D. Year: 2008 The American road network is a major economic investment that is a major organizing force for human activity. The road system has profoundly altered ecological processes and, as a result, it is also an important organizing force for ecosystems. Understanding the ecological consequences of road system design and use is critical to effective engineering and management of road systems to minimize impacts. I propose a new course within Environmental Science and Resources at PSU that will expose upper division undergraduate and graduate students to the fundamental concepts of road ecology through discussion, guest speakers, and field trips. I propose a student-led discussion course focusing on the book Road Ecology, edited by Forman et al. This book provides an introduction to the basic concepts underlying road impacts on the environment. Students will be required to read the book; discussion of each chapter will be led by a student who will make a presentation on results of a search of the primary literature on the subject of each chapter. The classroom discussion will be supplemented with guest speakers that will bring real- world examples to the classroom. Potential topics for guest speakers include: impacts of highway runoff and roadside spraying on water quality, impacts of roads on erosional processes, impacts of culverts on anadromous fish passage, mitigation of habitat fragmentation and road impacts on animal movements, and role of maintenance activities in facilitating weed invasions. Invited speakers will also be requested to provide a departmental seminar to communicate road ecology principles to a wider audience of undergraduate and graduate students in Environmental Science and Resources. Field trips to observe road impacts and mitigation measures will add to students understanding of road ecology. We will schedule field trips to observe impacts as well as efforts to mitigate environmental effects of roads. Students performance will be evaluated by participation in class discussions and field trips, and a term paper. Undergraduates will be required to write a literature review on an aspect of road ecology of their choosing. Graduate students will be required to write a proposal to investigate some aspect of road ecology. Following completion of the course students will have a broad understanding of the environmental impacts of road systems on ecological processes and methods of mitigating impacts. Graduate students will have a research proposal that may be appropriate for submittal to OTREC for funding. Close

• 40. [Article] Carbon Sequestration in the New Jersey Pine Barrens Under Different Scenarios of Fire Management

  The New Jersey Pine Barrens (NJPB) is the largest forested area along the northeastern coast of the United States. The NJPB are dominated by pine (Pinus spp.) and oak (Quercus spp.) stands that are fragmented ...

  Citation × Citation Citation Abstract Copy Title: Carbon Sequestration in the New Jersey Pine Barrens Under Different Scenarios of Fire Management Author: Scheller, Robert M., Van Tuyl, Steve, Clark, Kenneth L., Hom, John, La Puma, Inga Year: 2011 The New Jersey Pine Barrens (NJPB) is the largest forested area along the northeastern coast of the United States. The NJPB are dominated by pine (Pinus spp.) and oak (Quercus spp.) stands that are fragmented and subject to frequent disturbance and forest management. Over long time periods (>50 years), the balance between oak and pine dominance is determined by fire frequency. As a consequence, the ability of the NJPB to sequester carbon may be contingent upon management activities as well as patterns of historic land use. We simulated 100 years of carbon change using three scenarios: (1) contemporary management as reflected in the recent (1991–2006) fire records, (2) an increase in the fire ignitions within the wildland urban interface areas of the NJPB reflecting increased prescribed fires, and (3) a longer response time to wildfires, reflecting a more liberal burning policy by the New Jersey Forest Fire Service. We used the LANDIS-II model coupled with CENTURY and the Dynamic Fire and Dynamic Biomass Fuels extensions to estimate forest carbon sequestration based on these three scenarios. Calibration and validation via comparison to monthly flux tower data indicated that the model reasonably captured the timing and magnitude of net ecosystem exchange in the absence of Gypsy moth defoliation (r2 = 0.89). Under all scenarios, our simulations suggest that forests of the NJPB will continue to accumulate carbon over the next 100 years under current climatic conditions. Although above ground net primary productivity, live carbon, and detrital carbon were roughly constant or increased only modestly, soil organic carbon continued to increase through time for all forest types except the highly xeric pine plains. Our simulated changes in management reflected only minor alterations to the fire regime and thus management may have only minor effects on total forest carbon budgets in the immediate future particularly when compared to recovery from historic disturbance patterns. Title: Carbon Sequestration in the New Jersey Pine Barrens Under Different Scenarios of Fire Management Author: Scheller, Robert M., Van Tuyl, Steve, Clark, Kenneth L., Hom, John, La Puma, Inga Year: 2011 The New Jersey Pine Barrens (NJPB) is the largest forested area along the northeastern coast of the United States. The NJPB are dominated by pine (Pinus spp.) and oak (Quercus spp.) stands that are fragmented and subject to frequent disturbance and forest management. Over long time periods (>50 years), the balance between oak and pine dominance is determined by fire frequency. As a consequence, the ability of the NJPB to sequester carbon may be contingent upon management activities as well as patterns of historic land use. We simulated 100 years of carbon change using three scenarios: (1) contemporary management as reflected in the recent (1991–2006) fire records, (2) an increase in the fire ignitions within the wildland urban interface areas of the NJPB reflecting increased prescribed fires, and (3) a longer response time to wildfires, reflecting a more liberal burning policy by the New Jersey Forest Fire Service. We used the LANDIS-II model coupled with CENTURY and the Dynamic Fire and Dynamic Biomass Fuels extensions to estimate forest carbon sequestration based on these three scenarios. Calibration and validation via comparison to monthly flux tower data indicated that the model reasonably captured the timing and magnitude of net ecosystem exchange in the absence of Gypsy moth defoliation (r2 = 0.89). Under all scenarios, our simulations suggest that forests of the NJPB will continue to accumulate carbon over the next 100 years under current climatic conditions. Although above ground net primary productivity, live carbon, and detrital carbon were roughly constant or increased only modestly, soil organic carbon continued to increase through time for all forest types except the highly xeric pine plains. Our simulated changes in management reflected only minor alterations to the fire regime and thus management may have only minor effects on total forest carbon budgets in the immediate future particularly when compared to recovery from historic disturbance patterns. Close

• 41. [Article] Creating Useful and Usable Climate Tools for Sagebrush Land Management Through Scientist and Manager Collaboration

  The sagebrush ecosystem, home to numerous plant and animal species including big sagebrush (Artemisia tridentata) and the endemic greater sage-grouse (Centrocercus urophasianus), has endured fragmentation ...

  Citation × Citation Citation Abstract Copy Title: Creating Useful and Usable Climate Tools for Sagebrush Land Management Through Scientist and Manager Collaboration Author: Brown, Melanie The sagebrush ecosystem, home to numerous plant and animal species including big sagebrush (Artemisia tridentata) and the endemic greater sage-grouse (Centrocercus urophasianus), has endured fragmentation and degradation of both quantity and quality due to the cumulative and synergistic relationships between an abundance of individual disturbances including grazing, invasive annuals and fire. Climate change may now be an additional threat that poses the greatest risk to these imperiled habitats. Natural resource agencies such as the Bureau of Land Management (BLM) seek to conserve sagelands through land management activities that ensure the survival of sage-grouse and continuity of the sagebrush biome. Web-based climate tools can help convey climate information that may be necessary for long-term land management, but these tools may not agree with the needs of land managers, may be too complex, or may be misinterpreted. To overcome barriers of user compatibility, the participation of both climate scientists and land managers is necessary during tool development. With the collaboration of Oregon and Idaho BLM sagebrush land managers and climate scientists, this study sought to assess land manager needs and define the criteria for useful and useable climate tools. Using an initial online survey, individual phone interviews with land managers, and a follow-up online survey, a series of land management activities and related climate variables were identified, and several web-based climate tools were assessed. Most managers perform vegetation management through a variety of means including seeding and herbicide application. Such activities are affected by the magnitude and timing of precipitation and temperature, as well as other variables, on seasonal and annual timeframes. For planning purposes land managers also need information on long-term 10-20 year climate trends. The act of listening to the needs of land managers uncovered communication barriers, and provided feedback on existing climate tools emphasizing accessibility, dependability and consistency, clear explanation of terminology, effective visualizations, and relevant spatial and temporal scales to the scope of management activities. We also identified a need for basic information and education on the location of existing climate tools and climate impacts, and a need for near-term forecasting tools that could bridge the gap between weather (≤ 6 months) and climate (≥ 30 years) projections. Title: Creating Useful and Usable Climate Tools for Sagebrush Land Management Through Scientist and Manager Collaboration Author: Brown, Melanie The sagebrush ecosystem, home to numerous plant and animal species including big sagebrush (Artemisia tridentata) and the endemic greater sage-grouse (Centrocercus urophasianus), has endured fragmentation and degradation of both quantity and quality due to the cumulative and synergistic relationships between an abundance of individual disturbances including grazing, invasive annuals and fire. Climate change may now be an additional threat that poses the greatest risk to these imperiled habitats. Natural resource agencies such as the Bureau of Land Management (BLM) seek to conserve sagelands through land management activities that ensure the survival of sage-grouse and continuity of the sagebrush biome. Web-based climate tools can help convey climate information that may be necessary for long-term land management, but these tools may not agree with the needs of land managers, may be too complex, or may be misinterpreted. To overcome barriers of user compatibility, the participation of both climate scientists and land managers is necessary during tool development. With the collaboration of Oregon and Idaho BLM sagebrush land managers and climate scientists, this study sought to assess land manager needs and define the criteria for useful and useable climate tools. Using an initial online survey, individual phone interviews with land managers, and a follow-up online survey, a series of land management activities and related climate variables were identified, and several web-based climate tools were assessed. Most managers perform vegetation management through a variety of means including seeding and herbicide application. Such activities are affected by the magnitude and timing of precipitation and temperature, as well as other variables, on seasonal and annual timeframes. For planning purposes land managers also need information on long-term 10-20 year climate trends. The act of listening to the needs of land managers uncovered communication barriers, and provided feedback on existing climate tools emphasizing accessibility, dependability and consistency, clear explanation of terminology, effective visualizations, and relevant spatial and temporal scales to the scope of management activities. We also identified a need for basic information and education on the location of existing climate tools and climate impacts, and a need for near-term forecasting tools that could bridge the gap between weather (≤ 6 months) and climate (≥ 30 years) projections. Close

• 42. [Article] Regional changes in landscape pattern and carbon stores in the interior of British Columbia as determined by satellite imagery

  The detection and mapping of large scale changes to forested landscapes is increasingly important in ecology and management. I used Landsat TM and MSS imagery to map forest cover in 1992 for a 4.2 million ...

  Citation × Citation Citation Abstract Copy Title: Regional changes in landscape pattern and carbon stores in the interior of British Columbia as determined by satellite imagery Author: Sachs, Donald The detection and mapping of large scale changes to forested landscapes is increasingly important in ecology and management. I used Landsat TM and MSS imagery to map forest cover in 1992 for a 4.2 million ha area of the interior of British Columbia with an overall classification accuracy of 79%. A combination of unsupervised and supervised classification allowed mapping of forested pixels into closed conifer, semi-open conifer, deciduous, and mixed deciduous and coniferous forest classes. The closed conifer class was further subdivided into old (> 140 years), mature (60 - 140 years), and young (< 60 years) age classes. Unsupervised classification of a combined image containing the Tasseled Cap indices from 1975 and 1992 imagery allowed accurate mapping of disturbances during the period. I examined changes in landscape pattern from 1975 - 1992 by calculating indices that describe overall landscape pattern and that of each cover class. The B.C. biogeoclimatic ecosystem classification map was used as an overlay to calculate landscape pattern and changes by biogeoclimatic zone. Analysis showed that 11.4% of the forested area outside provincial parks was disturbed during the 17 - year period. Disturbed areas were consistently much smaller than conifer patches in all biogeoclimatic zones and had a lower percentage of interior area. Conifer patch shape complexity varied between zones, but the size and shape of disturbed areas were similar in all zones. Results indicated the early stages of fragmentation of this landscape. Models of stand level carbon storage and carbon stores in forest products were used in conjunction with British Columbia Ministry of Forests inventory data to calculate a carbon budget for the Engelmann spruce - subalpine fir forests of the study area. Based on inventory data, the net flux of carbon from these forests to the atmosphere averaged 0.14 Mg ha⁻¹year⁻¹ between 1975 and 1992. Assuming no harvesting during this period, these forests would have been a net carbon sink averaging 0.39 Mg ha⁻¹year⁻¹. Calculating the carbon budget using satellite data showed a slight net carbon sink of 0.15 Mg ha⁻¹year⁻¹ for 1975 - 1992. The major difference between satellite- and inventory-based carbon budgets was due to differences in the simulation of the detrital carbon pool and timing of harvests. The satellite-based estimate failed to account for recovering areas harvested before 1975, and assumed a constant harvest rate. Title: Regional changes in landscape pattern and carbon stores in the interior of British Columbia as determined by satellite imagery Author: Sachs, Donald The detection and mapping of large scale changes to forested landscapes is increasingly important in ecology and management. I used Landsat TM and MSS imagery to map forest cover in 1992 for a 4.2 million ha area of the interior of British Columbia with an overall classification accuracy of 79%. A combination of unsupervised and supervised classification allowed mapping of forested pixels into closed conifer, semi-open conifer, deciduous, and mixed deciduous and coniferous forest classes. The closed conifer class was further subdivided into old (> 140 years), mature (60 - 140 years), and young (< 60 years) age classes. Unsupervised classification of a combined image containing the Tasseled Cap indices from 1975 and 1992 imagery allowed accurate mapping of disturbances during the period. I examined changes in landscape pattern from 1975 - 1992 by calculating indices that describe overall landscape pattern and that of each cover class. The B.C. biogeoclimatic ecosystem classification map was used as an overlay to calculate landscape pattern and changes by biogeoclimatic zone. Analysis showed that 11.4% of the forested area outside provincial parks was disturbed during the 17 - year period. Disturbed areas were consistently much smaller than conifer patches in all biogeoclimatic zones and had a lower percentage of interior area. Conifer patch shape complexity varied between zones, but the size and shape of disturbed areas were similar in all zones. Results indicated the early stages of fragmentation of this landscape. Models of stand level carbon storage and carbon stores in forest products were used in conjunction with British Columbia Ministry of Forests inventory data to calculate a carbon budget for the Engelmann spruce - subalpine fir forests of the study area. Based on inventory data, the net flux of carbon from these forests to the atmosphere averaged 0.14 Mg ha⁻¹year⁻¹ between 1975 and 1992. Assuming no harvesting during this period, these forests would have been a net carbon sink averaging 0.39 Mg ha⁻¹year⁻¹. Calculating the carbon budget using satellite data showed a slight net carbon sink of 0.15 Mg ha⁻¹year⁻¹ for 1975 - 1992. The major difference between satellite- and inventory-based carbon budgets was due to differences in the simulation of the detrital carbon pool and timing of harvests. The satellite-based estimate failed to account for recovering areas harvested before 1975, and assumed a constant harvest rate. Close

• 43. [Article] Impacts of post-fire salvage logging on soil chemistry, physical properties and bacterial community composition in a mixed-conifer forest in central Oregon

  Land managers, scientists, and the interested public are confronted with uncertainty about the impacts of salvage logging on soil productivity. In recent years, stand-replacing wildfires in the western ...

  Citation × Citation Citation Abstract Copy Title: Impacts of post-fire salvage logging on soil chemistry, physical properties and bacterial community composition in a mixed-conifer forest in central Oregon Author: Jennings, Tara Nicolle Land managers, scientists, and the interested public are confronted with uncertainty about the impacts of salvage logging on soil productivity. In recent years, stand-replacing wildfires in the western United States have increased in frequency, prompting the need to evaluate the effect of post-fire treatments on forest ecosystem recovery. This study examined whether compaction and subsoiling after post-fire salvage logging impacted the structure, metabolism, and function of soil bacterial communities, which were assessed using terminal restriction fragment length polymorphism (T-RFLP) and Biolog Ecoplates. Supporting data for these salvage logging effects included soil biological activities, soil chemistry, and physical soil properties in a mixed conifer forest in central Oregon. Post-fire salvage logging treatments had little effect on soil bacterial community richness, suggesting that soil bacteria in these post-fire landscapes are both tolerant of the occurrence of fires and resilient to disturbance. However, even though a statistically significant difference in bacterial species richness was not detected among treatments, a greater cumulative mean number of species was found across six sampling seasons in the compacted soil treatment compared to the fire-only and subsoiled treatments. This trend may be due to less predation on the soil bacteria by microbivores in the soil pores reduced in size by compaction. Additionally, there was a significant decline in the mean number of species after the spring 2007 sampling. A NMS ordination of the mean number of species suggested that the bacterial community composition changed after spring 2007. Furthermore, the lack of difference in mean number of species among treatments suggests that time since fire had a stronger effect on the structure of the soil bacterial community than did logging disturbance. Plant-available phosphorus and nitrogen concentrations were lower in the mechanically disturbed treatments than in the fire-only treatment. Soil respiration and soil phosphatase rates both were higher in the fire-only treatment. Mechanical disturbance may have long-lasting effects in an already nutrient limited system. No other differences among treatments for soil chemical properties were detected. Soil bulk density was lowest in the subsoiled treatment. Microbial community responses to burning and compaction caused by harvesting can be negative, neutral, or positive. While many factors, in addition to soil chemical and physical properties, affect the microbial community richness and functional diversity, our results support other recent studies showing that soil bacteria are resilient in disturbed environments. Title: Impacts of post-fire salvage logging on soil chemistry, physical properties and bacterial community composition in a mixed-conifer forest in central Oregon Author: Jennings, Tara Nicolle Land managers, scientists, and the interested public are confronted with uncertainty about the impacts of salvage logging on soil productivity. In recent years, stand-replacing wildfires in the western United States have increased in frequency, prompting the need to evaluate the effect of post-fire treatments on forest ecosystem recovery. This study examined whether compaction and subsoiling after post-fire salvage logging impacted the structure, metabolism, and function of soil bacterial communities, which were assessed using terminal restriction fragment length polymorphism (T-RFLP) and Biolog Ecoplates. Supporting data for these salvage logging effects included soil biological activities, soil chemistry, and physical soil properties in a mixed conifer forest in central Oregon. Post-fire salvage logging treatments had little effect on soil bacterial community richness, suggesting that soil bacteria in these post-fire landscapes are both tolerant of the occurrence of fires and resilient to disturbance. However, even though a statistically significant difference in bacterial species richness was not detected among treatments, a greater cumulative mean number of species was found across six sampling seasons in the compacted soil treatment compared to the fire-only and subsoiled treatments. This trend may be due to less predation on the soil bacteria by microbivores in the soil pores reduced in size by compaction. Additionally, there was a significant decline in the mean number of species after the spring 2007 sampling. A NMS ordination of the mean number of species suggested that the bacterial community composition changed after spring 2007. Furthermore, the lack of difference in mean number of species among treatments suggests that time since fire had a stronger effect on the structure of the soil bacterial community than did logging disturbance. Plant-available phosphorus and nitrogen concentrations were lower in the mechanically disturbed treatments than in the fire-only treatment. Soil respiration and soil phosphatase rates both were higher in the fire-only treatment. Mechanical disturbance may have long-lasting effects in an already nutrient limited system. No other differences among treatments for soil chemical properties were detected. Soil bulk density was lowest in the subsoiled treatment. Microbial community responses to burning and compaction caused by harvesting can be negative, neutral, or positive. While many factors, in addition to soil chemical and physical properties, affect the microbial community richness and functional diversity, our results support other recent studies showing that soil bacteria are resilient in disturbed environments. Close

• 44. [Article] Factors affecting female space use in ten populations of prairie chickens

  Supplemental material (Appendices A and B) available online at: http://dx.doi.org/10.1890/ES14-00536.1.sm

  Citation × Citation Citation Abstract Copy Title: Factors affecting female space use in ten populations of prairie chickens Author: Carrlson, Kaylan M., Winder, Virginia L., Hagen, Christian A., et al., Gregory, Andrew J. Supplemental material (Appendices A and B) available online at: http://dx.doi.org/10.1890/ES14-00536.1.sm Title: Factors affecting female space use in ten populations of prairie chickens Author: Carrlson, Kaylan M., Winder, Virginia L., Hagen, Christian A., et al., Gregory, Andrew J. Supplemental material (Appendices A and B) available online at: http://dx.doi.org/10.1890/ES14-00536.1.sm Close

• 45. [Article] Habitat selection, reproductive success, and site fidelity of burrowing owls in a grassland ecosystem

  I used a comparative and experimental approach to examine nest habitat selection, reproductive success, and nest site fidelity of burrowing owls (Athene cunicularia) in a large, non-fragmented grassland ...

  Citation × Citation Citation Abstract Copy Title: Habitat selection, reproductive success, and site fidelity of burrowing owls in a grassland ecosystem Author: Ronan, Noelle A. I used a comparative and experimental approach to examine nest habitat selection, reproductive success, and nest site fidelity of burrowing owls (Athene cunicularia) in a large, non-fragmented grassland in southwest California. In 1999, I compared habitat characteristics between nest sites (n = 31) and randomly selected, unoccupied burrows (n = 31) in the local vicinity of the nest (paired burrows). In 2000, I compared habitat characteristics between nest sites (n = 33) and randomly selected, unoccupied burrows (n 32) within the study area (unpaired burrows). I examined reproductive success and variation in nest habitat characteristics, diet quality, and intraspecific competition using data from 1998, 1999, and 2000. I experimentally (n = 11 control and 11 treatment nests) assessed the effect of satellite burrow (multiple auxiliary burrows near the nest) use on productivity and behavior. I found little variation in habitat between nest sites and unoccupied burrows. Habitat selection was not strong when nests and unoccupied burrows were spatially correlated (paired burrows). However, nest sites had a larger number of large diameter burrows, satellite burrows, and perches than the unpaired burrows. Nest success ( I young fledged) and productivity (the number of young alive at 14 -21 days) varied substantially among some years, though the habitat variables I tested did not explain reproductive success when both failed and successful nests were evaluated. When nests were successful, productivity was influenced by rodent consumption. Nest fidelity within the breeding season was highly correlated with nest success. Nest abandonment occurred at 83% (n = 15 of 18), 92% (n = 12 of 13), and 83% (n = 20 of 24) of the failed nests in 1998, 1999, and 2000, respectively. Results of the experimental manipulation of satellite burrow access showed that productivity did not differ between groups but demonstrated that burrowing owls will adjust their behavior to use satellites. Owls in the treatment group (71%; n = 5 of 7) responded by moving their families to areas with access to satellite burrows but none of the control group owl families moved. This study illustrates the importance of identifying critical factors affecting reproductive success of burrowing owls in large grasslands. Maintenance of burrowing mammal populations to provide nest and satellite burrows will be important for protecting burrowing owls. Also, temporal dynamics influenced reproductive success. Habitat characteristics that enhance foraging ability may benefit productivity, especially in years of low rodent numbers. Furthermore, temporal variation in nest success may lead to low nest site fidelity. Title: Habitat selection, reproductive success, and site fidelity of burrowing owls in a grassland ecosystem Author: Ronan, Noelle A. I used a comparative and experimental approach to examine nest habitat selection, reproductive success, and nest site fidelity of burrowing owls (Athene cunicularia) in a large, non-fragmented grassland in southwest California. In 1999, I compared habitat characteristics between nest sites (n = 31) and randomly selected, unoccupied burrows (n = 31) in the local vicinity of the nest (paired burrows). In 2000, I compared habitat characteristics between nest sites (n = 33) and randomly selected, unoccupied burrows (n 32) within the study area (unpaired burrows). I examined reproductive success and variation in nest habitat characteristics, diet quality, and intraspecific competition using data from 1998, 1999, and 2000. I experimentally (n = 11 control and 11 treatment nests) assessed the effect of satellite burrow (multiple auxiliary burrows near the nest) use on productivity and behavior. I found little variation in habitat between nest sites and unoccupied burrows. Habitat selection was not strong when nests and unoccupied burrows were spatially correlated (paired burrows). However, nest sites had a larger number of large diameter burrows, satellite burrows, and perches than the unpaired burrows. Nest success ( I young fledged) and productivity (the number of young alive at 14 -21 days) varied substantially among some years, though the habitat variables I tested did not explain reproductive success when both failed and successful nests were evaluated. When nests were successful, productivity was influenced by rodent consumption. Nest fidelity within the breeding season was highly correlated with nest success. Nest abandonment occurred at 83% (n = 15 of 18), 92% (n = 12 of 13), and 83% (n = 20 of 24) of the failed nests in 1998, 1999, and 2000, respectively. Results of the experimental manipulation of satellite burrow access showed that productivity did not differ between groups but demonstrated that burrowing owls will adjust their behavior to use satellites. Owls in the treatment group (71%; n = 5 of 7) responded by moving their families to areas with access to satellite burrows but none of the control group owl families moved. This study illustrates the importance of identifying critical factors affecting reproductive success of burrowing owls in large grasslands. Maintenance of burrowing mammal populations to provide nest and satellite burrows will be important for protecting burrowing owls. Also, temporal dynamics influenced reproductive success. Habitat characteristics that enhance foraging ability may benefit productivity, especially in years of low rodent numbers. Furthermore, temporal variation in nest success may lead to low nest site fidelity. Close

• 46. [Article] Potential carbon storage at the landscape scale in the Pacific Northwest, U.S.A.

  Estimates of potential carbon (C) storage can be used to constrain predictions of future carbon sequestration and to understand the degree to which disturbances, both natural aid anthropogenic, affect ...

  Citation × Citation Citation Abstract Copy Title: Potential carbon storage at the landscape scale in the Pacific Northwest, U.S.A. Author: Smithwick, Erica Ann Hoffa Estimates of potential carbon (C) storage can be used to constrain predictions of future carbon sequestration and to understand the degree to which disturbances, both natural aid anthropogenic, affect C storage. An upper bound on C storage in the Pacific Northwest (PNW) of the United States was estimated using field data from old-growth forests, which are near steady-state conditions and have been relatively undisturbed for long periods of time. The sites were located across a broad, biogeographical gradient in western Washington and Oregon, allowing comparison of potential carbon storage given a wide range of climate, soils, and vegetation conditions. Total ecosystem carbon (TEC) ranged from 195 Mg C ha' in eastern Oregon to 1127 Mg C ha1 at the Oregon coast. A simple, area-weighted average of TEC to a soil depth of 1 m was 671 Mg C ha1. Compared to estimates of current C storage, up to 338 Mg C ha1 could be stored in addition to current stores in this region. A new model called MAXCARB was developed to predict potential carbon storage over a large area (approximately 1 0 ha), in part to better understand the role of disturbances on potential carbon storage. MAXCARB simulates the effects of climate, soils, or vegetation on potential carbon storage at steady state, for a range of natural and anthropogenic disturbance regimes. Initial results indicate that as the average interval between disturbance events increased, the steady-state C stores at the landscape scale increased. Predictions were well correlated to observed C stores in the PNW. Spatial interactions affect C flux processes at multiple levels of spatial interactions. Using another model, STANDCARB, the relative effect of edge-induced, tree mortality (mainly due to wind), and light limitations, on C dynamics were assessed for several artificial forest landscapes. Emergent behaviors resulting from the interaction of these processes were present at all levels of spatial interaction (stand and landscape). However, the magnitude of the emergent behaviors depended on the spatial structure of the landscape and the level of spatial interaction that was considered. When wind- mortality was high (8 times above natural mortality rates), the dynamics of C processes in fragmented landscapes was not captured using an additive approach. The spatial arrangement of patches on the landscape led to emergent behaviors for one case. However, in many cases, emergent behaviors were mt significant or could be accounted for with traditional modeling methods. Title: Potential carbon storage at the landscape scale in the Pacific Northwest, U.S.A. Author: Smithwick, Erica Ann Hoffa Estimates of potential carbon (C) storage can be used to constrain predictions of future carbon sequestration and to understand the degree to which disturbances, both natural aid anthropogenic, affect C storage. An upper bound on C storage in the Pacific Northwest (PNW) of the United States was estimated using field data from old-growth forests, which are near steady-state conditions and have been relatively undisturbed for long periods of time. The sites were located across a broad, biogeographical gradient in western Washington and Oregon, allowing comparison of potential carbon storage given a wide range of climate, soils, and vegetation conditions. Total ecosystem carbon (TEC) ranged from 195 Mg C ha' in eastern Oregon to 1127 Mg C ha1 at the Oregon coast. A simple, area-weighted average of TEC to a soil depth of 1 m was 671 Mg C ha1. Compared to estimates of current C storage, up to 338 Mg C ha1 could be stored in addition to current stores in this region. A new model called MAXCARB was developed to predict potential carbon storage over a large area (approximately 1 0 ha), in part to better understand the role of disturbances on potential carbon storage. MAXCARB simulates the effects of climate, soils, or vegetation on potential carbon storage at steady state, for a range of natural and anthropogenic disturbance regimes. Initial results indicate that as the average interval between disturbance events increased, the steady-state C stores at the landscape scale increased. Predictions were well correlated to observed C stores in the PNW. Spatial interactions affect C flux processes at multiple levels of spatial interactions. Using another model, STANDCARB, the relative effect of edge-induced, tree mortality (mainly due to wind), and light limitations, on C dynamics were assessed for several artificial forest landscapes. Emergent behaviors resulting from the interaction of these processes were present at all levels of spatial interaction (stand and landscape). However, the magnitude of the emergent behaviors depended on the spatial structure of the landscape and the level of spatial interaction that was considered. When wind- mortality was high (8 times above natural mortality rates), the dynamics of C processes in fragmented landscapes was not captured using an additive approach. The spatial arrangement of patches on the landscape led to emergent behaviors for one case. However, in many cases, emergent behaviors were mt significant or could be accounted for with traditional modeling methods. Close

• 47. [Article] The Influence of fire on the Radiocarbon Signature and Character of Soil Organic Matter in the Siskiyou National Forest, Oregon, USA

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Association ...

  Citation × Citation Citation Abstract Copy Title: The Influence of fire on the Radiocarbon Signature and Character of Soil Organic Matter in the Siskiyou National Forest, Oregon, USA Author: Powers, Heath, Swanston, Chris, Law, Beverly, Campbell, John, Heckman, Katherine To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Association for Fire Ecology and can be found at: http://fireecology.org/. Title: The Influence of fire on the Radiocarbon Signature and Character of Soil Organic Matter in the Siskiyou National Forest, Oregon, USA Author: Powers, Heath, Swanston, Chris, Law, Beverly, Campbell, John, Heckman, Katherine To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Association for Fire Ecology and can be found at: http://fireecology.org/. Close

• 48. [Article] Areal distribution, change, and restoration potential of wetlands within the lower Columbia River riparian zone, 1948-1991

  The lower Columbia River (LCR) riparian zone is rich in habitat diversity. However, the natural beauty and species diversity along the river have increasingly become affected by human activity. This study ...

  Citation × Citation Citation Abstract Copy Title: Areal distribution, change, and restoration potential of wetlands within the lower Columbia River riparian zone, 1948-1991 Author: Allen, Tracy Hunter The lower Columbia River (LCR) riparian zone is rich in habitat diversity. However, the natural beauty and species diversity along the river have increasingly become affected by human activity. This study quantifies the areal extent and degree of wetlands change and associated causes along the LCR over the past 44 years. This research examines the distribution of wetland types and their patterns of change, developing regional models which rank areas most conducive to potential wetland recovery or restoration efforts. The length of the study area totals 234 river kilometers, from the mouth of the Columbia River to Bonneville Dam. The width includes the active channel and an approximately three-kilometer swath on either side of the river. Aerial photography was the primary means for interpreting historical extent of wetlands, using five photo dates (1948, 1961, 1973, 1983, and 1991), based upon their time interval, coverage, and photo quality. For each photo throughout the entire study site, land uses and wetland habitats greater than one hectare were identified and classified. Each classified polygon was digitized and spatially analyzed using a Geographic Information System. This study indicates that wetland habitats which were once contiguously draped upon the linear features of the river are decreasing in size and becoming fragmented. There have been both increases and decreases in specific wetland habitat areas which vary by river reach, even though wetlands have diminished overall. The estuarine section of the LCR experienced a 25% net decrease in wetland area between 1948 and 1991, while the riverine tidal section fostered a 1% increase. The riverine lower perennial section sustained the greatest loss of wetlands, which decreased by 37%. Causes for wetland losses in the estuarine section were largely related to in-water activities, such as channelization, while the causes for declines in the riverine lower perennial section were correlated with rapid urbanization. Wetland increases in the riverine tidal section were generally influenced by significant growth in palustrine and forested wetlands associated with the establishment of wildlife refuges and the incremental increase of upstream flood storage capacity. This research provides a template for identifying degraded or displaced wetlands. Through the use of a GIS, each historical wetland was ranked in either low, moderate, or high categories for restoration potential. GIS technology permits focused, sequentially-refined queries to identify potential restoration or recovery sites. In the estuarine section, 74 historical wetland sites were ranked high for restoration potential, while in the riverine tidal and riverine lower perennial sections, there were 178 and 105, respectively. Overall, these sites represent only 25% of the area occupied by wetlands in 1948. While this study advocates restoration potential, restoration is not a surrogate for responsible ecosystem-wide stewardship of the riparian zone. Restoration will not succeed unless degrading factors are mitigated or eliminated. Title: Areal distribution, change, and restoration potential of wetlands within the lower Columbia River riparian zone, 1948-1991 Author: Allen, Tracy Hunter The lower Columbia River (LCR) riparian zone is rich in habitat diversity. However, the natural beauty and species diversity along the river have increasingly become affected by human activity. This study quantifies the areal extent and degree of wetlands change and associated causes along the LCR over the past 44 years. This research examines the distribution of wetland types and their patterns of change, developing regional models which rank areas most conducive to potential wetland recovery or restoration efforts. The length of the study area totals 234 river kilometers, from the mouth of the Columbia River to Bonneville Dam. The width includes the active channel and an approximately three-kilometer swath on either side of the river. Aerial photography was the primary means for interpreting historical extent of wetlands, using five photo dates (1948, 1961, 1973, 1983, and 1991), based upon their time interval, coverage, and photo quality. For each photo throughout the entire study site, land uses and wetland habitats greater than one hectare were identified and classified. Each classified polygon was digitized and spatially analyzed using a Geographic Information System. This study indicates that wetland habitats which were once contiguously draped upon the linear features of the river are decreasing in size and becoming fragmented. There have been both increases and decreases in specific wetland habitat areas which vary by river reach, even though wetlands have diminished overall. The estuarine section of the LCR experienced a 25% net decrease in wetland area between 1948 and 1991, while the riverine tidal section fostered a 1% increase. The riverine lower perennial section sustained the greatest loss of wetlands, which decreased by 37%. Causes for wetland losses in the estuarine section were largely related to in-water activities, such as channelization, while the causes for declines in the riverine lower perennial section were correlated with rapid urbanization. Wetland increases in the riverine tidal section were generally influenced by significant growth in palustrine and forested wetlands associated with the establishment of wildlife refuges and the incremental increase of upstream flood storage capacity. This research provides a template for identifying degraded or displaced wetlands. Through the use of a GIS, each historical wetland was ranked in either low, moderate, or high categories for restoration potential. GIS technology permits focused, sequentially-refined queries to identify potential restoration or recovery sites. In the estuarine section, 74 historical wetland sites were ranked high for restoration potential, while in the riverine tidal and riverine lower perennial sections, there were 178 and 105, respectively. Overall, these sites represent only 25% of the area occupied by wetlands in 1948. While this study advocates restoration potential, restoration is not a surrogate for responsible ecosystem-wide stewardship of the riparian zone. Restoration will not succeed unless degrading factors are mitigated or eliminated. Close

• 49. [Article] Spatial modeling of carnivore distribution and viability

  Viability analysis of well-selected focal species can complement other types of conservation planning by revealing thresholds in habitat area and landscape connectivity that may not be evident from ecosystem-level ...

  Citation × Citation Citation Abstract Copy Title: Spatial modeling of carnivore distribution and viability Author: Carroll, Carlos Viability analysis of well-selected focal species can complement other types of conservation planning by revealing thresholds in habitat area and landscape connectivity that may not be evident from ecosystem-level evaluations. I used focal species analysis of five carnivores to suggest conservation planning guidelines for the Rocky Mountains and adjacent areas in the Pacific states. I compared static and dynamic viability models for carnivores to assess whether the increased complexity of dynamic simulation models was appropriate given the level of demographic field data typical for carnivore species. I developed models of fisher, lynx, and wolverine distribution from a data set of trapping and sighting records and predicted grizzly bear habitat by adapting previously published regional-scale habitat models. Highquality habitats for wolverine and grizzly bear are strongly associated with low levels of human population and roads. High-quality habitats for fisher and lynx are naturally fragmented by topography and vegetation gradients and are poorly represented in existing protected areas. A comprehensive conservation strategy for carnivores in the region must therefore consider the needs of several species rather than a singie presumed umbrella species. Because many large carnivores are habitat generalists limited by human persecution, and easily-measured attributes such as road density provide robust surrogates for human impact, extrapolation of static habitat models to potential reintroduction areas provides useful guidance for prioritizing carnivore restoration efforts. I also incorporated the static modeling results into a dynamic model. For the wolf; incorporating pack structure into the simulation model increased resilience to human-associated landscape change. For the lynx, relatively low levels of population cycling were found to greatly increase extinction risk when the region was isolated from boreal lynx populations. For the fisher, mortality levels were found to interact strongly with habitat area and isolation to create tlaeshold effects on distribution. Further range contraction is predicted for all species unless coordinated regional planning for habitat restoration occurs. The results suggest that a hybrid approach that builds complex dynamic models on a foundation of data from empirical static models can provide complementary results with a range of perspectives on species' persistence. Title: Spatial modeling of carnivore distribution and viability Author: Carroll, Carlos Viability analysis of well-selected focal species can complement other types of conservation planning by revealing thresholds in habitat area and landscape connectivity that may not be evident from ecosystem-level evaluations. I used focal species analysis of five carnivores to suggest conservation planning guidelines for the Rocky Mountains and adjacent areas in the Pacific states. I compared static and dynamic viability models for carnivores to assess whether the increased complexity of dynamic simulation models was appropriate given the level of demographic field data typical for carnivore species. I developed models of fisher, lynx, and wolverine distribution from a data set of trapping and sighting records and predicted grizzly bear habitat by adapting previously published regional-scale habitat models. Highquality habitats for wolverine and grizzly bear are strongly associated with low levels of human population and roads. High-quality habitats for fisher and lynx are naturally fragmented by topography and vegetation gradients and are poorly represented in existing protected areas. A comprehensive conservation strategy for carnivores in the region must therefore consider the needs of several species rather than a singie presumed umbrella species. Because many large carnivores are habitat generalists limited by human persecution, and easily-measured attributes such as road density provide robust surrogates for human impact, extrapolation of static habitat models to potential reintroduction areas provides useful guidance for prioritizing carnivore restoration efforts. I also incorporated the static modeling results into a dynamic model. For the wolf; incorporating pack structure into the simulation model increased resilience to human-associated landscape change. For the lynx, relatively low levels of population cycling were found to greatly increase extinction risk when the region was isolated from boreal lynx populations. For the fisher, mortality levels were found to interact strongly with habitat area and isolation to create tlaeshold effects on distribution. Further range contraction is predicted for all species unless coordinated regional planning for habitat restoration occurs. The results suggest that a hybrid approach that builds complex dynamic models on a foundation of data from empirical static models can provide complementary results with a range of perspectives on species' persistence. Close

• 50. [Article] Creating Useful and Useable Climate Tools for Sagebrush Land Management Through Scientist and Manager Collaboration

  The sagebrush ecosystem, home to numerous plant and animal species including big sagebrush (Artemisia tridentata) and the endemic greater sage-grouse (Centrocercus urophasianus), has endured fragmentation ...

  Citation × Citation Citation Abstract Copy Title: Creating Useful and Useable Climate Tools for Sagebrush Land Management Through Scientist and Manager Collaboration Author: Brown, Melanie The sagebrush ecosystem, home to numerous plant and animal species including big sagebrush (Artemisia tridentata) and the endemic greater sage-grouse (Centrocercus urophasianus), has endured fragmentation and degradation of both quantity and quality due to the cumulative and synergistic relationships between an abundance of individual disturbances including grazing, invasive annuals and fire. Climate change may now be an additional threat that poses the greatest risk to these imperiled habitats. Natural resource agencies such as the Bureau of Land Management (BLM) seek to conserve sagelands through land management activities that ensure the survival of sage-grouse and continuity of the sagebrush biome. Web-based climate tools can help convey climate information that may be necessary for long-term land management, but these tools may not agree with the needs of land managers, may be too complex, or may be misinterpreted. To overcome barriers of user compatibility, the participation of both climate scientists and land managers is necessary during tool development. With the collaboration of Oregon and Idaho BLM sagebrush land managers and climate scientists, this study sought to assess land manager needs and define the criteria for useful and useable climate tools. Using an initial online survey, individual phone interviews with land managers, and a follow-up online survey, a series of land management activities and related climate variables were identified, and several web-based climate tools were assessed. Most managers perform vegetation management through a variety of means including seeding and herbicide application. Such activities are affected by the magnitude and timing of precipitation and temperature, as well as other variables, on seasonal and annual timeframes. For planning purposes land managers also need information on long-term 10-20 year climate trends. The act of listening to the needs of land managers uncovered communication barriers, and provided feedback on existing climate tools emphasizing accessibility, dependability and consistency, clear explanation of terminology, effective visualizations, and relevant spatial and temporal scales to the scope of management activities. We also identified a need for basic information and education on the location of existing climate tools and climate impacts, and a need for near-term forecasting tools that could bridge the gap between weather (≤ 6 months) and climate (≥ 30 years) projections. Keywords: sagebrush, sage-grouse, Oregon State University, climate change, BLM, Oregon, Idaho, Great Basin, land management, collaboration, climate tools Title: Creating Useful and Useable Climate Tools for Sagebrush Land Management Through Scientist and Manager Collaboration Author: Brown, Melanie The sagebrush ecosystem, home to numerous plant and animal species including big sagebrush (Artemisia tridentata) and the endemic greater sage-grouse (Centrocercus urophasianus), has endured fragmentation and degradation of both quantity and quality due to the cumulative and synergistic relationships between an abundance of individual disturbances including grazing, invasive annuals and fire. Climate change may now be an additional threat that poses the greatest risk to these imperiled habitats. Natural resource agencies such as the Bureau of Land Management (BLM) seek to conserve sagelands through land management activities that ensure the survival of sage-grouse and continuity of the sagebrush biome. Web-based climate tools can help convey climate information that may be necessary for long-term land management, but these tools may not agree with the needs of land managers, may be too complex, or may be misinterpreted. To overcome barriers of user compatibility, the participation of both climate scientists and land managers is necessary during tool development. With the collaboration of Oregon and Idaho BLM sagebrush land managers and climate scientists, this study sought to assess land manager needs and define the criteria for useful and useable climate tools. Using an initial online survey, individual phone interviews with land managers, and a follow-up online survey, a series of land management activities and related climate variables were identified, and several web-based climate tools were assessed. Most managers perform vegetation management through a variety of means including seeding and herbicide application. Such activities are affected by the magnitude and timing of precipitation and temperature, as well as other variables, on seasonal and annual timeframes. For planning purposes land managers also need information on long-term 10-20 year climate trends. The act of listening to the needs of land managers uncovered communication barriers, and provided feedback on existing climate tools emphasizing accessibility, dependability and consistency, clear explanation of terminology, effective visualizations, and relevant spatial and temporal scales to the scope of management activities. We also identified a need for basic information and education on the location of existing climate tools and climate impacts, and a need for near-term forecasting tools that could bridge the gap between weather (≤ 6 months) and climate (≥ 30 years) projections. Keywords: sagebrush, sage-grouse, Oregon State University, climate change, BLM, Oregon, Idaho, Great Basin, land management, collaboration, climate tools Close