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• 1. [Article] Stability of International Fisheries Agreements: Implications of Non-Market Benefits and the Country of Origin

  Salmon is an anadromous species that during its feeding and spawning migrations is sought after catch for commercial and recreational fisheries. The management of salmon fisheries is complicated by the ...

  Citation × Citation Citation Abstract Copy Title: Stability of International Fisheries Agreements: Implications of Non-Market Benefits and the Country of Origin Author: Levontin, Polina, Kulmala, Soile, Laukkanen, Marita, Lindroos, Marko, Romakkaniemi, Atso, Nieminen, Emmi, Parkkila, Katja, Pintassilgo, Pedro, Pulkkinen, Henni Salmon is an anadromous species that during its feeding and spawning migrations is sought after catch for commercial and recreational fisheries. The management of salmon fisheries is complicated by the combination of mixed and single stock fisheries. Thus, the country of origin has sovereign control over harvest of a salmon stock only at the last steps of the gauntlet. This paper addresses the stability of an international fisheries agreement on the Baltic salmon. This setting is modeled through a coalition game in the partition function form with four asymmetric players. Countries payoffs depend both on commercial fishery’s profits and net benefits from recreational harvest. Moreover, the country of origin must ensure that each of the salmon stocks achieve or maintain a sustainable size. The economic sub-model is calibrated using commercial fisheries statistics and existing non-market valuation studies. The underlying population dynamics model accounts for 15 salmon stocks and it is used in the actual stock assessment. The results indicate that by considering economic aspects of recreational fisheries it is possible to stabilize the grand coalition. However, the cooperative strategies of the grand coalition do not ensure biologically sound harvesting of all salmon stocks. Title: Stability of International Fisheries Agreements: Implications of Non-Market Benefits and the Country of Origin Author: Levontin, Polina, Kulmala, Soile, Laukkanen, Marita, Lindroos, Marko, Romakkaniemi, Atso, Nieminen, Emmi, Parkkila, Katja, Pintassilgo, Pedro, Pulkkinen, Henni Salmon is an anadromous species that during its feeding and spawning migrations is sought after catch for commercial and recreational fisheries. The management of salmon fisheries is complicated by the combination of mixed and single stock fisheries. Thus, the country of origin has sovereign control over harvest of a salmon stock only at the last steps of the gauntlet. This paper addresses the stability of an international fisheries agreement on the Baltic salmon. This setting is modeled through a coalition game in the partition function form with four asymmetric players. Countries payoffs depend both on commercial fishery’s profits and net benefits from recreational harvest. Moreover, the country of origin must ensure that each of the salmon stocks achieve or maintain a sustainable size. The economic sub-model is calibrated using commercial fisheries statistics and existing non-market valuation studies. The underlying population dynamics model accounts for 15 salmon stocks and it is used in the actual stock assessment. The results indicate that by considering economic aspects of recreational fisheries it is possible to stabilize the grand coalition. However, the cooperative strategies of the grand coalition do not ensure biologically sound harvesting of all salmon stocks. Close

• 2. [Article] Ecological and evolutionary patterns of freshwater maturation in Pacific and Atlantic salmonines

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer ...

  Citation × Citation Citation Abstract Copy Title: Ecological and evolutionary patterns of freshwater maturation in Pacific and Atlantic salmonines Author: Ohms, Haley A., Sloat, Matthew R., et al., Dunham, Jason B., Fraser, Dylan J. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer and can be found at: http://link.springer.com/journal/11160. Title: Ecological and evolutionary patterns of freshwater maturation in Pacific and Atlantic salmonines Author: Ohms, Haley A., Sloat, Matthew R., et al., Dunham, Jason B., Fraser, Dylan J. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer and can be found at: http://link.springer.com/journal/11160. Close

• 3. [Article] The Economics of Conservation Targeting Strategies

  This is an abbreviated version of a paper published in the American Journal of Agricultural Economics, Vol. 82 #2, pages 400-413.

  Citation × Citation Citation Abstract Copy Title: The Economics of Conservation Targeting Strategies Author: Boggess, William G., Wu, JunJie, Adams, Richard M. Year: 2001 This is an abbreviated version of a paper published in the American Journal of Agricultural Economics, Vol. 82 #2, pages 400-413. Title: The Economics of Conservation Targeting Strategies Author: Boggess, William G., Wu, JunJie, Adams, Richard M. Year: 2001 This is an abbreviated version of a paper published in the American Journal of Agricultural Economics, Vol. 82 #2, pages 400-413. Close

• 4. [Article] Risk assessment : introduction and establishment of Myxobolus cerebralis in the Deschutes River Basin, Oregon, USA

  A plan for the reestablishment of anadromous salmonid stocks into waters above the Pelton Round Butte Hydroelectric Project (PRB) located on the Deschutes River, Oregon has been under development since ...

  Citation × Citation Citation Abstract Copy Title: Risk assessment : introduction and establishment of Myxobolus cerebralis in the Deschutes River Basin, Oregon, USA Author: Zielinski, Christopher A plan for the reestablishment of anadromous salmonid stocks into waters above the Pelton Round Butte Hydroelectric Project (PRB) located on the Deschutes River, Oregon has been under development since 1996. The PRB complex, starting at river kilometer (Rkm) 103, blocks the further upstream migration of anadromous salmonids into upper river basin waters. Part of the process for evaluating the passage plan was the development of a Fish Health Management Program. The program is, in part, designed to minimize and communicate the magnitude of risk associated with passing serious fish pathogens upstream of the PRB until evaluation of the fish passage effort confirms that the reestablishment of anadromous fish species can be successful. With the goal of minimizing the risk of passing serious fish pathogens along with evaluating the magnitude of risk, several potential pathogens of concern were identified. At the forefront of concern is the fish pathogen Myxobolus cerebralis which is responsible for whirling disease in salmonids. Annual, or at least periodic, introduction of M. cerebralis into the lower DRB has occurred since at least 1984 as a result of infected stray adult salmonids spawning and dying there. These are primarily hatchery summer steelhead (Oncorhynchus mykiss), and to a lesser extent spring Chinook salmon (O. tshawytscha), from the Snake River system. Because of its' potential to cause significant mortality, the risk associated with M. cerebralis establishment in the DRB needed to be addressed. The risk of the parasite being transmitted and established upstream of the PRB with the renewal of fish passage at the PRB, and the potential for M. cerebralis to affect survival of reintroduced anadromous fish species required investigation. To evaluate these potential risks, we conducted multi-year studies (1998-2007) that informed a risk-assessment model. Tailored specifically for M. cerebralis, this model was used to identify key characteristics that will influence pathways of M. cerebralis introduction, the potential for establishment, and the consequences of establishment. The three steps involved in the risk assessment model are the release, exposure and consequence assessments. Using this model we evaluated the probability of introduction and establishment of M. cerebralis in the lower Deschutes River Basin (DRB) as well as in waters above the PRB with future passage of anadromous fish. The current study gives evidence that M. cerebralis is established in the lower DRB. The current study also gives evidence not all areas of the DRB can be classified as having the same likelihood for establishment and the potential impact the parasite poses will not be identical from one location to another. It is clear, however, that M. cerebralis could become established above the PRB if infected fish carry the myxospore into these waters. Title: Risk assessment : introduction and establishment of Myxobolus cerebralis in the Deschutes River Basin, Oregon, USA Author: Zielinski, Christopher A plan for the reestablishment of anadromous salmonid stocks into waters above the Pelton Round Butte Hydroelectric Project (PRB) located on the Deschutes River, Oregon has been under development since 1996. The PRB complex, starting at river kilometer (Rkm) 103, blocks the further upstream migration of anadromous salmonids into upper river basin waters. Part of the process for evaluating the passage plan was the development of a Fish Health Management Program. The program is, in part, designed to minimize and communicate the magnitude of risk associated with passing serious fish pathogens upstream of the PRB until evaluation of the fish passage effort confirms that the reestablishment of anadromous fish species can be successful. With the goal of minimizing the risk of passing serious fish pathogens along with evaluating the magnitude of risk, several potential pathogens of concern were identified. At the forefront of concern is the fish pathogen Myxobolus cerebralis which is responsible for whirling disease in salmonids. Annual, or at least periodic, introduction of M. cerebralis into the lower DRB has occurred since at least 1984 as a result of infected stray adult salmonids spawning and dying there. These are primarily hatchery summer steelhead (Oncorhynchus mykiss), and to a lesser extent spring Chinook salmon (O. tshawytscha), from the Snake River system. Because of its' potential to cause significant mortality, the risk associated with M. cerebralis establishment in the DRB needed to be addressed. The risk of the parasite being transmitted and established upstream of the PRB with the renewal of fish passage at the PRB, and the potential for M. cerebralis to affect survival of reintroduced anadromous fish species required investigation. To evaluate these potential risks, we conducted multi-year studies (1998-2007) that informed a risk-assessment model. Tailored specifically for M. cerebralis, this model was used to identify key characteristics that will influence pathways of M. cerebralis introduction, the potential for establishment, and the consequences of establishment. The three steps involved in the risk assessment model are the release, exposure and consequence assessments. Using this model we evaluated the probability of introduction and establishment of M. cerebralis in the lower Deschutes River Basin (DRB) as well as in waters above the PRB with future passage of anadromous fish. The current study gives evidence that M. cerebralis is established in the lower DRB. The current study also gives evidence not all areas of the DRB can be classified as having the same likelihood for establishment and the potential impact the parasite poses will not be identical from one location to another. It is clear, however, that M. cerebralis could become established above the PRB if infected fish carry the myxospore into these waters. Close

• 5. [Article] Environmental variables as predictors of fish assemblages in the Tillamook Basin, Oregon

  The Tillamook and Kilchis subbasins of the Tillamook Bay watershed (Oregon Coast Range) have differences in geology and land use history and, therefore, varied environmental conditions that could affect ...

  Citation × Citation Citation Abstract Copy Title: Environmental variables as predictors of fish assemblages in the Tillamook Basin, Oregon Author: Rose, Cathleen E. The Tillamook and Kilchis subbasins of the Tillamook Bay watershed (Oregon Coast Range) have differences in geology and land use history and, therefore, varied environmental conditions that could affect fish assemblages. Fifty-two randomly selected wadeable stream reaches in these two subbasins were surveyed for stream habitat and fish assemblage composition during two summer field seasons. Data collected were used to identify environmental variables most associated with fish assemblages, determine major predictors of fish species relative abundance, and test for any differences in fish assemblages between the Tillamook and Kilchis subbasins and among three stream orders. Eleven fish species were encountered, including four salmonid species (Oncorhynchus spp.) and five species of sculpin (Cottus spp.). For three sites visited twice in both field seasons, variability in fish assemblages and environmental measurements between sample years and between early and late-summer visits to sites was low. Fish assemblages differed between the two subbasins. The assemblages in the Kilchis subbasin contained higher proportions of trout species, while the Tillamook subbasin assemblages contained higher proportions of sculpin species. A breakdown of fish assemblages by stream order showed that different fish species dominated the assemblages in different sized streams. With data from both subbasins combined, trout species composed a smaller proportion of the fish assemblages in larger streams, and sculpin species composition also varied with stream order. Results from non-metric multidimensional scaling ordinations indicated that substrate type, stream gradient, and variables related to stream size were the most highly correlated to fish species relative abundances. Multiple linear regression was used to assess the usefulness of an uncorrelated subset of the environmental variables as predictors of relative abundance of the six most common fish species. The environmental variables were good predictors of relative abundance for some species, but were less successful in predicting abundances of entirely anadromous species. These results provide valuable insight into fish-habitat relationships using a fish assemblage perspective and incorporating species, such as sculpins, for which there is little existing information. There has been recent interest in quantifying the effects of human land use on stream fish, and these data can be used in future studies to explore potential links between important environmental variables and land use practices. Title: Environmental variables as predictors of fish assemblages in the Tillamook Basin, Oregon Author: Rose, Cathleen E. The Tillamook and Kilchis subbasins of the Tillamook Bay watershed (Oregon Coast Range) have differences in geology and land use history and, therefore, varied environmental conditions that could affect fish assemblages. Fifty-two randomly selected wadeable stream reaches in these two subbasins were surveyed for stream habitat and fish assemblage composition during two summer field seasons. Data collected were used to identify environmental variables most associated with fish assemblages, determine major predictors of fish species relative abundance, and test for any differences in fish assemblages between the Tillamook and Kilchis subbasins and among three stream orders. Eleven fish species were encountered, including four salmonid species (Oncorhynchus spp.) and five species of sculpin (Cottus spp.). For three sites visited twice in both field seasons, variability in fish assemblages and environmental measurements between sample years and between early and late-summer visits to sites was low. Fish assemblages differed between the two subbasins. The assemblages in the Kilchis subbasin contained higher proportions of trout species, while the Tillamook subbasin assemblages contained higher proportions of sculpin species. A breakdown of fish assemblages by stream order showed that different fish species dominated the assemblages in different sized streams. With data from both subbasins combined, trout species composed a smaller proportion of the fish assemblages in larger streams, and sculpin species composition also varied with stream order. Results from non-metric multidimensional scaling ordinations indicated that substrate type, stream gradient, and variables related to stream size were the most highly correlated to fish species relative abundances. Multiple linear regression was used to assess the usefulness of an uncorrelated subset of the environmental variables as predictors of relative abundance of the six most common fish species. The environmental variables were good predictors of relative abundance for some species, but were less successful in predicting abundances of entirely anadromous species. These results provide valuable insight into fish-habitat relationships using a fish assemblage perspective and incorporating species, such as sculpins, for which there is little existing information. There has been recent interest in quantifying the effects of human land use on stream fish, and these data can be used in future studies to explore potential links between important environmental variables and land use practices. Close

• 6. [Article] Distribution of fishes and their relationships to environments in selected coastal streams, Douglas and Coos Counties, Oregon

  This study was an investigation to determine the distribution, abundance and general food habits of salmonids and non-game fishes residing in several selected coastal streams, Douglas and Coos Counties, ...

  Citation × Citation Citation Abstract Copy Title: Distribution of fishes and their relationships to environments in selected coastal streams, Douglas and Coos Counties, Oregon Author: Duke, Stephen D'Wayne This study was an investigation to determine the distribution, abundance and general food habits of salmonids and non-game fishes residing in several selected coastal streams, Douglas and Coos Counties, Oregon. The physical conditions that characterize the various stream habitat types were monitored and the influence of physical conditions on the distribution and abundance of fish was investigated. Study of fish distribution began fall 1979 and continued through summer 1980. Biological and ecological monitoring was undertaken at eleven permanent study sites on four occasions during the summer period of decreasing flow. Population size of each species and several environmental characteristics were measured during these occasions. The study streams were characterized by good populations of juvenile coho salmon (Oncorhynchus kisutch) and steelhead (rainbow) trout (Salmo gairdneri), demonstrating the importance of these streams as rearing areas for anadromous species. Reticulate sculpin (Cottus perplexus) were the most abundant and widespread non-game fish collected during this study. Insects were the principal prey organisms in the diets of all species studied, with Ephemeroptera and Diptera larvae the major components. There was also considerable overlap in the diets. Coho salmon and steelhead trout were most versatile in acceptance of foods, taking 19 and 15 of the separate categories tabulated. The benthic fishes, speckled dace (Rhinichthys osculus), reticulate and riffle sculpin (Cottus gulosus) were more restricted in diet, taking 12, 7 and 14 food types respectively. Diversity of habitat within the various study streams was attributable not only to geological and climatic factors, but disturbances by fire, erosion, activity of beaver and logging. Mostly such disturbances are deleterious, though in at least one instance effects of disturbances were beneficial. Beaver ponds created excellent rearing pools for coho salmon in one stream. Instream distribution of fishes appears to be mainly a function of habitat selection. Coho salmon were found in loose schools throughout pools in areas where short pools alternated with riffles. Steelhead and cutthroat trout (Salmo clarki) were found most often associated with instream cover. The reticulate and riffle sculpin, both of which are benthic and cryptic and select the same kind of cover, were usually associated in gravelly riffles. Speckled dace were common throughout large streams and almost always in the vicinity of adequate cover. Redside shiners (Richardsonius balteatus) were associated with pool like environments and were excluded from upstream areas with increasing gradient and shallow pools. Limitations on populations of salmonid fishes in the study streams are probably associated with lack of adequate summer flow, lack of instream cover, siltation of gravel and rubble bottoms and departure from an approximate 1:1 pool to riffle ratio. Title: Distribution of fishes and their relationships to environments in selected coastal streams, Douglas and Coos Counties, Oregon Author: Duke, Stephen D'Wayne This study was an investigation to determine the distribution, abundance and general food habits of salmonids and non-game fishes residing in several selected coastal streams, Douglas and Coos Counties, Oregon. The physical conditions that characterize the various stream habitat types were monitored and the influence of physical conditions on the distribution and abundance of fish was investigated. Study of fish distribution began fall 1979 and continued through summer 1980. Biological and ecological monitoring was undertaken at eleven permanent study sites on four occasions during the summer period of decreasing flow. Population size of each species and several environmental characteristics were measured during these occasions. The study streams were characterized by good populations of juvenile coho salmon (Oncorhynchus kisutch) and steelhead (rainbow) trout (Salmo gairdneri), demonstrating the importance of these streams as rearing areas for anadromous species. Reticulate sculpin (Cottus perplexus) were the most abundant and widespread non-game fish collected during this study. Insects were the principal prey organisms in the diets of all species studied, with Ephemeroptera and Diptera larvae the major components. There was also considerable overlap in the diets. Coho salmon and steelhead trout were most versatile in acceptance of foods, taking 19 and 15 of the separate categories tabulated. The benthic fishes, speckled dace (Rhinichthys osculus), reticulate and riffle sculpin (Cottus gulosus) were more restricted in diet, taking 12, 7 and 14 food types respectively. Diversity of habitat within the various study streams was attributable not only to geological and climatic factors, but disturbances by fire, erosion, activity of beaver and logging. Mostly such disturbances are deleterious, though in at least one instance effects of disturbances were beneficial. Beaver ponds created excellent rearing pools for coho salmon in one stream. Instream distribution of fishes appears to be mainly a function of habitat selection. Coho salmon were found in loose schools throughout pools in areas where short pools alternated with riffles. Steelhead and cutthroat trout (Salmo clarki) were found most often associated with instream cover. The reticulate and riffle sculpin, both of which are benthic and cryptic and select the same kind of cover, were usually associated in gravelly riffles. Speckled dace were common throughout large streams and almost always in the vicinity of adequate cover. Redside shiners (Richardsonius balteatus) were associated with pool like environments and were excluded from upstream areas with increasing gradient and shallow pools. Limitations on populations of salmonid fishes in the study streams are probably associated with lack of adequate summer flow, lack of instream cover, siltation of gravel and rubble bottoms and departure from an approximate 1:1 pool to riffle ratio. Close

• 7. [Article] An economic analysis of alternative water allocations and habitat investments for anadromous fish production, John Day Basin, Oregon

  This thesis focuses on the economic efficiency of alternative habitat, streamfiow, and land use allocations within a major drainage of the Pacific Northwest. Specifically, productivity of present habitat and ...

  Citation × Citation Citation Abstract Copy Title: An economic analysis of alternative water allocations and habitat investments for anadromous fish production, John Day Basin, Oregon Author: Cerda, Arcadio Alberto This thesis focuses on the economic efficiency of alternative habitat, streamfiow, and land use allocations within a major drainage of the Pacific Northwest. Specifically, productivity of present habitat and streamfiow conditions is evaluated with respect to anadromous fish species (chinook salmon and steelhead trout) in the John Day River Basin, a high-desert system in the Pacific Northwest. Alternative water allocations and fish habitat enhancements to improve anadromous fish productivity are also examined. This biophysical information is combined with economics concepts to evaluate comparative benefits and economic efficiency of selected technically feasible habitat/streamflow alternatives. The John Day River Basin is used as a case study because it provides one of the most significant runs of wild summer steelhead trout and spring chinook salmon in the Columbia River Basin. Specific categories of alternative enhancements included water transfers to instream flow, habitat vegetation improvements, and habitat structural measures. The benefits of changes in fish numbers are valued through the travel cost method. Costs of habitat investments are derived from a synthesis of cost data for similar investments for the region. For most lower-reach stream sections, benefits of habitat investments/streamflow alternatives do not equal costs. However, changes in water allocations and habitat do appear economically efficient to some upper reaches. The main results suggest that economic efficiency varies greatly within the system, indicating the need for site-specific proposals for streamfiow or habitat alterations within this or similar high desert streams. In addition to providing preliminary estimates of the economic efficency of streamfiow and habitat investments, this thesis serves as a methodological template for future integrated bioeconomic evaluations of water and fishery management issues. Title: An economic analysis of alternative water allocations and habitat investments for anadromous fish production, John Day Basin, Oregon Author: Cerda, Arcadio Alberto This thesis focuses on the economic efficiency of alternative habitat, streamfiow, and land use allocations within a major drainage of the Pacific Northwest. Specifically, productivity of present habitat and streamfiow conditions is evaluated with respect to anadromous fish species (chinook salmon and steelhead trout) in the John Day River Basin, a high-desert system in the Pacific Northwest. Alternative water allocations and fish habitat enhancements to improve anadromous fish productivity are also examined. This biophysical information is combined with economics concepts to evaluate comparative benefits and economic efficiency of selected technically feasible habitat/streamflow alternatives. The John Day River Basin is used as a case study because it provides one of the most significant runs of wild summer steelhead trout and spring chinook salmon in the Columbia River Basin. Specific categories of alternative enhancements included water transfers to instream flow, habitat vegetation improvements, and habitat structural measures. The benefits of changes in fish numbers are valued through the travel cost method. Costs of habitat investments are derived from a synthesis of cost data for similar investments for the region. For most lower-reach stream sections, benefits of habitat investments/streamflow alternatives do not equal costs. However, changes in water allocations and habitat do appear economically efficient to some upper reaches. The main results suggest that economic efficiency varies greatly within the system, indicating the need for site-specific proposals for streamfiow or habitat alterations within this or similar high desert streams. In addition to providing preliminary estimates of the economic efficency of streamfiow and habitat investments, this thesis serves as a methodological template for future integrated bioeconomic evaluations of water and fishery management issues. Close

• 8. [Article] Proposed regional general permit for stream restoration : programmatic biological opinion

  On April 7, 2000, the National Marine Fisheries Service (NMFS) received a request from the U.S. Army Corps of Engineers, Portland District (COE) for Endangered Species Act (ESA) section 7 formal consultation ...

  Citation × Citation Citation Abstract Copy Title: Proposed regional general permit for stream restoration : programmatic biological opinion On April 7, 2000, the National Marine Fisheries Service (NMFS) received a request from the U.S. Army Corps of Engineers, Portland District (COE) for Endangered Species Act (ESA) section 7 formal consultation for programmatic coverage for a proposed Regional General Permit (RGP) for certain stream restoration activities (habitat restoration using large wood or boulders throughout the State of Oregon) requiring COE approval. The Biological Assessment (BA) provided with the request determined that the proposed activities covered under the RGP would be likely to adversely affect anadromous fish species listed under the ESA (Table 1). Species considered in this Biological Opinion (Opinion) are: Snake River sockeye salmon (Oncorhynchus nerka); Snake River spring /summer chinook salmon (O. tshawytscha); Snake River fall chinook salmon (O. tshawytscha); Lower Columbia River steelhead (O. mykiss); Upper Columbia River steelhead (O. mykiss); Snake River steelhead (O. mykiss); Upper Willamette River steelhead (O. mykiss); Middle Columbia River steelhead (O. mykiss); Columbia River chum salmon (O. keta); Lower Columbia River chinook salmon (O. tshawytscha); Upper Willamette River chinook salmon (O. tshawytscha); Upper Columbia River spring run chinook salmon (O. tshawytscha); S. Oregon/N. California Coast coho salmon (O. kisutch); and Oregon Coast coho salmon (O. kisutch). On April 18, 2000, NMFS received a letter from COE providing additional information on proposed criteria to be included in the RGP. On May 12, 2000, NMFS received an e-mail with further proposed criteria to be included in the RGP. Title: Proposed regional general permit for stream restoration : programmatic biological opinion On April 7, 2000, the National Marine Fisheries Service (NMFS) received a request from the U.S. Army Corps of Engineers, Portland District (COE) for Endangered Species Act (ESA) section 7 formal consultation for programmatic coverage for a proposed Regional General Permit (RGP) for certain stream restoration activities (habitat restoration using large wood or boulders throughout the State of Oregon) requiring COE approval. The Biological Assessment (BA) provided with the request determined that the proposed activities covered under the RGP would be likely to adversely affect anadromous fish species listed under the ESA (Table 1). Species considered in this Biological Opinion (Opinion) are: Snake River sockeye salmon (Oncorhynchus nerka); Snake River spring /summer chinook salmon (O. tshawytscha); Snake River fall chinook salmon (O. tshawytscha); Lower Columbia River steelhead (O. mykiss); Upper Columbia River steelhead (O. mykiss); Snake River steelhead (O. mykiss); Upper Willamette River steelhead (O. mykiss); Middle Columbia River steelhead (O. mykiss); Columbia River chum salmon (O. keta); Lower Columbia River chinook salmon (O. tshawytscha); Upper Willamette River chinook salmon (O. tshawytscha); Upper Columbia River spring run chinook salmon (O. tshawytscha); S. Oregon/N. California Coast coho salmon (O. kisutch); and Oregon Coast coho salmon (O. kisutch). On April 18, 2000, NMFS received a letter from COE providing additional information on proposed criteria to be included in the RGP. On May 12, 2000, NMFS received an e-mail with further proposed criteria to be included in the RGP. Close

• 9. [Article] Statistical analysis on otolith data of anadromous fishes

  To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer ...

  Citation × Citation Citation Abstract Copy Title: Statistical analysis on otolith data of anadromous fishes Author: Bean, David, Noakes, David L. G., Conrad, Robert, Gao, Yongwen To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer and can be found at: http://www.springer.com/?SGWID=0-102-0-0-0. Title: Statistical analysis on otolith data of anadromous fishes Author: Bean, David, Noakes, David L. G., Conrad, Robert, Gao, Yongwen To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. This is the publisher’s final pdf. The published article is copyrighted by Springer and can be found at: http://www.springer.com/?SGWID=0-102-0-0-0. Close

• 10. [Article] Salmon Abundances and Effects of Harvest: Implications for Rebuilding Stocks of Wild Coho Salmon in Oregon

  This report was prepared by the Independent Multidisciplinary Science Team of the Oregon Plan for Salmon and Watersheds.

  Citation × Citation Citation Abstract Copy Title: Salmon Abundances and Effects of Harvest: Implications for Rebuilding Stocks of Wild Coho Salmon in Oregon Author: Independent Multidisciplinary Science Team (Or.) This report was prepared by the Independent Multidisciplinary Science Team of the Oregon Plan for Salmon and Watersheds. Title: Salmon Abundances and Effects of Harvest: Implications for Rebuilding Stocks of Wild Coho Salmon in Oregon Author: Independent Multidisciplinary Science Team (Or.) This report was prepared by the Independent Multidisciplinary Science Team of the Oregon Plan for Salmon and Watersheds. Close

• 11. [Article] Sandy beach surf zones: An alternative nursery habitat for 0-age Chinook salmon

  This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/estuarine-coasta...

  Citation × Citation Citation Abstract Copy Title: Sandy beach surf zones: An alternative nursery habitat for 0-age Chinook salmon Author: Miller, J. A., Marin Jarrin, J. R. This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/estuarine-coastal-and-shelf-science/. Title: Sandy beach surf zones: An alternative nursery habitat for 0-age Chinook salmon Author: Miller, J. A., Marin Jarrin, J. R. This is an author's peer-reviewed final manuscript, as accepted by the publisher. The published article is copyrighted by Elsevier and can be found at: http://www.journals.elsevier.com/estuarine-coastal-and-shelf-science/. Close

• 12. [Article] Sandy beach surf zones : what is their role in the early life history of Chinook salmon (Oncorhynchus tshawytscha)?

  Early life stages of many marine and diadromous fish species use sandy beach surf zones, which occur along >50% of the world's marine coastlines. This extensive habitat can provide juvenile fishes with ...

  Citation × Citation Citation Abstract Copy Title: Sandy beach surf zones : what is their role in the early life history of Chinook salmon (Oncorhynchus tshawytscha)? Author: Marin Jarrin, Jose R., 1980- Early life stages of many marine and diadromous fish species use sandy beach surf zones, which occur along >50% of the world's marine coastlines. This extensive habitat can provide juvenile fishes with an abundant supply of potential prey and the ability to hide from predators in its shallow turbid waters. Chinook salmon is an anadromous species that migrates to the ocean during their first (subyearlings) or second (yearlings) year of life. The majority of subyearlings reside in estuaries during their first summer season; however, a small number of juveniles also use surf zones. Early marine residence is considered a critical period for Chinook salmon due to high mortality rates; however the role of surf zones in Chinook salmon life history is unclear. Therefore, I determined the distribution of juvenile Chinook salmon on beaches of the eastern North Pacific, compared the migration and growth patterns observed in surf zones and estuaries, identified the factors that accounted for variation in juvenile surf zone catch, explored the factors that influence growth rate variation in surf zones and estuaries, and modeled how growth rates in these coastal habitats may vary in the near future with predicted changes in climate. The majority (94%) of juveniles were caught in surf zones adjacent to estuaries with trough areas, which are beach sections where sand moved by currents and waves produce a trench-like shape. Surf zone fish were collected in significantly lower numbers than estuarine juveniles but entered brackish/ocean waters at similar sizes. Juveniles in surf zones consumed similar organisms (gammarid amphipods, crustacean larvae and insects) as in estuaries. Furthermore, stomach fullness indices (average = 2% of body weight) and growth rates (average = 0.4 mm day⁻¹) were similar in surf zones and estuaries. At one surf zone, juvenile catch was positively correlated to short-term specific growth rates (14 days prior to capture). A bioenergetics modeling approach indicated that given current conditions, consumption rates accounted for more of the variation in growth than prey energetic content and temperature. Climate models predict future increases in fresh water temperature (1.5 to 5.8°C), sea surface temperature (1.2°C) and wave height (0.75 m) that could influence estuarine and surf zone use. Therefore, I developed a local mixing model based on these predictions to estimate future surf zone and estuarine water temperatures in two of the watersheds studied. Based on these temperature projections and the bioenergetics model, I predicted how juvenile specific growth rates would vary in both habitats. I determined that increases in water temperature in both habitats would reduce specific growth rates by 9 to 40% in surf zones and estuaries if diet composition and consumption rates remain similar to present conditions. To compensate for the decline in growth, juveniles may increase their consumption rates or consume more energetically rich prey, if available. If they are not able to compensate, their size at the end of the season may be reduced, which could reduce their overall survival. These results confirm that a small number of suyearling Chinook salmon use sandy beach surf zones, mostly adjacent to estuary mouths, where they experience growth conditions comparable to estuaries. My findings indicate that, in certain situations, juvenile Chinook salmon surf zone use can be influenced by surf zone growth conditions, while variation in growth rates are themselves most strongly influenced by variation in consumption rates in surf zones and estuaries. Predicted changes in coastal western North American climate will likely modify juvenile growth conditions in the next 50 years, and potentially reduce overall survival. Additional insights into the potential impacts of climate change on juvenile salmon will require estimates of changes in the composition, energetic quality and abundance of prey communities inhabiting coastal environments. Title: Sandy beach surf zones : what is their role in the early life history of Chinook salmon (Oncorhynchus tshawytscha)? Author: Marin Jarrin, Jose R., 1980- Early life stages of many marine and diadromous fish species use sandy beach surf zones, which occur along >50% of the world's marine coastlines. This extensive habitat can provide juvenile fishes with an abundant supply of potential prey and the ability to hide from predators in its shallow turbid waters. Chinook salmon is an anadromous species that migrates to the ocean during their first (subyearlings) or second (yearlings) year of life. The majority of subyearlings reside in estuaries during their first summer season; however, a small number of juveniles also use surf zones. Early marine residence is considered a critical period for Chinook salmon due to high mortality rates; however the role of surf zones in Chinook salmon life history is unclear. Therefore, I determined the distribution of juvenile Chinook salmon on beaches of the eastern North Pacific, compared the migration and growth patterns observed in surf zones and estuaries, identified the factors that accounted for variation in juvenile surf zone catch, explored the factors that influence growth rate variation in surf zones and estuaries, and modeled how growth rates in these coastal habitats may vary in the near future with predicted changes in climate. The majority (94%) of juveniles were caught in surf zones adjacent to estuaries with trough areas, which are beach sections where sand moved by currents and waves produce a trench-like shape. Surf zone fish were collected in significantly lower numbers than estuarine juveniles but entered brackish/ocean waters at similar sizes. Juveniles in surf zones consumed similar organisms (gammarid amphipods, crustacean larvae and insects) as in estuaries. Furthermore, stomach fullness indices (average = 2% of body weight) and growth rates (average = 0.4 mm day⁻¹) were similar in surf zones and estuaries. At one surf zone, juvenile catch was positively correlated to short-term specific growth rates (14 days prior to capture). A bioenergetics modeling approach indicated that given current conditions, consumption rates accounted for more of the variation in growth than prey energetic content and temperature. Climate models predict future increases in fresh water temperature (1.5 to 5.8°C), sea surface temperature (1.2°C) and wave height (0.75 m) that could influence estuarine and surf zone use. Therefore, I developed a local mixing model based on these predictions to estimate future surf zone and estuarine water temperatures in two of the watersheds studied. Based on these temperature projections and the bioenergetics model, I predicted how juvenile specific growth rates would vary in both habitats. I determined that increases in water temperature in both habitats would reduce specific growth rates by 9 to 40% in surf zones and estuaries if diet composition and consumption rates remain similar to present conditions. To compensate for the decline in growth, juveniles may increase their consumption rates or consume more energetically rich prey, if available. If they are not able to compensate, their size at the end of the season may be reduced, which could reduce their overall survival. These results confirm that a small number of suyearling Chinook salmon use sandy beach surf zones, mostly adjacent to estuary mouths, where they experience growth conditions comparable to estuaries. My findings indicate that, in certain situations, juvenile Chinook salmon surf zone use can be influenced by surf zone growth conditions, while variation in growth rates are themselves most strongly influenced by variation in consumption rates in surf zones and estuaries. Predicted changes in coastal western North American climate will likely modify juvenile growth conditions in the next 50 years, and potentially reduce overall survival. Additional insights into the potential impacts of climate change on juvenile salmon will require estimates of changes in the composition, energetic quality and abundance of prey communities inhabiting coastal environments. Close

• 13. [Article] Evaluating "nuisance" beaver relocation as a tool to increase coho salmon habitat in the Alsea Basin of the central Oregon Coast Range

  Many stakeholders involved with stream restoration in the Pacific Northwest have discussed the potential benefits of using beaver dam construction activities (Castor canadensis) as a management tool to ...

  Citation × Citation Citation Abstract Copy Title: Evaluating "nuisance" beaver relocation as a tool to increase coho salmon habitat in the Alsea Basin of the central Oregon Coast Range Author: Petro, Vanessa M. Many stakeholders involved with stream restoration in the Pacific Northwest have discussed the potential benefits of using beaver dam construction activities (Castor canadensis) as a management tool to improve degraded stream habitat for anadromous salmon species. In addition, there has been growing interest in using nuisance beavers, primarily controlled by lethal methods, to alleviate human-wildlife conflict issues and subsequently improve Coho salmon (Oncorhynchus kisutch) rearing habitat. To understand the efficacy of using beavers as a stream restoration tool, I studied the movement, survival, and dam construction of nuisance beavers relocated to the Alsea Basin of the Oregon Coast Range. I trapped and transported 38 individuals to the nine release sites where dams constructed by beavers would benefit coho salmon productivity. All adult and sub-adult beavers were equipped with tail-mount transmitters. Beavers moved an average of 3.3 ± 0.2 (SE) stream km from release sites. The maximum distance moved from a release site was 29.2 stream km. Post-release movements did not differ significantly by age or sex. All radio-tagged individuals dispersed from their release sites. Survival 16 weeks post-release was 47%. Twelve of the radio-tagged beavers died within 90 days of release. Three cause-specific sources of mortality were identified including predation, natural causes, and human related. Mountain lions (Puma concolor) were responsible for the majority of predation based mortalities. Of the 38 nuisance beavers relocated, only five individuals contributed to the nine dams constructed post-release. Six of these dams were built by one male: female pair. All dams constructed by relocated beavers were ephemeral. In addition to monitoring post-release beaver responses, I also assessed the utility of pre-existing models to identify release site locations. Thus, I explored dam habitat relationships of extant and relocated colonies throughout the Alsea Basin. I determined the utility of locally developed spatial models was reasonable. Analysis revealed primary pool habitat and valley floor width variables as strong indicators of beaver dam sites. I conclude beaver relocation as a tool for stream restoration in the Alsea Basin may not offer an effective solution to lethal control measures and coho habitat enhancement due to low survival rates, unwanted movement and establishment outside of suitable release sites, and lack of dam construction. Title: Evaluating "nuisance" beaver relocation as a tool to increase coho salmon habitat in the Alsea Basin of the central Oregon Coast Range Author: Petro, Vanessa M. Many stakeholders involved with stream restoration in the Pacific Northwest have discussed the potential benefits of using beaver dam construction activities (Castor canadensis) as a management tool to improve degraded stream habitat for anadromous salmon species. In addition, there has been growing interest in using nuisance beavers, primarily controlled by lethal methods, to alleviate human-wildlife conflict issues and subsequently improve Coho salmon (Oncorhynchus kisutch) rearing habitat. To understand the efficacy of using beavers as a stream restoration tool, I studied the movement, survival, and dam construction of nuisance beavers relocated to the Alsea Basin of the Oregon Coast Range. I trapped and transported 38 individuals to the nine release sites where dams constructed by beavers would benefit coho salmon productivity. All adult and sub-adult beavers were equipped with tail-mount transmitters. Beavers moved an average of 3.3 ± 0.2 (SE) stream km from release sites. The maximum distance moved from a release site was 29.2 stream km. Post-release movements did not differ significantly by age or sex. All radio-tagged individuals dispersed from their release sites. Survival 16 weeks post-release was 47%. Twelve of the radio-tagged beavers died within 90 days of release. Three cause-specific sources of mortality were identified including predation, natural causes, and human related. Mountain lions (Puma concolor) were responsible for the majority of predation based mortalities. Of the 38 nuisance beavers relocated, only five individuals contributed to the nine dams constructed post-release. Six of these dams were built by one male: female pair. All dams constructed by relocated beavers were ephemeral. In addition to monitoring post-release beaver responses, I also assessed the utility of pre-existing models to identify release site locations. Thus, I explored dam habitat relationships of extant and relocated colonies throughout the Alsea Basin. I determined the utility of locally developed spatial models was reasonable. Analysis revealed primary pool habitat and valley floor width variables as strong indicators of beaver dam sites. I conclude beaver relocation as a tool for stream restoration in the Alsea Basin may not offer an effective solution to lethal control measures and coho habitat enhancement due to low survival rates, unwanted movement and establishment outside of suitable release sites, and lack of dam construction. Close

• 14. [Article] Policy Patterns across Riverscapes : Riparian Land Standards in the Oregon Coast Range

  Land management policies are ideas about nature projected onto the landscape. Culminations of social, economic, and scientific influences, these policies create standards affecting the function of ecological ...

  Citation × Citation Citation Abstract Copy Title: Policy Patterns across Riverscapes : Riparian Land Standards in the Oregon Coast Range Author: Boisjolie, Brett A. Land management policies are ideas about nature projected onto the landscape. Culminations of social, economic, and scientific influences, these policies create standards affecting the function of ecological systems. In the case of riparian lands in the Oregon Coast Range, policy requirements vary considerably across federal, state, and private land ownerships. Protective measures, such as the adoption of fixed buffer widths for maintaining vegetation, are intended to preserve natural processes important to instream habitat and water quality. However, the "policy landscape" of variable management standards can result in a fragmented approach to policy protection. This fragmentation complicates recovery efforts for threatened anadromous fish species such as coho salmon (Oncorhynchus kisutch), which require connectivity in suitable habitat segments throughout the river network. Taking a riverscape perspective, this study evaluates how aquatic protections vary across the river and its valley. To explore the patterns created from variable standard, the study first delineates categories of riparian management standards in the Oregon Coast Range. Using a geographic information system (GIS), these standards are mapped across the river networks of this region. The extent of each policy category is quantified in stream kilometers in order to evaluate policy efforts within and across the riverscapes of the Oregon Coast Range. A model of intrinsic potential for coho salmon habitat is used to identify the policy standards in stream reaches where flow, valley constraint, and channel gradient are appropriate for coho salmon habitat. A second model of the stream network (hydrography) is developed to evaluate how differences in stream delineation can influence policy patterns. Results indicate that riparian lands of the Oregon Coast Range are protected by variable standards for land management, including buffer widths that range from 0 to ~152m. In streams important to coho salmon, a lower proportion of the stream network was found to be protected by policy standards compared to the watershed and ESU scale. A lack of stream data central to policy application (fish use, stream flow duration, mean annual flow, water use) complicated our ability to attribute stream segments with specific land management requirements. The choice of hydrographic data was also found to influence the proportional extent of policies within riverscapes. This framework for assessing the spatial distribution of policies can provide insights for future studies of riverscape systems. Title: Policy Patterns across Riverscapes : Riparian Land Standards in the Oregon Coast Range Author: Boisjolie, Brett A. Land management policies are ideas about nature projected onto the landscape. Culminations of social, economic, and scientific influences, these policies create standards affecting the function of ecological systems. In the case of riparian lands in the Oregon Coast Range, policy requirements vary considerably across federal, state, and private land ownerships. Protective measures, such as the adoption of fixed buffer widths for maintaining vegetation, are intended to preserve natural processes important to instream habitat and water quality. However, the "policy landscape" of variable management standards can result in a fragmented approach to policy protection. This fragmentation complicates recovery efforts for threatened anadromous fish species such as coho salmon (Oncorhynchus kisutch), which require connectivity in suitable habitat segments throughout the river network. Taking a riverscape perspective, this study evaluates how aquatic protections vary across the river and its valley. To explore the patterns created from variable standard, the study first delineates categories of riparian management standards in the Oregon Coast Range. Using a geographic information system (GIS), these standards are mapped across the river networks of this region. The extent of each policy category is quantified in stream kilometers in order to evaluate policy efforts within and across the riverscapes of the Oregon Coast Range. A model of intrinsic potential for coho salmon habitat is used to identify the policy standards in stream reaches where flow, valley constraint, and channel gradient are appropriate for coho salmon habitat. A second model of the stream network (hydrography) is developed to evaluate how differences in stream delineation can influence policy patterns. Results indicate that riparian lands of the Oregon Coast Range are protected by variable standards for land management, including buffer widths that range from 0 to ~152m. In streams important to coho salmon, a lower proportion of the stream network was found to be protected by policy standards compared to the watershed and ESU scale. A lack of stream data central to policy application (fish use, stream flow duration, mean annual flow, water use) complicated our ability to attribute stream segments with specific land management requirements. The choice of hydrographic data was also found to influence the proportional extent of policies within riverscapes. This framework for assessing the spatial distribution of policies can provide insights for future studies of riverscape systems. Close

• 15. [Article] The influence of sex, migration distance, and latitude on expression of anadromy in Oncorhynchus mykiss

  The evolution and maintenance of multiple phenotypes within populations is an interesting evolutionary and ecological question, and is becoming increasingly important for the restoration and conservation ...

  Citation × Citation Citation Abstract Copy Title: The influence of sex, migration distance, and latitude on expression of anadromy in Oncorhynchus mykiss Author: Ohms, Haley The evolution and maintenance of multiple phenotypes within populations is an interesting evolutionary and ecological question, and is becoming increasingly important for the restoration and conservation of partially migratory species. In particular, why individuals adopt a particular phenotype has been the focus of numerous research efforts. In the partially migratory species Oncorhynchys mykiss (commonly known as steelhead and rainbow trout), I found that sex plays a strong role in whether an individual adopts an anadromous or resident life history. I observed a higher frequency of females adopting the anadromous tactic throughout the species range, and I attributed this to the relationship between body size and reproductive success in females that is not present in males. I did not observe differences in female bias along gradients of latitude or migration distance, and I hypothesize that while the frequency of tactics among populations may change along these gradients, either the change is equal between males and females, or an interaction of latitude and migration distance may control changes. I did not observe a corresponding male bias in the one resident population I sampled, and suggest that this is the result of a high frequency of residents in that population. I propose that sex ratios could be a valuable metric for measuring frequency of life-history tactics in populations and could facilitate the investigation into the controls of life-history tactics across broad scales. Title: The influence of sex, migration distance, and latitude on expression of anadromy in Oncorhynchus mykiss Author: Ohms, Haley The evolution and maintenance of multiple phenotypes within populations is an interesting evolutionary and ecological question, and is becoming increasingly important for the restoration and conservation of partially migratory species. In particular, why individuals adopt a particular phenotype has been the focus of numerous research efforts. In the partially migratory species Oncorhynchys mykiss (commonly known as steelhead and rainbow trout), I found that sex plays a strong role in whether an individual adopts an anadromous or resident life history. I observed a higher frequency of females adopting the anadromous tactic throughout the species range, and I attributed this to the relationship between body size and reproductive success in females that is not present in males. I did not observe differences in female bias along gradients of latitude or migration distance, and I hypothesize that while the frequency of tactics among populations may change along these gradients, either the change is equal between males and females, or an interaction of latitude and migration distance may control changes. I did not observe a corresponding male bias in the one resident population I sampled, and suggest that this is the result of a high frequency of residents in that population. I propose that sex ratios could be a valuable metric for measuring frequency of life-history tactics in populations and could facilitate the investigation into the controls of life-history tactics across broad scales. Close

• 16. [Article] The Umpqua River study : a joint report

  The species of anadromous fish entering the Umpqua River are: chinook salmon (spring, summer, and fall runs), silver salmon, steelhead trout (summer and winter runs), cutthroat trout, shad, striped bass ...

  Citation × Citation Citation Abstract Copy Title: The Umpqua River study : a joint report Author: Oregon. State Game Commission, Oregon. Fish Commission The species of anadromous fish entering the Umpqua River are: chinook salmon (spring, summer, and fall runs), silver salmon, steelhead trout (summer and winter runs), cutthroat trout, shad, striped bass and sturgeon. The winter steelhead, striped bass, and shad runs appear to be maintaining themselves. The runs of the other species appear to be in various stages of depletion. Major factors which have contributed to this depletion are: overfishing by the commercial fishery and more recently the sport fishery; and barriers such as log jams, "splash" damns, mill dam and hydroelectric developments. Other factors which may be of importance are: predation; unscreened gravity water diversions; water removal by pumping for irrigation, industrial, and municipal use; and fluctuations in water levels due to meterological conditions. The total run of spring chinook salmon into the Umpqua River in 1946 was estimated to be 4,400 fish of which approximately 935 were jacks. (fish under 20 inches). The fishing mortality of shad of commercial size, in the 1946 Umpqua River run, was estimated to be 50 percent. The total catch of the various species of salmon by the commercial and sport fisheries in 1946 were as follows: Under a proper management program, an average sustained yield of several times in the 1946 catches may be expected. During 1946, anglers spent at least $292,700 in this area while fishing for salmon and trout in the Umpqua River. The capital investment was estimated to be $2,402,500. The commercial fishery products of salmon steelhead, shad, striped bass, pilchards, and crabs taken from the Umpqua River in 1946 had a wholesale valur of $652,400. The capital investment was not determined. Title: The Umpqua River study : a joint report Author: Oregon. State Game Commission, Oregon. Fish Commission The species of anadromous fish entering the Umpqua River are: chinook salmon (spring, summer, and fall runs), silver salmon, steelhead trout (summer and winter runs), cutthroat trout, shad, striped bass and sturgeon. The winter steelhead, striped bass, and shad runs appear to be maintaining themselves. The runs of the other species appear to be in various stages of depletion. Major factors which have contributed to this depletion are: overfishing by the commercial fishery and more recently the sport fishery; and barriers such as log jams, "splash" damns, mill dam and hydroelectric developments. Other factors which may be of importance are: predation; unscreened gravity water diversions; water removal by pumping for irrigation, industrial, and municipal use; and fluctuations in water levels due to meterological conditions. The total run of spring chinook salmon into the Umpqua River in 1946 was estimated to be 4,400 fish of which approximately 935 were jacks. (fish under 20 inches). The fishing mortality of shad of commercial size, in the 1946 Umpqua River run, was estimated to be 50 percent. The total catch of the various species of salmon by the commercial and sport fisheries in 1946 were as follows: Under a proper management program, an average sustained yield of several times in the 1946 catches may be expected. During 1946, anglers spent at least $292,700 in this area while fishing for salmon and trout in the Umpqua River. The capital investment was estimated to be $2,402,500. The commercial fishery products of salmon steelhead, shad, striped bass, pilchards, and crabs taken from the Umpqua River in 1946 had a wholesale valur of $652,400. The capital investment was not determined. Close

• 17. [Article] Aspects of the early life history of juvenile salmonids in the Dungeness River Estuary

  The decline of many Pacific salmon stocks has stimulated interest in the early life history and habitat requirements of juvenile salmon. Although estuarine habitat associations of juvenile salmon have ...

  Citation × Citation Citation Abstract Copy Title: Aspects of the early life history of juvenile salmonids in the Dungeness River Estuary Author: Sather, Nichole K. The decline of many Pacific salmon stocks has stimulated interest in the early life history and habitat requirements of juvenile salmon. Although estuarine habitat associations of juvenile salmon have been investigated in many coastal areas of the eastern Pacific Ocean, until recently, little was known about juvenile salmonid ecology within the Straits of Juan de Fuca. During the Spring/Summer outmigration period in 2006 and 2007, I examined the early life history of the five species of anadromous salmon in the Dungeness River estuary on the north Olympic Peninsula, Washington. I sampled multiple spatial scales within several habitat types to characterize salmon distribution and habitat use. My results presented in this thesis are segregated into two components: 1) tidal marsh ecology of juvenile salmonids in the Dungeness River estuary, and 2) the landscape-scale distribution of juvenile salmonids within the Dungeness River estuary. I examined the population of juvenile salmonids within blind tidal sloughs near the vicinity of the Dungeness River delta. Salmonids were present within the tidal marshes throughout the entire outmigration period (e.g., March through July). Juvenile Chinook salmon (Oncorhynchus tshawytscha) were the most abundant salmonid species within the marshes. Based on the temporal distribution and size structure of juvenile Chinook salmon in the estuary I identified at least four life history types: 1) a fry strategy included a large pulse of fish emigrating from the river at a small size (e.g., 35-45mm FL) during late winter and early spring months; 2) the second group of fish was the least abundant group emigrating from the river from April through mid May at sizes ranging from 50-75mm FL; 3) the third group of migrants entered the estuary between from late spring through the summer months at larger sizes than the initial groups (e.g., 60-90mm FL); and 4) the final group of Chinook salmon included a stream-type yearling strategy. In addition to the four life history strategies identified for Chinook salmon, I detected at least three groups of chum salmon migrating into the estuary. These groups were distinguished by their size and timing of migration and are further described according to different rearing strategies. The distribution of juvenile salmonids was most strongly influenced by the degree of connectivity (i.e., distance) between the tidal marshes and the mouth of the Dungeness River. Habitat complexity and opportunity also governed the distribution of juvenile salmonids within the tidal marshes. I also sampled three regions of the estuary with a beach seine to investigate the nearshore distribution of juvenile salmonids within the Dungeness River estuary: the delta face, inner Bay, and outer Bay. Among the three regions, species composition was highly variable between 2006 and 2007. The most common salmonids encountered within the beach seine sites included Chinook salmon, chum salmon (O. kisutch), and pink salmon (O. gorbuscha). The relative abundance of salmonids was highest near the delta face and lowest within the outer bay area. The landscape-scale distribution and habitat use of juvenile salmonids within the Dungeness River estuary is largely influenced by ecosystem connectivity, but is also linked to biotic characteristics of the fish (e.g., life history type and fish size). Although the Dungeness includes hydrogeomorphic characteristics (e.g., steep river gradient, composition of sand spits in the estuary) unique to other Pacific Northwest watersheds, this system produces a variety of life history types comparable to other estuaries. Understanding the mechanisms that drive the distribution of juvenile salmonids within the Dungeness will supply local resource managers with a baseline with which to establish ecosystem restoration goals. Title: Aspects of the early life history of juvenile salmonids in the Dungeness River Estuary Author: Sather, Nichole K. The decline of many Pacific salmon stocks has stimulated interest in the early life history and habitat requirements of juvenile salmon. Although estuarine habitat associations of juvenile salmon have been investigated in many coastal areas of the eastern Pacific Ocean, until recently, little was known about juvenile salmonid ecology within the Straits of Juan de Fuca. During the Spring/Summer outmigration period in 2006 and 2007, I examined the early life history of the five species of anadromous salmon in the Dungeness River estuary on the north Olympic Peninsula, Washington. I sampled multiple spatial scales within several habitat types to characterize salmon distribution and habitat use. My results presented in this thesis are segregated into two components: 1) tidal marsh ecology of juvenile salmonids in the Dungeness River estuary, and 2) the landscape-scale distribution of juvenile salmonids within the Dungeness River estuary. I examined the population of juvenile salmonids within blind tidal sloughs near the vicinity of the Dungeness River delta. Salmonids were present within the tidal marshes throughout the entire outmigration period (e.g., March through July). Juvenile Chinook salmon (Oncorhynchus tshawytscha) were the most abundant salmonid species within the marshes. Based on the temporal distribution and size structure of juvenile Chinook salmon in the estuary I identified at least four life history types: 1) a fry strategy included a large pulse of fish emigrating from the river at a small size (e.g., 35-45mm FL) during late winter and early spring months; 2) the second group of fish was the least abundant group emigrating from the river from April through mid May at sizes ranging from 50-75mm FL; 3) the third group of migrants entered the estuary between from late spring through the summer months at larger sizes than the initial groups (e.g., 60-90mm FL); and 4) the final group of Chinook salmon included a stream-type yearling strategy. In addition to the four life history strategies identified for Chinook salmon, I detected at least three groups of chum salmon migrating into the estuary. These groups were distinguished by their size and timing of migration and are further described according to different rearing strategies. The distribution of juvenile salmonids was most strongly influenced by the degree of connectivity (i.e., distance) between the tidal marshes and the mouth of the Dungeness River. Habitat complexity and opportunity also governed the distribution of juvenile salmonids within the tidal marshes. I also sampled three regions of the estuary with a beach seine to investigate the nearshore distribution of juvenile salmonids within the Dungeness River estuary: the delta face, inner Bay, and outer Bay. Among the three regions, species composition was highly variable between 2006 and 2007. The most common salmonids encountered within the beach seine sites included Chinook salmon, chum salmon (O. kisutch), and pink salmon (O. gorbuscha). The relative abundance of salmonids was highest near the delta face and lowest within the outer bay area. The landscape-scale distribution and habitat use of juvenile salmonids within the Dungeness River estuary is largely influenced by ecosystem connectivity, but is also linked to biotic characteristics of the fish (e.g., life history type and fish size). Although the Dungeness includes hydrogeomorphic characteristics (e.g., steep river gradient, composition of sand spits in the estuary) unique to other Pacific Northwest watersheds, this system produces a variety of life history types comparable to other estuaries. Understanding the mechanisms that drive the distribution of juvenile salmonids within the Dungeness will supply local resource managers with a baseline with which to establish ecosystem restoration goals. Close

• 18. [Article] The future of the Columbia River salmon fisheries

  The salmon runs of the Columbia River constitute one of the most important natural resources of the states of Oregon and Washington. Thousands of people are dependent, wholly or in part, upon these resources ...

  Citation × Citation Citation Abstract Copy Title: The future of the Columbia River salmon fisheries Author: Oregon. Fish Commission, Rich, Willis H. (Willis Horton), 1885-1972 The salmon runs of the Columbia River constitute one of the most important natural resources of the states of Oregon and Washington. Thousands of people are dependent, wholly or in part, upon these resources for their livelihood; and their welfare is dependent upon the maintenance of the salmon runs. It is to the special interest of these people that the salmon supply be maintained at the level of maximum productivity and, indirectly, it is to the interest of all the citizens of these states that this be done. In recent years it has become increasingly evident that these resources, taken as. a whole, are decreasing in productivity and it has been frequently pointed out that this decrease is due primarily to the operation of two factors-- a very intensive fishery and a reduction in spawning and rearing areas due to the utilization of the water resources of the Columbia basin for other purposes. These influences, however, have affected the several species of salmon that are found in the Columbia River quite differently; the more valuable species, Chinooks and bluebacks, showing more evidence of depletion than do the other species, silver salmon, chums and steelhead. Title: The future of the Columbia River salmon fisheries Author: Oregon. Fish Commission, Rich, Willis H. (Willis Horton), 1885-1972 The salmon runs of the Columbia River constitute one of the most important natural resources of the states of Oregon and Washington. Thousands of people are dependent, wholly or in part, upon these resources for their livelihood; and their welfare is dependent upon the maintenance of the salmon runs. It is to the special interest of these people that the salmon supply be maintained at the level of maximum productivity and, indirectly, it is to the interest of all the citizens of these states that this be done. In recent years it has become increasingly evident that these resources, taken as. a whole, are decreasing in productivity and it has been frequently pointed out that this decrease is due primarily to the operation of two factors-- a very intensive fishery and a reduction in spawning and rearing areas due to the utilization of the water resources of the Columbia basin for other purposes. These influences, however, have affected the several species of salmon that are found in the Columbia River quite differently; the more valuable species, Chinooks and bluebacks, showing more evidence of depletion than do the other species, silver salmon, chums and steelhead. Close

• 19. [Article] Fish Inves 1947 pt 1.pdf

  Citation × Citation Citation Abstract Copy Title: Fish Inves 1947 pt 1.pdf Author: Oregon. State Game Commission Title: Fish Inves 1947 pt 1.pdf Author: Oregon. State Game Commission Close

• 20. [Article] Fish Invest 1946 pt 2.pdf

  Citation × Citation Citation Abstract Copy Title: Fish Invest 1946 pt 2.pdf Author: Oregon. State Game Commission Title: Fish Invest 1946 pt 2.pdf Author: Oregon. State Game Commission Close

• 21. [Article] Variability in expression of anadromy by female Oncorhynchus mykiss within a river network

  We described and predicted spatial variation in marine migration (anadromy) of female Oncorhynchus mykiss in the John Day River watershed, Oregon. We collected 149 juvenile O. mykiss across 72 sites and ...

  Citation × Citation Citation Abstract Copy Title: Variability in expression of anadromy by female Oncorhynchus mykiss within a river network Author: Zimmerman, Christian E., Jordan, Chris E., McMillan, John R., Mills, Justin S., Dunham, Jason B., Reeves, Gordon H. We described and predicted spatial variation in marine migration (anadromy) of female Oncorhynchus mykiss in the John Day River watershed, Oregon. We collected 149 juvenile O. mykiss across 72 sites and identified locations used by anadromous females by assigning maternal origin (anadromous versus non-anadromous) to each juvenile. These assignments used comparisons of strontium to calcium ratios in otolith primordia and freshwater growth regions to indicate maternal origin. We used logistic regression to predict probability of anadromy in relation to mean annual stream runoff using data from a subset of individuals. This model correctly predicted anadromy in a second sample of individuals with a moderate level of accuracy (e.g., 68% correctly predicted with a 0.5 classification threshold). Residuals from the models were not spatially autocorrelated, suggesting that remaining variability in the expression of anadromy was due to localized influences, as opposed to broad-scale gradients unrelated to mean annual stream runoff. These results are important for the management of O. mykiss because anadromous individuals (steelhead) within the John Day River watershed are listed as a threatened species, and it is difficult to discern juvenile steelhead from non-anadromous individuals (rainbow trout) in the field. Our results provide a broad-scale description and prediction of locations supporting anadromy, and new insight for habitat restoration, monitoring, and research to better manage and understand the expression of anadromy in O. mykiss. Title: Variability in expression of anadromy by female Oncorhynchus mykiss within a river network Author: Zimmerman, Christian E., Jordan, Chris E., McMillan, John R., Mills, Justin S., Dunham, Jason B., Reeves, Gordon H. We described and predicted spatial variation in marine migration (anadromy) of female Oncorhynchus mykiss in the John Day River watershed, Oregon. We collected 149 juvenile O. mykiss across 72 sites and identified locations used by anadromous females by assigning maternal origin (anadromous versus non-anadromous) to each juvenile. These assignments used comparisons of strontium to calcium ratios in otolith primordia and freshwater growth regions to indicate maternal origin. We used logistic regression to predict probability of anadromy in relation to mean annual stream runoff using data from a subset of individuals. This model correctly predicted anadromy in a second sample of individuals with a moderate level of accuracy (e.g., 68% correctly predicted with a 0.5 classification threshold). Residuals from the models were not spatially autocorrelated, suggesting that remaining variability in the expression of anadromy was due to localized influences, as opposed to broad-scale gradients unrelated to mean annual stream runoff. These results are important for the management of O. mykiss because anadromous individuals (steelhead) within the John Day River watershed are listed as a threatened species, and it is difficult to discern juvenile steelhead from non-anadromous individuals (rainbow trout) in the field. Our results provide a broad-scale description and prediction of locations supporting anadromy, and new insight for habitat restoration, monitoring, and research to better manage and understand the expression of anadromy in O. mykiss. Close

• 22. [Article] The effects on salmon populations of the partial elimination of fixed fishing gear on the Columbia River in 1935

  1. Six major types of commercial gear have been used to take salmon and steelhead on the Columbia River; namely, gill nets, set nets, seines, traps, fish wheels and dip nets. 2. The five important commercial ...

  Citation × Citation Citation Abstract Copy Title: The effects on salmon populations of the partial elimination of fixed fishing gear on the Columbia River in 1935 Author: Johnson, Donald R., Chapman, Wilbert McLeod, 1910-1970, Oregon. Fish Commission, Schoning, Robert W. 1. Six major types of commercial gear have been used to take salmon and steelhead on the Columbia River; namely, gill nets, set nets, seines, traps, fish wheels and dip nets. 2. The five important commercial species in the Columbia River are chinook, silver, blueback and chum salmon and steelhead trout. 3. In 1927 fish wheels were eliminated in Oregon. In 1935 fish wheels, traps, seines and set nets were eliminated in Washington. 4. The effect of the gear change in Washington in 1935 has been to increase the take in other gears, while the previous trend of total landings has not changed appreciably. 5. There is no conclusive evidence that the elimination of fixed gear in Washington increased the escapement of salmon and steelhead. 6. There is no conclusive evidence that any gear is injurious to the run except by subtracting fish from the run. 7. Measures designed to increase the escapement should apply to all gears and cover the entire life history of each species. Title: The effects on salmon populations of the partial elimination of fixed fishing gear on the Columbia River in 1935 Author: Johnson, Donald R., Chapman, Wilbert McLeod, 1910-1970, Oregon. Fish Commission, Schoning, Robert W. 1. Six major types of commercial gear have been used to take salmon and steelhead on the Columbia River; namely, gill nets, set nets, seines, traps, fish wheels and dip nets. 2. The five important commercial species in the Columbia River are chinook, silver, blueback and chum salmon and steelhead trout. 3. In 1927 fish wheels were eliminated in Oregon. In 1935 fish wheels, traps, seines and set nets were eliminated in Washington. 4. The effect of the gear change in Washington in 1935 has been to increase the take in other gears, while the previous trend of total landings has not changed appreciably. 5. There is no conclusive evidence that the elimination of fixed gear in Washington increased the escapement of salmon and steelhead. 6. There is no conclusive evidence that any gear is injurious to the run except by subtracting fish from the run. 7. Measures designed to increase the escapement should apply to all gears and cover the entire life history of each species. Close

• 23. [Article] A report of fisheries investigations in Oregon coastal streams south of the Columbia River and exclusive of the Umpqua River

  This report is a summarization of investigational work that has been completed since 1941 by the Oregon State Game Commission on the coast streams south of the Columbia River, exclusive of the Umqpua ...

  Citation × Citation Citation Abstract Copy Title: A report of fisheries investigations in Oregon coastal streams south of the Columbia River and exclusive of the Umpqua River Author: Oregon. State Game Commission This report is a summarization of investigational work that has been completed since 1941 by the Oregon State Game Commission on the coast streams south of the Columbia River, exclusive of the Umqpua and its tributaries. The Umpqua river system was covered in another report prepared jointly by biologists of the Fish Commission of Oregon and the Oregon State Game Commission. It is hoped that the facts presented here will help in an understanding of the problems involved and serve to make known the critically low levels of runs of anadromous fishes in the coastal streams. The report covers general observations, fish population studies, barriers and stream improvements, salvage problems, hatcheries and economic evaluations. Title: A report of fisheries investigations in Oregon coastal streams south of the Columbia River and exclusive of the Umpqua River Author: Oregon. State Game Commission This report is a summarization of investigational work that has been completed since 1941 by the Oregon State Game Commission on the coast streams south of the Columbia River, exclusive of the Umqpua and its tributaries. The Umpqua river system was covered in another report prepared jointly by biologists of the Fish Commission of Oregon and the Oregon State Game Commission. It is hoped that the facts presented here will help in an understanding of the problems involved and serve to make known the critically low levels of runs of anadromous fishes in the coastal streams. The report covers general observations, fish population studies, barriers and stream improvements, salvage problems, hatcheries and economic evaluations. Close

• 24. [Article] Pool habitat characteristics and juvenile anadromous salmonids in two Oregon coastal streams

  Relations between the diversity of juvenile anadromous salmonids and pool features were examined in a managed and a pristine watershed in Oregon during the summer of 1990. There were no differences (p>0.05) ...

  Citation × Citation Citation Abstract Copy Title: Pool habitat characteristics and juvenile anadromous salmonids in two Oregon coastal streams Author: Haapala, Deborah L. Relations between the diversity of juvenile anadromous salmonids and pool features were examined in a managed and a pristine watershed in Oregon during the summer of 1990. There were no differences (p>0.05) in pool depth, velocity or pool wood volumes between streams. However, the pristine system had twice the number of pools within similar lineal distances. Pools in the pristine system also had larger substrates (percent dominant within pools) and smaller pool area (p=0.01). Fish diversity was found to be greater in pools in the pristine system than in the managed system using the Simpson's Diversity and Shannon Evenness indicies (p=0.01). The Shannon-Wiener Diversity index did not show any differences between streams. The difference in assemblage diversity was due to differences in relative abundance and not species richness. Relative abundance of juvenile steelhead and cutthroat trout and coho salmon was more even in Cummins Creek, the pristine system, than in Cape Creek, the managed system. Relative abundance of coho increased in the managed system possibly due to a change in pool habitat characteristics, whose conditions favored coho salmon, but this relationship was not clear. This study emphasizes the importance of assessing communities of juvenile anadromous salmonids as opposed to studies involving a single species. Past land management activities have focused upon single species' with regards to a particular habitat component, which has decreased biodiversity and changed stream habitat characteristics through cumulative effects. Resource managers should examine interactions between habitat characteristics and salmonid communities in order to maintain biological diversity or risk creating favored habitat for a single species within stream systems. Title: Pool habitat characteristics and juvenile anadromous salmonids in two Oregon coastal streams Author: Haapala, Deborah L. Relations between the diversity of juvenile anadromous salmonids and pool features were examined in a managed and a pristine watershed in Oregon during the summer of 1990. There were no differences (p>0.05) in pool depth, velocity or pool wood volumes between streams. However, the pristine system had twice the number of pools within similar lineal distances. Pools in the pristine system also had larger substrates (percent dominant within pools) and smaller pool area (p=0.01). Fish diversity was found to be greater in pools in the pristine system than in the managed system using the Simpson's Diversity and Shannon Evenness indicies (p=0.01). The Shannon-Wiener Diversity index did not show any differences between streams. The difference in assemblage diversity was due to differences in relative abundance and not species richness. Relative abundance of juvenile steelhead and cutthroat trout and coho salmon was more even in Cummins Creek, the pristine system, than in Cape Creek, the managed system. Relative abundance of coho increased in the managed system possibly due to a change in pool habitat characteristics, whose conditions favored coho salmon, but this relationship was not clear. This study emphasizes the importance of assessing communities of juvenile anadromous salmonids as opposed to studies involving a single species. Past land management activities have focused upon single species' with regards to a particular habitat component, which has decreased biodiversity and changed stream habitat characteristics through cumulative effects. Resource managers should examine interactions between habitat characteristics and salmonid communities in order to maintain biological diversity or risk creating favored habitat for a single species within stream systems. Close

• 25. [Article] Description and prediction of broad-scale spatial variability in expression of anadromy in female Oncorhynchus mykiss in the John Day River, Oregon, USA

  Partial migration is a common migratory behavior where some individuals in a population migrate and others do not. Patterns of partial migration can vary dramatically, especially for species that inhabit ...

  Citation × Citation Citation Abstract Copy Title: Description and prediction of broad-scale spatial variability in expression of anadromy in female Oncorhynchus mykiss in the John Day River, Oregon, USA Author: Mills, Justin S. Partial migration is a common migratory behavior where some individuals in a population migrate and others do not. Patterns of partial migration can vary dramatically, especially for species that inhabit a wide range of environments. I described and predicted spatial variation in marine migration (anadromy) of female Oncorhynchus mykiss (steelhead and rainbow trout) in the John Day River catchment basin, Oregon. This large catchment (20,500 km2), where O. mykiss is known to exhibit partial migration, encompasses a broad range of environmental conditions. I collected 149 juvenile O. mykiss across 72 sites and identified locations used by anadromous females by assigning maternal origin (anadromous versus non-anadromous) to each juvenile. These assignments used comparisons of strontium to calcium ratios in otolith primordia and freshwater growth regions to indicate maternal origin. Individuals within sites were more likely to have the same maternal origin than expected at random (χ2 test, P < 0.001). I used logistic regression to predict probability of anadromy among sites in relation to stream size (as indicated by mean annual runoff). Stream size and maternal origin for single fish collected from 69 sites were used to construct a predictive model. I examined the ability of this model to predict new individuals at 47 of these sites (where a second fish was collected) using a variety of diagnostics, including kappa statistics and receiver operating characteristic curves. The model predicted anadromy in this second set of individuals with a moderate level of accuracy (e.g. 68% correctly predicted with a 0.5 classification threshold). Residuals from the models were not spatially autocorrelated, as indicated by Mantel tests. This result also suggested that remaining variability in the expression of anadromy was due to localized influences, as opposed to broad-scale gradients unrelated to stream size. The importance of stream size implies that occurrence of migratory females was related to environmental variability, but stream size is an indicator of many potential processes, and I was unable to identify a specific likely mechanism. However, I was able to demonstrate 1) that it is possible to predict the probability of anadromy across broad environmental gradients, and 2) the validity of a sampling approach that minimizes the need for sacrificial sampling of individuals. These results are important for the management of O. mykiss because anadromous individuals (steelhead) within the John Day River watershed are listed as a threatened species, and it is difficult to discern steelhead from non-anadromous individuals (rainbow trout). My results provide managers with the first broad-scale description and prediction of locations supporting anadromy, and provide a broad template that can guide future habitat restoration, monitoring, and research to better manage and understand the expression of anadromy in O. mykiss. Title: Description and prediction of broad-scale spatial variability in expression of anadromy in female Oncorhynchus mykiss in the John Day River, Oregon, USA Author: Mills, Justin S. Partial migration is a common migratory behavior where some individuals in a population migrate and others do not. Patterns of partial migration can vary dramatically, especially for species that inhabit a wide range of environments. I described and predicted spatial variation in marine migration (anadromy) of female Oncorhynchus mykiss (steelhead and rainbow trout) in the John Day River catchment basin, Oregon. This large catchment (20,500 km2), where O. mykiss is known to exhibit partial migration, encompasses a broad range of environmental conditions. I collected 149 juvenile O. mykiss across 72 sites and identified locations used by anadromous females by assigning maternal origin (anadromous versus non-anadromous) to each juvenile. These assignments used comparisons of strontium to calcium ratios in otolith primordia and freshwater growth regions to indicate maternal origin. Individuals within sites were more likely to have the same maternal origin than expected at random (χ2 test, P < 0.001). I used logistic regression to predict probability of anadromy among sites in relation to stream size (as indicated by mean annual runoff). Stream size and maternal origin for single fish collected from 69 sites were used to construct a predictive model. I examined the ability of this model to predict new individuals at 47 of these sites (where a second fish was collected) using a variety of diagnostics, including kappa statistics and receiver operating characteristic curves. The model predicted anadromy in this second set of individuals with a moderate level of accuracy (e.g. 68% correctly predicted with a 0.5 classification threshold). Residuals from the models were not spatially autocorrelated, as indicated by Mantel tests. This result also suggested that remaining variability in the expression of anadromy was due to localized influences, as opposed to broad-scale gradients unrelated to stream size. The importance of stream size implies that occurrence of migratory females was related to environmental variability, but stream size is an indicator of many potential processes, and I was unable to identify a specific likely mechanism. However, I was able to demonstrate 1) that it is possible to predict the probability of anadromy across broad environmental gradients, and 2) the validity of a sampling approach that minimizes the need for sacrificial sampling of individuals. These results are important for the management of O. mykiss because anadromous individuals (steelhead) within the John Day River watershed are listed as a threatened species, and it is difficult to discern steelhead from non-anadromous individuals (rainbow trout). My results provide managers with the first broad-scale description and prediction of locations supporting anadromy, and provide a broad template that can guide future habitat restoration, monitoring, and research to better manage and understand the expression of anadromy in O. mykiss. Close

• 26. [Article] Predicted Redistribution of Ceratomyxa shasta Genotypes with Salmonid Passage in the Deschutes River, Oregon

  This is the publisher’s final pdf. The published article is copyrighted by Taylor & Francis and can be found at: http://www.tandf.co.uk/journals/.

  Citation × Citation Citation Abstract Copy Title: Predicted Redistribution of Ceratomyxa shasta Genotypes with Salmonid Passage in the Deschutes River, Oregon Author: Bartholomew, Jerri L., Stinson, Matthew E. T. This is the publisher’s final pdf. The published article is copyrighted by Taylor & Francis and can be found at: http://www.tandf.co.uk/journals/. Title: Predicted Redistribution of Ceratomyxa shasta Genotypes with Salmonid Passage in the Deschutes River, Oregon Author: Bartholomew, Jerri L., Stinson, Matthew E. T. This is the publisher’s final pdf. The published article is copyrighted by Taylor & Francis and can be found at: http://www.tandf.co.uk/journals/. Close

• 27. [Article] A study of Myxobolus cerebralis in the Lostine River, Oregon : epizootiology, distribution and implications for resident and anadromous salmonids

  Myxobolus cerebralis, a myxozoan parasite that infects almost all species of wild and cultured salmonids, was first identified in northeastern Oregon in the Lostine River in 1986. Fish that are heavily ...

  Citation × Citation Citation Abstract Copy Title: A study of Myxobolus cerebralis in the Lostine River, Oregon : epizootiology, distribution and implications for resident and anadromous salmonids Author: Sandell, Todd A. Myxobolus cerebralis, a myxozoan parasite that infects almost all species of wild and cultured salmonids, was first identified in northeastern Oregon in the Lostine River in 1986. Fish that are heavily infected with M. cerebralis develop whirling disease, which was determined to be the cause of catastrophic declines in rainbow trout populations in Montana and Colorado in 1996. There have been no reports of severe salmonid population declines attributed to whirling disease in Oregon, although many populations are at levels that warrant protective listing under the Endangered Species Act. This study was undertaken to I) establish the distribution and seasonal prevalence of M. cerebralis infection in the Lostine River, 2) investigate the epizootiological factors influencing the occurrence of whirling disease in this river, 3) determine the potential impact of the parasite on chinook salmon, and 4) evaluate the suitability of a non-lethal tissue sampling protocol for the detection of M. cerebralis in juvenile and adult salmonids for use in monitoring the effects and distribution of the parasite in protected salmonid populations. Results of this study indicate that the parasite is present throughout the main stem of the Lostine River during all seasons, and that rainbow and anadromous steelhead trout fry emerging in the lower river are most likely to become infected and develop whirling disease. Chinook salmon were shown to be relatively resistant to M. cerebralis at low levels of exposure, but this effect was less evident when the parasite exposure increased. The non-lethal sampling protocol was successful in detecting M. cerebralis in juvenile salmonids but not in adults, where it is most needed. The information gathered during these studies was then correlated with the life histories of resident and anadromous salmonids in the Lostine River to determine the potential impact of M. cerebralis On salmonid populations in this region. Title: A study of Myxobolus cerebralis in the Lostine River, Oregon : epizootiology, distribution and implications for resident and anadromous salmonids Author: Sandell, Todd A. Myxobolus cerebralis, a myxozoan parasite that infects almost all species of wild and cultured salmonids, was first identified in northeastern Oregon in the Lostine River in 1986. Fish that are heavily infected with M. cerebralis develop whirling disease, which was determined to be the cause of catastrophic declines in rainbow trout populations in Montana and Colorado in 1996. There have been no reports of severe salmonid population declines attributed to whirling disease in Oregon, although many populations are at levels that warrant protective listing under the Endangered Species Act. This study was undertaken to I) establish the distribution and seasonal prevalence of M. cerebralis infection in the Lostine River, 2) investigate the epizootiological factors influencing the occurrence of whirling disease in this river, 3) determine the potential impact of the parasite on chinook salmon, and 4) evaluate the suitability of a non-lethal tissue sampling protocol for the detection of M. cerebralis in juvenile and adult salmonids for use in monitoring the effects and distribution of the parasite in protected salmonid populations. Results of this study indicate that the parasite is present throughout the main stem of the Lostine River during all seasons, and that rainbow and anadromous steelhead trout fry emerging in the lower river are most likely to become infected and develop whirling disease. Chinook salmon were shown to be relatively resistant to M. cerebralis at low levels of exposure, but this effect was less evident when the parasite exposure increased. The non-lethal sampling protocol was successful in detecting M. cerebralis in juvenile salmonids but not in adults, where it is most needed. The information gathered during these studies was then correlated with the life histories of resident and anadromous salmonids in the Lostine River to determine the potential impact of M. cerebralis On salmonid populations in this region. Close

• 28. [Article] The present state of the Columbia River salmon resources

  Reprinted from Proceedings of the Sixth Pacific Science Congress, University of California Press, Berkeley, California, 1941.

  Citation × Citation Citation Abstract Copy Title: The present state of the Columbia River salmon resources Author: Oregon. Fish Commission. Research Dept., Rich, Willis H. (Willis Horton), 1885-1972 Reprinted from Proceedings of the Sixth Pacific Science Congress, University of California Press, Berkeley, California, 1941. Title: The present state of the Columbia River salmon resources Author: Oregon. Fish Commission. Research Dept., Rich, Willis H. (Willis Horton), 1885-1972 Reprinted from Proceedings of the Sixth Pacific Science Congress, University of California Press, Berkeley, California, 1941. Close

• 29. [Article] The salmon runs of the Columbia River in 1938

  Exceptional data are available for the study of the salmon runs of the Columbia River in 1938. Detailed figures on catch were supplied by Oregon and Washington in such form that they could readily be combined ...

  Citation × Citation Citation Abstract Copy Title: The salmon runs of the Columbia River in 1938 Author: Oregon. Fish Commission, Rich, Willis H. (Willis Horton), 1885-1972 Exceptional data are available for the study of the salmon runs of the Columbia River in 1938. Detailed figures on catch were supplied by Oregon and Washington in such form that they could readily be combined with the counts at Bonneville Dam to provide a basis for estimating the escapement. Tables show the catch of each species for each week in each of six zones, and the counts at Bonneville and Rock Island dams. The general course of the run of each species is shown. The numbers of fish bound for the spawning grounds above Rock Island Dam are estimated as follows: Chinook salmon entering Columbia River before May 1, 4 percent; during May, 6 percent; June and July, 15 percent; and August to December, 1 percent. Blueback salmon entering the river during the above periods, 40 percent. Steelhead trout entering the river during June to September, 1 percent; during the rest of the year, 10 percent. Fishing intensities are shown by escapement to catch ratios. Percentages of chinook salmon escapement are less than 15 during May; 17 during June and July; and 33 during the remainder of the year. The June and July runs are now greatly depleted, and an important part of these runs spawns above Rock Island Dam. The blueback salmon escapement is about 20 percent, and of steelhead trout about 33 percent. Weekly and seasonal closed periods are shown to be almost entirely ineffective for increasing the spawning escapement. Exploitation is further increased by the intensive troll fishery conducted from Monterey Bay to southeastern Alaska. Chinook salmon are also subjected to a sport fishery of considerable importance. Main runs of salmon to the Columbia River are practically unprotected and are fished with destructive intensity. Title: The salmon runs of the Columbia River in 1938 Author: Oregon. Fish Commission, Rich, Willis H. (Willis Horton), 1885-1972 Exceptional data are available for the study of the salmon runs of the Columbia River in 1938. Detailed figures on catch were supplied by Oregon and Washington in such form that they could readily be combined with the counts at Bonneville Dam to provide a basis for estimating the escapement. Tables show the catch of each species for each week in each of six zones, and the counts at Bonneville and Rock Island dams. The general course of the run of each species is shown. The numbers of fish bound for the spawning grounds above Rock Island Dam are estimated as follows: Chinook salmon entering Columbia River before May 1, 4 percent; during May, 6 percent; June and July, 15 percent; and August to December, 1 percent. Blueback salmon entering the river during the above periods, 40 percent. Steelhead trout entering the river during June to September, 1 percent; during the rest of the year, 10 percent. Fishing intensities are shown by escapement to catch ratios. Percentages of chinook salmon escapement are less than 15 during May; 17 during June and July; and 33 during the remainder of the year. The June and July runs are now greatly depleted, and an important part of these runs spawns above Rock Island Dam. The blueback salmon escapement is about 20 percent, and of steelhead trout about 33 percent. Weekly and seasonal closed periods are shown to be almost entirely ineffective for increasing the spawning escapement. Exploitation is further increased by the intensive troll fishery conducted from Monterey Bay to southeastern Alaska. Chinook salmon are also subjected to a sport fishery of considerable importance. Main runs of salmon to the Columbia River are practically unprotected and are fished with destructive intensity. Close

• 30. [Article] Siletz Bay National Fish and Wildlife Refuge : environmental assessment

  "The Service is proposing to establish the Siletz Bay National Fish and Wildlife Refuge by initially accepting a donation of 46 acres of salt marsh, acquiring up to 793 acres of land, and cooperatively ...

  Citation × Citation Citation Abstract Copy Title: Siletz Bay National Fish and Wildlife Refuge : environmental assessment Author: U.S. Fish and Wildlife Service. Region 1, Harrison, Ben "The Service is proposing to establish the Siletz Bay National Fish and Wildlife Refuge by initially accepting a donation of 46 acres of salt marsh, acquiring up to 793 acres of land, and cooperatively managing 1,060 acres of tidelands with the State of Oregon... The primary need for the Service acquisition is to preserve diverse coastal wetland habitats and upland buffers for a variety of waterfowl, shorebirds, marine mammals, endangered species, raptors, songbirds, anadromous and resident fish, and other wildlife." (p.2) The report describes the area and discusses potential issues. It includes maps, an aerial photograph, and a species list. Title: Siletz Bay National Fish and Wildlife Refuge : environmental assessment Author: U.S. Fish and Wildlife Service. Region 1, Harrison, Ben "The Service is proposing to establish the Siletz Bay National Fish and Wildlife Refuge by initially accepting a donation of 46 acres of salt marsh, acquiring up to 793 acres of land, and cooperatively managing 1,060 acres of tidelands with the State of Oregon... The primary need for the Service acquisition is to preserve diverse coastal wetland habitats and upland buffers for a variety of waterfowl, shorebirds, marine mammals, endangered species, raptors, songbirds, anadromous and resident fish, and other wildlife." (p.2) The report describes the area and discusses potential issues. It includes maps, an aerial photograph, and a species list. Close

• 31. [Article] The Scientific Basis for Artificial Propagation in the Recovery of Wild Anadromous Salmonids in Oregon

  This report was prepared by the Independent Multidisciplinary Science Team of the Oregon Plan for Salmon and Watersheds

  Citation × Citation Citation Abstract Copy Title: The Scientific Basis for Artificial Propagation in the Recovery of Wild Anadromous Salmonids in Oregon Author: Independent Multidisciplinary Science Team (Or.) This report was prepared by the Independent Multidisciplinary Science Team of the Oregon Plan for Salmon and Watersheds Title: The Scientific Basis for Artificial Propagation in the Recovery of Wild Anadromous Salmonids in Oregon Author: Independent Multidisciplinary Science Team (Or.) This report was prepared by the Independent Multidisciplinary Science Team of the Oregon Plan for Salmon and Watersheds Close

• 32. [Article] Differential use of salmon by vertebrate consumers: implications for conservation

  This is the publisher’s final pdf. The published article is copyrighted by the author(s) and published by PeerJ. The published article can be found at: https://peerj.com/

  Citation × Citation Citation Abstract Copy Title: Differential use of salmon by vertebrate consumers: implications for conservation Author: Allen, Jennifer M., Wilmers, Christopher C., Wheat, Rachel E., Levi, Taal This is the publisher’s final pdf. The published article is copyrighted by the author(s) and published by PeerJ. The published article can be found at: https://peerj.com/ Title: Differential use of salmon by vertebrate consumers: implications for conservation Author: Allen, Jennifer M., Wilmers, Christopher C., Wheat, Rachel E., Levi, Taal This is the publisher’s final pdf. The published article is copyrighted by the author(s) and published by PeerJ. The published article can be found at: https://peerj.com/ Close

• 33. [Article] Habitat Improvement Project, No. 17

  "Fall Creek Reservoir is a unit of the comprehensive plan of the U. S. Army Corps of Engineers for flood control and multiple use of the water resources of the upper Willamette Basin. The reservoir is ...

  Citation × Citation Citation Abstract Copy Title: Habitat Improvement Project, No. 17 Author: Swan, Ralph L., Oregon. State Game Commission. Fishery Division "Fall Creek Reservoir is a unit of the comprehensive plan of the U. S. Army Corps of Engineers for flood control and multiple use of the water resources of the upper Willamette Basin. The reservoir is located on Big Fall Creek, nineteen miles southeast of Eugene in Lane County...Undesirable fish species that compete with anadromous fish were to be eliminated from the watershed. Rough fish problems could conceivably make expensive fish facilities useless by depleting runs...Funds for the project were provided to the Oregon State Game Commission by the U. S. Army Corps of Engineers." (Abstract from Author) Title: Habitat Improvement Project, No. 17 Author: Swan, Ralph L., Oregon. State Game Commission. Fishery Division "Fall Creek Reservoir is a unit of the comprehensive plan of the U. S. Army Corps of Engineers for flood control and multiple use of the water resources of the upper Willamette Basin. The reservoir is located on Big Fall Creek, nineteen miles southeast of Eugene in Lane County...Undesirable fish species that compete with anadromous fish were to be eliminated from the watershed. Rough fish problems could conceivably make expensive fish facilities useless by depleting runs...Funds for the project were provided to the Oregon State Game Commission by the U. S. Army Corps of Engineers." (Abstract from Author) Close

• 34. [Article] An application of the control chart method to the analysis of fisheries data

  In a paper now in press as a Bulletin of the U. S. Fish and Wildlife Services the writer has discussed the downward trend of the catch of Columbia River Chinook salmon since 1920 and has stated that the ...

  Citation × Citation Citation Abstract Copy Title: An application of the control chart method to the analysis of fisheries data Author: Oregon. Fish Commission, Rich, Willis H. (Willis Horton), 1885-1972 In a paper now in press as a Bulletin of the U. S. Fish and Wildlife Services the writer has discussed the downward trend of the catch of Columbia River Chinook salmon since 1920 and has stated that the decline is doubtless an indication that the runs of this species are seriously depleted. Recently the same data have been studied by means of the control chart method and the results show so clearly that the productivity has dropped to a new and much lower level that it has seemed worth while to present a brief note amplifying the discussion in the bulletin. Title: An application of the control chart method to the analysis of fisheries data Author: Oregon. Fish Commission, Rich, Willis H. (Willis Horton), 1885-1972 In a paper now in press as a Bulletin of the U. S. Fish and Wildlife Services the writer has discussed the downward trend of the catch of Columbia River Chinook salmon since 1920 and has stated that the decline is doubtless an indication that the runs of this species are seriously depleted. Recently the same data have been studied by means of the control chart method and the results show so clearly that the productivity has dropped to a new and much lower level that it has seemed worth while to present a brief note amplifying the discussion in the bulletin. Close

• 35. [Article] Bycatch Avoidance Programs in Two New England (USA) Fisheries

  We conducted similar bycatch avoidance programs in two New England fisheries: yellowtail flounder bycatch in the sea scallop closed area fisheries and river herring bycatch in the Atlantic herring and ...

  Citation × Citation Citation Abstract Copy Title: Bycatch Avoidance Programs in Two New England (USA) Fisheries Author: Bethoney, David, DeCelles, Greg, Stokesbury, Kevin, Cadrin, Steven, O'Keefe, Catherine, Georgianna, Daniel We conducted similar bycatch avoidance programs in two New England fisheries: yellowtail flounder bycatch in the sea scallop closed area fisheries and river herring bycatch in the Atlantic herring and mackerel fishery. These species have different behavioral properties: sea scallops are sessile; yellowtail flounder are demersal with limited but unknown range. Herring and mackerel are pelagic species with wide spatial and temporal ranges; river herring are anadromous. Management actions also differed between the two fisheries. The sea scallop closed area fisheries were limited in time and space with hard quotas for each species. The herring and mackerel fishery was limited in large scale areas by hard quotas, without quotas for river herring. Fishermen were motivated to avoid river herring by strong conservation pressure, river herring are listed as a species of concern and under review for endangered species by the U.S. agencies, which designation would effectively close the Atlantic herring fishery. In the scallop fisheries, we mapped and updated densities for target and bycatch stocks through real-time monitoring and communicated hotspots with captains at sea. In the herring and mackerel fishery we identified densities through port sampling and communicated hotspots to captains at sea. We also reported environmental factors of river herring density such as depth to captains. Percent participation exceeded 75% in each fisheries with evidence of avoidance behavior in both fisheries. Yellowtail bycatch was sharply reduced in the scallop closed area fisheries with some reduction of river herring bycatch in the herring and mackerel fisheries. Title: Bycatch Avoidance Programs in Two New England (USA) Fisheries Author: Bethoney, David, DeCelles, Greg, Stokesbury, Kevin, Cadrin, Steven, O'Keefe, Catherine, Georgianna, Daniel We conducted similar bycatch avoidance programs in two New England fisheries: yellowtail flounder bycatch in the sea scallop closed area fisheries and river herring bycatch in the Atlantic herring and mackerel fishery. These species have different behavioral properties: sea scallops are sessile; yellowtail flounder are demersal with limited but unknown range. Herring and mackerel are pelagic species with wide spatial and temporal ranges; river herring are anadromous. Management actions also differed between the two fisheries. The sea scallop closed area fisheries were limited in time and space with hard quotas for each species. The herring and mackerel fishery was limited in large scale areas by hard quotas, without quotas for river herring. Fishermen were motivated to avoid river herring by strong conservation pressure, river herring are listed as a species of concern and under review for endangered species by the U.S. agencies, which designation would effectively close the Atlantic herring fishery. In the scallop fisheries, we mapped and updated densities for target and bycatch stocks through real-time monitoring and communicated hotspots with captains at sea. In the herring and mackerel fishery we identified densities through port sampling and communicated hotspots to captains at sea. We also reported environmental factors of river herring density such as depth to captains. Percent participation exceeded 75% in each fisheries with evidence of avoidance behavior in both fisheries. Yellowtail bycatch was sharply reduced in the scallop closed area fisheries with some reduction of river herring bycatch in the herring and mackerel fisheries. Close

• 36. [Article] Species Extinction and Cultural Arrogance: The Perils of Ignoring History

  For more than a century, politicians, newspaper editors, tate and federal fisheries managers, and their various constituencies with an interest in salmon have complained,lobbied and petitioned, sought ...

  Citation × Citation Citation Abstract Copy Title: Species Extinction and Cultural Arrogance: The Perils of Ignoring History Author: Robbins, William G. Year: 2001 For more than a century, politicians, newspaper editors, tate and federal fisheries managers, and their various constituencies with an interest in salmon have complained,lobbied and petitioned, sought legislative relief, and studied to near extinction the declining runs of the Pacific Northwest’s anadromous fish runs. All to no avail, of course. Management efforts have failed; the attempt to the establish reliable, predictive models has failed; massive federal expenditures to rescue crashing stocks have failed; and endeavors to achieve consensus about future policymaking have failed. And today one can add to the now familiar story of extinct and endangered salmon, similar tales about ocean groundfish, newly impoverished fishing communities, reductions in fleet size, and more federal bailouts. What is fascinating in the unveiling of this most recent story is the sense of history repeating itself as commercial fishers shift from one species to the next. Title: Species Extinction and Cultural Arrogance: The Perils of Ignoring History Author: Robbins, William G. Year: 2001 For more than a century, politicians, newspaper editors, tate and federal fisheries managers, and their various constituencies with an interest in salmon have complained,lobbied and petitioned, sought legislative relief, and studied to near extinction the declining runs of the Pacific Northwest’s anadromous fish runs. All to no avail, of course. Management efforts have failed; the attempt to the establish reliable, predictive models has failed; massive federal expenditures to rescue crashing stocks have failed; and endeavors to achieve consensus about future policymaking have failed. And today one can add to the now familiar story of extinct and endangered salmon, similar tales about ocean groundfish, newly impoverished fishing communities, reductions in fleet size, and more federal bailouts. What is fascinating in the unveiling of this most recent story is the sense of history repeating itself as commercial fishers shift from one species to the next. Close

• 37. [Article] Biological and Conference Opinion : Bureau of Land Management Activities Affecting Oregon Coast Coho Salmon and Oregon Coast Steelhead

  The Coos Bay, Medford, and Roseburg Bureau of Land Management (BLM) District Offices submitted May 25 and June 20, 2000, letters requesting formal consultation for a number of proposed activities that ...

  Citation × Citation Citation Abstract Copy Title: Biological and Conference Opinion : Bureau of Land Management Activities Affecting Oregon Coast Coho Salmon and Oregon Coast Steelhead The Coos Bay, Medford, and Roseburg Bureau of Land Management (BLM) District Offices submitted May 25 and June 20, 2000, letters requesting formal consultation for a number of proposed activities that were determined likely to adversely affect Oregon Coast (OC) coho salmon (Oncorhynchus kisutch) and OC steelhead (O. mykiss). Biological assessments (BA) addressing potential affects to these species accompanied the May 25 and June 20, 2000, letters. The BAs describe the environmental baseline and effects of actions in the Umpqua River basin which may have short-term adverse effects on OC coho salmon and OC steelhead and their habitat, but which are expected to provide long-term beneficial effects. Title: Biological and Conference Opinion : Bureau of Land Management Activities Affecting Oregon Coast Coho Salmon and Oregon Coast Steelhead The Coos Bay, Medford, and Roseburg Bureau of Land Management (BLM) District Offices submitted May 25 and June 20, 2000, letters requesting formal consultation for a number of proposed activities that were determined likely to adversely affect Oregon Coast (OC) coho salmon (Oncorhynchus kisutch) and OC steelhead (O. mykiss). Biological assessments (BA) addressing potential affects to these species accompanied the May 25 and June 20, 2000, letters. The BAs describe the environmental baseline and effects of actions in the Umpqua River basin which may have short-term adverse effects on OC coho salmon and OC steelhead and their habitat, but which are expected to provide long-term beneficial effects. Close

• 38. [Article] Genetic diversity in coastal cutthroat trout and Dolly Varden in Prince William Sound, Alaska

  Genetic diversity of two salmonid species, Dolly Varden (Salvelinus malma) and coastal cutthroat trout (Oncorhynchus clarki clarki) in Prince William Sound, Alaska were examined at multiple spatial scales ...

  Citation × Citation Citation Abstract Copy Title: Genetic diversity in coastal cutthroat trout and Dolly Varden in Prince William Sound, Alaska Author: Griswold, Kitty E. Genetic diversity of two salmonid species, Dolly Varden (Salvelinus malma) and coastal cutthroat trout (Oncorhynchus clarki clarki) in Prince William Sound, Alaska were examined at multiple spatial scales with three molecular markers. Pleistocene glaciers covered what is now Prince William Sound 8,000- 12,000 years ago and both species colonized the region subsequently. Because these species have different migratory behavior and historic ranges I expected to see different patterns of genetic diversity within Prince William Sound. Haplotype frequency and nucleotide diversity in sixteen mitochondrial DNA (mtDNA) restriction fragment enzymes combinations in Prince William Sound were higher in Dolly Varden than in coastal cutthroat trout, which were close to fixation. Low estimates of these measures in coastal cutthroat trout may reflect a founder effect resulting from colonization of this region from a single glacial refuge (Cascadia) and low dispersal capacity. To examine if genetic diversity in coastal cutthroat trout was higher in areas of glacial refuge, three additional locations from throughout their distributional range were examined with the same restriction enzymes. Haplotype frequency and nucleotide diversity were lower in areas that were glaciated than estimates of the same measures in a glacial refuge. Again, founder effects during colonization of the three glaciated sites may account for these results. Genetic variation of Dolly Varden and coastal cutthroat trout was also examined within and among populations in Prince William Sound with microsatellites and allozymes. The pattern of genetic variation in coastal cutthroat trout was complex. There was lower genetic diversity within populations that were recently colonized following recent de-glaciation (150-350 ybp). Genetic diversity among some coastal cutthroat trout populations was high, which possibly reflects restricted migration. In other trout populations there was low diversity among populations, possible reflecting historic gene flow. In Dolly Varden, genetic variation among anadromous populations in Prince William Sound was low. There were large differences among resident and anadromous populations. These data provide information for the management for both species, which may reduce the risk of the loss of genetic diversity within local populations. Title: Genetic diversity in coastal cutthroat trout and Dolly Varden in Prince William Sound, Alaska Author: Griswold, Kitty E. Genetic diversity of two salmonid species, Dolly Varden (Salvelinus malma) and coastal cutthroat trout (Oncorhynchus clarki clarki) in Prince William Sound, Alaska were examined at multiple spatial scales with three molecular markers. Pleistocene glaciers covered what is now Prince William Sound 8,000- 12,000 years ago and both species colonized the region subsequently. Because these species have different migratory behavior and historic ranges I expected to see different patterns of genetic diversity within Prince William Sound. Haplotype frequency and nucleotide diversity in sixteen mitochondrial DNA (mtDNA) restriction fragment enzymes combinations in Prince William Sound were higher in Dolly Varden than in coastal cutthroat trout, which were close to fixation. Low estimates of these measures in coastal cutthroat trout may reflect a founder effect resulting from colonization of this region from a single glacial refuge (Cascadia) and low dispersal capacity. To examine if genetic diversity in coastal cutthroat trout was higher in areas of glacial refuge, three additional locations from throughout their distributional range were examined with the same restriction enzymes. Haplotype frequency and nucleotide diversity were lower in areas that were glaciated than estimates of the same measures in a glacial refuge. Again, founder effects during colonization of the three glaciated sites may account for these results. Genetic variation of Dolly Varden and coastal cutthroat trout was also examined within and among populations in Prince William Sound with microsatellites and allozymes. The pattern of genetic variation in coastal cutthroat trout was complex. There was lower genetic diversity within populations that were recently colonized following recent de-glaciation (150-350 ybp). Genetic diversity among some coastal cutthroat trout populations was high, which possibly reflects restricted migration. In other trout populations there was low diversity among populations, possible reflecting historic gene flow. In Dolly Varden, genetic variation among anadromous populations in Prince William Sound was low. There were large differences among resident and anadromous populations. These data provide information for the management for both species, which may reduce the risk of the loss of genetic diversity within local populations. Close

• 39. [Article] Quantifying Salmon-Derived Nutrient Loads from the Mortality of Hatchery-Origin Juvenile Chinook Salmon in the Snake River Basin

  This is the publisher’s final pdf. The article is copyrighted by the American Fisheries Society and published by Taylor & Francis. It can be found at: http://www.tandfonline.com/toc/utaf20/current. To ...

  Citation × Citation Citation Abstract Copy Title: Quantifying Salmon-Derived Nutrient Loads from the Mortality of Hatchery-Origin Juvenile Chinook Salmon in the Snake River Basin Author: Warren, Dana R., McClure, Michelle M. This is the publisher’s final pdf. The article is copyrighted by the American Fisheries Society and published by Taylor & Francis. It can be found at: http://www.tandfonline.com/toc/utaf20/current. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. Title: Quantifying Salmon-Derived Nutrient Loads from the Mortality of Hatchery-Origin Juvenile Chinook Salmon in the Snake River Basin Author: Warren, Dana R., McClure, Michelle M. This is the publisher’s final pdf. The article is copyrighted by the American Fisheries Society and published by Taylor & Francis. It can be found at: http://www.tandfonline.com/toc/utaf20/current. To the best of our knowledge, one or more authors of this paper were federal employees when contributing to this work. Close

• 40. [Article] Susceptibility of select salmonids to Myxobolus cerebralis and effects of exposure on anadromous salmonids in Oregon

  Myxobolus cerebralis, a myxozoan parasite of salmonids, is the causative agent of whirling disease. The parasite is considered widespread throughout northeastern Oregon in the Grande Ronde and Imnaha River ...

  Citation × Citation Citation Abstract Copy Title: Susceptibility of select salmonids to Myxobolus cerebralis and effects of exposure on anadromous salmonids in Oregon Author: Sollid, Sarah A. Myxobolus cerebralis, a myxozoan parasite of salmonids, is the causative agent of whirling disease. The parasite is considered widespread throughout northeastern Oregon in the Grande Ronde and Imnaha River basins where threatened and endangered salmonid populations exist. The work presented in this thesis comprises several studies that assess the effects of M. cerebralis on resident and anadromous salmonids in Oregon. Laboratory challenges to determine the susceptibility of indigenous Deschutes River, Oregon, salmonids to M. cerebralis were conducted as part of a study to assess the risk of reintroducing anadromous salmon above a migration barrier on that river. This study was the first to assess the susceptibility of kokanee salmon Oncorhynchus nerka to M. cerebralis, and results contribute to the understanding of salmonid susceptibility as it relates to species and age. Further, this study demonstrates that the indigenous salmonids present in the river are susceptible to infection, but resident rainbow trout Oncorhynchus mykiss would be most at risk should introduction of M. cerebralis occur in this system. The susceptibility of chinook salmon Oncorhynchus tshawytscha to M. cerebralis was assessed following laboratory challenges at different ages to different parasite levels. Results from this study indicate that chinook salmon are more resistant to M. cerebralis infection than susceptible rainbow trout; resistance to disease developed in chinook salmon exposed after 3 weeks of age. Sustained exposures to a low parasite dose were performed to model the conditions chinook salmon would encounter in a natural exposure. Following continuous exposure to low parasite densities, chinook salmon were very resistant to infection and disease. Managed populations of chinook salmon and steelhead Oncorhynchus mykiss juveniles are held in acclimation facilities on their natal streams before release to commence their seaward migration. Rainbow trout fry were held at each of the acclimation facilities to detect the presence of M. cerebralis and to assess the prevalence of exposure during the acclimation period. Results from these exposures indicate that the Wallowa acclimation facility is a high-risk site for parasite exposure of steelhead juveniles. Further, detection was confirmed in several rivers and streams where the presence of M. cerebralis has been suggested. Finally, a preliminary study on the effect of M. cerebralis on survival of steelhead juveniles upon transfer to saltwater was conducted. Results from this study indicate that M. cerebralis exposure and infection may increase mortality among juvenile steelhead during saltwater adaptation. Title: Susceptibility of select salmonids to Myxobolus cerebralis and effects of exposure on anadromous salmonids in Oregon Author: Sollid, Sarah A. Myxobolus cerebralis, a myxozoan parasite of salmonids, is the causative agent of whirling disease. The parasite is considered widespread throughout northeastern Oregon in the Grande Ronde and Imnaha River basins where threatened and endangered salmonid populations exist. The work presented in this thesis comprises several studies that assess the effects of M. cerebralis on resident and anadromous salmonids in Oregon. Laboratory challenges to determine the susceptibility of indigenous Deschutes River, Oregon, salmonids to M. cerebralis were conducted as part of a study to assess the risk of reintroducing anadromous salmon above a migration barrier on that river. This study was the first to assess the susceptibility of kokanee salmon Oncorhynchus nerka to M. cerebralis, and results contribute to the understanding of salmonid susceptibility as it relates to species and age. Further, this study demonstrates that the indigenous salmonids present in the river are susceptible to infection, but resident rainbow trout Oncorhynchus mykiss would be most at risk should introduction of M. cerebralis occur in this system. The susceptibility of chinook salmon Oncorhynchus tshawytscha to M. cerebralis was assessed following laboratory challenges at different ages to different parasite levels. Results from this study indicate that chinook salmon are more resistant to M. cerebralis infection than susceptible rainbow trout; resistance to disease developed in chinook salmon exposed after 3 weeks of age. Sustained exposures to a low parasite dose were performed to model the conditions chinook salmon would encounter in a natural exposure. Following continuous exposure to low parasite densities, chinook salmon were very resistant to infection and disease. Managed populations of chinook salmon and steelhead Oncorhynchus mykiss juveniles are held in acclimation facilities on their natal streams before release to commence their seaward migration. Rainbow trout fry were held at each of the acclimation facilities to detect the presence of M. cerebralis and to assess the prevalence of exposure during the acclimation period. Results from these exposures indicate that the Wallowa acclimation facility is a high-risk site for parasite exposure of steelhead juveniles. Further, detection was confirmed in several rivers and streams where the presence of M. cerebralis has been suggested. Finally, a preliminary study on the effect of M. cerebralis on survival of steelhead juveniles upon transfer to saltwater was conducted. Results from this study indicate that M. cerebralis exposure and infection may increase mortality among juvenile steelhead during saltwater adaptation. Close

• 41. [Article] Investigating Demographic and Evolutionary Factors Important for Fish Reintroduction

  Reintroduction programs are used to re-establish species back into their historical habitat. Most reintroduction programs have failed and few papers have evaluated factors that may be important to Pacific ...

  Citation × Citation Citation Abstract Copy Title: Investigating Demographic and Evolutionary Factors Important for Fish Reintroduction Author: Sard, Nicholas M. Reintroduction programs are used to re-establish species back into their historical habitat. Most reintroduction programs have failed and few papers have evaluated factors that may be important to Pacific salmon. The 158 meter tall Cougar Dam has blocked Chinook salmon (Oncorhynchus tshawytscha) from accessing 40 kilometers of historical spawning and rearing habitat for over 50 years. Here, I evaluated a Chinook salmon trap and transport reintroduction program above Cougar Dam on the South Fork McKenzie River, Oregon from 2007 to 2015. First, I evaluated if release location and date correlated with two measures of fitness based on adult assignments to age-0 juveniles collected above the dam and adult offspring returning to the South Fork McKenzie River, respectively. I found that release location and date had little to no effect on either measure of fitness. I also evaluated if there were fitness differences between hatchery and natural origin (HOR and NOR, respectively) adults. I found consistent fitness differences between males (RRS=0.48, p<0.001), but not females (RRS=0.72, p=0.81). In addition, I found that origin (p=0.352) no longer explained variation in fitness after accounting for variation in fork-length, which suggests that HOR fish may be less fit, in part, because they are 2-4 cm (95% CI) smaller, or perhaps younger. I also evaluated a measure of population productivity known as cohort replacement rate (CRR) - defined as the number of future spawners produced by a spawner. Based on genetic parentage assignments to NOR adults returning to the trap and transport facility, hereafter Cougar Trap, I found that adults reintroduced in 2007 and 2008 did not meet demographic replacement (CRR: 0.40 and 0.31, respectively). I also found a seasonal decline in the proportion of NOR adults produced above the dam that returned to the Cougar Trap. I also developed grandparentage assignment methods to determine how many precocial males and adfluvial Chinook salmon, two non-anadromous life history tactics, contributed to population productivity. I found 31 unsampled precocial males, as well as 48 age-4 and -5 probable adfluvial male and female adults contributed to the reintroduced population. My discovery of adfluvial Chinook salmon contributing to population productivity is significant because little is known about this life history tactic, and they provide resiliency to a reintroduced population. I show that adfluvial adults can be produced by anadromous mate pairs. Adfluvial adults increased CRR estimates; however, neither the 2007 or 2008 cohorts met replacement after incorporating this non-anadromous life history tactic (CRR: 0.46 and 0.35, respectively). Finally, I assessed if genetic variation in founding cohorts was maintained in their returning adult offspring returning to the South Fork McKenzie River, hereafter F₁ offspring. On average, 6 alleles were lost per locus between founding cohorts and their F₁ offspring. N[subscript e] estimates were high using either demographic or genetic methods (range: 344 to 893). My dissertation research provides valuable information on factors that may be important to population productivity, as well as the maintenance of genetic variation in nascent populations established through reintroduction. Title: Investigating Demographic and Evolutionary Factors Important for Fish Reintroduction Author: Sard, Nicholas M. Reintroduction programs are used to re-establish species back into their historical habitat. Most reintroduction programs have failed and few papers have evaluated factors that may be important to Pacific salmon. The 158 meter tall Cougar Dam has blocked Chinook salmon (Oncorhynchus tshawytscha) from accessing 40 kilometers of historical spawning and rearing habitat for over 50 years. Here, I evaluated a Chinook salmon trap and transport reintroduction program above Cougar Dam on the South Fork McKenzie River, Oregon from 2007 to 2015. First, I evaluated if release location and date correlated with two measures of fitness based on adult assignments to age-0 juveniles collected above the dam and adult offspring returning to the South Fork McKenzie River, respectively. I found that release location and date had little to no effect on either measure of fitness. I also evaluated if there were fitness differences between hatchery and natural origin (HOR and NOR, respectively) adults. I found consistent fitness differences between males (RRS=0.48, p<0.001), but not females (RRS=0.72, p=0.81). In addition, I found that origin (p=0.352) no longer explained variation in fitness after accounting for variation in fork-length, which suggests that HOR fish may be less fit, in part, because they are 2-4 cm (95% CI) smaller, or perhaps younger. I also evaluated a measure of population productivity known as cohort replacement rate (CRR) - defined as the number of future spawners produced by a spawner. Based on genetic parentage assignments to NOR adults returning to the trap and transport facility, hereafter Cougar Trap, I found that adults reintroduced in 2007 and 2008 did not meet demographic replacement (CRR: 0.40 and 0.31, respectively). I also found a seasonal decline in the proportion of NOR adults produced above the dam that returned to the Cougar Trap. I also developed grandparentage assignment methods to determine how many precocial males and adfluvial Chinook salmon, two non-anadromous life history tactics, contributed to population productivity. I found 31 unsampled precocial males, as well as 48 age-4 and -5 probable adfluvial male and female adults contributed to the reintroduced population. My discovery of adfluvial Chinook salmon contributing to population productivity is significant because little is known about this life history tactic, and they provide resiliency to a reintroduced population. I show that adfluvial adults can be produced by anadromous mate pairs. Adfluvial adults increased CRR estimates; however, neither the 2007 or 2008 cohorts met replacement after incorporating this non-anadromous life history tactic (CRR: 0.46 and 0.35, respectively). Finally, I assessed if genetic variation in founding cohorts was maintained in their returning adult offspring returning to the South Fork McKenzie River, hereafter F₁ offspring. On average, 6 alleles were lost per locus between founding cohorts and their F₁ offspring. N[subscript e] estimates were high using either demographic or genetic methods (range: 344 to 893). My dissertation research provides valuable information on factors that may be important to population productivity, as well as the maintenance of genetic variation in nascent populations established through reintroduction. Close

• 42. [Article] A bioeconomic analysis of altering instream flows anadromous fish production and competing demands for water in the John Day River basin, Oregon

  The growing demand for water in the arid regions of the West increases the need for optimal allocation of water among competing uses. An efficient allocation of water between instream and out-of-stream uses ...

  Citation × Citation Citation Abstract Copy Title: A bioeconomic analysis of altering instream flows anadromous fish production and competing demands for water in the John Day River basin, Oregon Author: Johnson, Neal S. The growing demand for water in the arid regions of the West increases the need for optimal allocation of water among competing uses. An efficient allocation of water between instream and out-of-stream uses has been impeded by institutional constraints and the scarcity of information regarding instream flow benefits. The objectives of this thesis were to provide preliminary economic data on the value of instream water in "producing" recreational fishing and to examine the effect of forestry, agriculture, and livestock practices on temporal streamflow patterns and anadromous fish production. The steelhead trout (Salmo gairdneri) sport fishery within the John Day River basin in north-central Oregon provided the setting for this research. The interdisciplinary methodology employed in estimating the marginal value of water with respect to steelhead production consisted of two tasks. The first task involved valuing a marginal change in the quality of the steelhead recreational fishery. The contingent valuation method (CVM) was selected for this purpose. Both open- and closed-ended willingness-to-pay (WTP) questions were included in a questionnaire administered to John Day River steelhead anglers during the 1986/87 steelhead fishing season. Survey data were analyzed to arrive at individual and aggregate bid functions relating WTP to expected angling success rates. Results indicate that, under current conditions, the average angler is willing to pay approximately $7.20 to catch an additional steelhead. The second task of the instream water valuation methodology was directed at deriving a streamflow/steelhead production relationship. By including variables influencing steelhead production in a Ricker stock-recruitment model, it was possible to develop a model which could be estimated using linear regression techniques. Some difficulty arose, however, with interpretation of the model due to the unavailability of cohort escapement data and the subsequent use of standing crop data. While possibly masking the true magnitude of streamflow's effect on fish production, this drawback was not deemed limiting within the general context of the interdisciplinary methodology. Results of the biological model conformed to a priori expectations. Increases in summer and winter streamflows led to increased steelhead survival, whereas higher spring flows increased mortality levels. Other results indicate that the John Day Dam was responsible for a 31.5 percent decline in the population index for the 1969-1983 period. Combining the economic and biological results into one equation yielded an estimate of the marginal value of summer instream water in "producing" recreational steelhead angling. Similar equations were developed for winter and spring flows. The marginal value of water in producing recreational steelhead fishing within the John Day basin was estimated at $0.56 per acre-foot for summer flows, $0.046 for winter flows, and -$0.075 for spring flows. By including out-of-basin benefits, these values increased to $2.26, $0.19, and -$0.30, respectively. In comparison, water's value in irrigation within the John Day basin has been estimated at between $10 to $24 per acre-foot. However, nonuse values of steelhead, as well as the increased production of other fish species (such as spring chinook salmon) were not included in the instream water values. In addition, no attempt was made at valuing instream water's contribution to boating, camping, or other benefit-producing activities. A secondary objective of this thesis was to briefly examine the possible benefits accruing to other instream and out-of-stream users due to an alteration in streamflow patterns. In addition, the impact of activities by other resource users -- namely forestry, agriculture, and livestock production --on anadromous fish production was reviewed. Improper management practices by these activities can negatively impact the aquatic and riparian ecosystems. While no firm conclusions were drawn, it appears the quality of these ecosystems, as opposed to the amount of streamflow, has the largest marginal impact on anadromous fish populations. Title: A bioeconomic analysis of altering instream flows anadromous fish production and competing demands for water in the John Day River basin, Oregon Author: Johnson, Neal S. The growing demand for water in the arid regions of the West increases the need for optimal allocation of water among competing uses. An efficient allocation of water between instream and out-of-stream uses has been impeded by institutional constraints and the scarcity of information regarding instream flow benefits. The objectives of this thesis were to provide preliminary economic data on the value of instream water in "producing" recreational fishing and to examine the effect of forestry, agriculture, and livestock practices on temporal streamflow patterns and anadromous fish production. The steelhead trout (Salmo gairdneri) sport fishery within the John Day River basin in north-central Oregon provided the setting for this research. The interdisciplinary methodology employed in estimating the marginal value of water with respect to steelhead production consisted of two tasks. The first task involved valuing a marginal change in the quality of the steelhead recreational fishery. The contingent valuation method (CVM) was selected for this purpose. Both open- and closed-ended willingness-to-pay (WTP) questions were included in a questionnaire administered to John Day River steelhead anglers during the 1986/87 steelhead fishing season. Survey data were analyzed to arrive at individual and aggregate bid functions relating WTP to expected angling success rates. Results indicate that, under current conditions, the average angler is willing to pay approximately $7.20 to catch an additional steelhead. The second task of the instream water valuation methodology was directed at deriving a streamflow/steelhead production relationship. By including variables influencing steelhead production in a Ricker stock-recruitment model, it was possible to develop a model which could be estimated using linear regression techniques. Some difficulty arose, however, with interpretation of the model due to the unavailability of cohort escapement data and the subsequent use of standing crop data. While possibly masking the true magnitude of streamflow's effect on fish production, this drawback was not deemed limiting within the general context of the interdisciplinary methodology. Results of the biological model conformed to a priori expectations. Increases in summer and winter streamflows led to increased steelhead survival, whereas higher spring flows increased mortality levels. Other results indicate that the John Day Dam was responsible for a 31.5 percent decline in the population index for the 1969-1983 period. Combining the economic and biological results into one equation yielded an estimate of the marginal value of summer instream water in "producing" recreational steelhead angling. Similar equations were developed for winter and spring flows. The marginal value of water in producing recreational steelhead fishing within the John Day basin was estimated at $0.56 per acre-foot for summer flows, $0.046 for winter flows, and -$0.075 for spring flows. By including out-of-basin benefits, these values increased to $2.26, $0.19, and -$0.30, respectively. In comparison, water's value in irrigation within the John Day basin has been estimated at between $10 to $24 per acre-foot. However, nonuse values of steelhead, as well as the increased production of other fish species (such as spring chinook salmon) were not included in the instream water values. In addition, no attempt was made at valuing instream water's contribution to boating, camping, or other benefit-producing activities. A secondary objective of this thesis was to briefly examine the possible benefits accruing to other instream and out-of-stream users due to an alteration in streamflow patterns. In addition, the impact of activities by other resource users -- namely forestry, agriculture, and livestock production --on anadromous fish production was reviewed. Improper management practices by these activities can negatively impact the aquatic and riparian ecosystems. While no firm conclusions were drawn, it appears the quality of these ecosystems, as opposed to the amount of streamflow, has the largest marginal impact on anadromous fish populations. Close

• 43. [Article] Over 50 Years of Government Fisheries Service—Then and Now

  As background, at the national level, for many years the U.S. Bureau of Commercial Fisheries concentrated on exploratory fishing, gear development, food science, catch statistics, scientific research, ...

  Citation × Citation Citation Abstract Copy Title: Over 50 Years of Government Fisheries Service—Then and Now Author: Schoning, Robert W. Year: 2001 As background, at the national level, for many years the U.S. Bureau of Commercial Fisheries concentrated on exploratory fishing, gear development, food science, catch statistics, scientific research, and marketing, and serviced the commercial industry in a variety of ways. In the early 1970's the name was changed to National Marine Fisheries Service and with the 200 mile law in 1976, its role changed very significantly to include major responsibilities in management and enforcement. It issues more than 400 rules a year. In 1978 its annual budget was about $150 million. In 2000 it is $383 million and the president has requested $657 million for 2001. I will discuss five examples of marine or anadromous fish life, the role government has played, the results to date, and some views about the future. The examples are Columbia River salmon, Pacific halibut, marine mammals, endangered species, and marine fish. Title: Over 50 Years of Government Fisheries Service—Then and Now Author: Schoning, Robert W. Year: 2001 As background, at the national level, for many years the U.S. Bureau of Commercial Fisheries concentrated on exploratory fishing, gear development, food science, catch statistics, scientific research, and marketing, and serviced the commercial industry in a variety of ways. In the early 1970's the name was changed to National Marine Fisheries Service and with the 200 mile law in 1976, its role changed very significantly to include major responsibilities in management and enforcement. It issues more than 400 rules a year. In 1978 its annual budget was about $150 million. In 2000 it is $383 million and the president has requested $657 million for 2001. I will discuss five examples of marine or anadromous fish life, the role government has played, the results to date, and some views about the future. The examples are Columbia River salmon, Pacific halibut, marine mammals, endangered species, and marine fish. Close

• 44. [Article] Biological opinion: Canby Ferry shoreside improvements project, Mountain Road, Clackamas County

  Citation × Citation Citation Abstract Copy Title: Biological opinion: Canby Ferry shoreside improvements project, Mountain Road, Clackamas County Author: U.S. Federal Highway Administration, National Marine Fisheries Service, Northwest Region Title: Biological opinion: Canby Ferry shoreside improvements project, Mountain Road, Clackamas County Author: U.S. Federal Highway Administration, National Marine Fisheries Service, Northwest Region Close

• 45. [Article] Estuary management in the Pacific Northwest

  Citation × Citation Citation Abstract Copy Title: Estuary management in the Pacific Northwest Title: Estuary management in the Pacific Northwest Close

• 46. [Article] River-dependent bird species as potential indicators of ecosystem response to removal of dams on the Elwha River, Washington

  If two dams on the Elwha River are removed, the ecosystem will be open to the downstream flow of sediments and the upstream flow of marine nutrients in the form of anadromous fish. Nutrient enrichment ...

  Citation × Citation Citation Abstract Copy Title: River-dependent bird species as potential indicators of ecosystem response to removal of dams on the Elwha River, Washington Author: Blackie, Barbara If two dams on the Elwha River are removed, the ecosystem will be open to the downstream flow of sediments and the upstream flow of marine nutrients in the form of anadromous fish. Nutrient enrichment may influence trophic dynamics of the entire ecosystem, extending beyond the aquatic boundary. I assessed the current relative densities of five river dependent bird species on the Elwha and three other rivers in Olympic National Park in Washington State to describe pretreatment reference conditions as a basis for assessment of post-treatment ecosystem responses. I also compared the amount of time that non-breeding and failed-breeding (NBIFB) female Harlequin Ducks (Histrionicus histrionicus) spent foraging on different rivers and on adjacent coastal habitat to determine whether time spent foraging could be used as an indicator of habitat preference. Surveys of key river-dependent bird species were conducted on two rivers in 1996 and on four rivers in 1997. The benthivorous foraging guild was represented by the American Dipper (Cinclus mexicanus), Harlequin Duck and Spotted Sandpiper (Actitis macularia). Harlequin Ducks had higher relative densities on the Elwha than the Hoh and Soleduck Rivers, but densities were similar to those found on the Duckabush River. There were greater numbers of Harlequin Ducks per linear kilometer above the two dams than between or below them. Relative densities of American Dippers on the Elwha were lower than on the Duckabush but not statistically different from those on the Hoh and Soleduck Rivers. Spotted Sandpipers had similar densities on all rivers except the Soleduck River where their numbers were lower. The piscivorous foraging guild was represented by the Common Merganser (Mergus merganser) and Belted Kingfisher (Ceryle alcyon). Common Mergansers were relatively more abundant on the Elwha River than on the other rivers due to their high numbers below the lowest dam where wild and hatchery anadromous fish are present. Belted Kingfisher relative densities showed no significant differences between rivers as they were found in low numbers on all rivers. Using relative abundances of some river dependent bird species as indicators of ecosystem recovery after dam removal may be useful tool surveys are continued throughout the process. In spite of lack of anadromy, the Elwha River ecosystem supports a significant population of Harlequin Ducks when compared to other Olympic rivers. American Dippers are also well represented on the Elwha River, specifically above the dams. Because of their relative abundance, these benthivore populations may have a measurable, upriver response to any nutrient enrichment as a result of dam removal and both species should be monitored. The Duckabush River, with a similar Harlequin Duck relative abundance in the lower reaches, should be monitored concurrently to account for region-wide population shifts of that species over time. The Common Merganser may be the best indicator of below dam effects on the Elwha River and numbers could be compared with the lower Soleduck River, however alternative survey methods, specifically drift boat surveys, should be considered to improve count accuracy. During June and July of 1997, time-activity budget data were gathered on NB/FB Harlequin Ducks on the Elwha, Duckabush and Dosewallips Rivers and at their mouths and at the mouth of Salt Creek. Time spent foraging by NBIFB Harlequin Ducks was similar among the three rivers examined. NB/FB Harlequin Ducks on the spent 33.1%, 33.2%, 36.4% of their time feeding, on the Elwha, Duckabush Dosewallips Rivers respectively. The Elwha River system was the only system where time activity budgets could be compared between river and adjacent coastal habitats. Time spent foraging at the mouth of the Elwha was significantly higher (52.5%) than on the Elwha River proper. Food availability on river habitat may be a factor in NB/FB females' decision to remain on river habitat during the breeding season or migrate to the ocean. Time-activity budgets provide information on Harlequin Duck life history but a better understanding of their foraging behavior and the role of food availability in habitat selection is needed before time-activity budgets alone can be used as a tool for assessing ecosystem response to dam removal. Time-activity budget data collected concurrently with food availability data from field studies, along with energetics and food preference data from field or laboratory studies may establish a clearer link between time spent feeding and habitat quality and preferences. Title: River-dependent bird species as potential indicators of ecosystem response to removal of dams on the Elwha River, Washington Author: Blackie, Barbara If two dams on the Elwha River are removed, the ecosystem will be open to the downstream flow of sediments and the upstream flow of marine nutrients in the form of anadromous fish. Nutrient enrichment may influence trophic dynamics of the entire ecosystem, extending beyond the aquatic boundary. I assessed the current relative densities of five river dependent bird species on the Elwha and three other rivers in Olympic National Park in Washington State to describe pretreatment reference conditions as a basis for assessment of post-treatment ecosystem responses. I also compared the amount of time that non-breeding and failed-breeding (NBIFB) female Harlequin Ducks (Histrionicus histrionicus) spent foraging on different rivers and on adjacent coastal habitat to determine whether time spent foraging could be used as an indicator of habitat preference. Surveys of key river-dependent bird species were conducted on two rivers in 1996 and on four rivers in 1997. The benthivorous foraging guild was represented by the American Dipper (Cinclus mexicanus), Harlequin Duck and Spotted Sandpiper (Actitis macularia). Harlequin Ducks had higher relative densities on the Elwha than the Hoh and Soleduck Rivers, but densities were similar to those found on the Duckabush River. There were greater numbers of Harlequin Ducks per linear kilometer above the two dams than between or below them. Relative densities of American Dippers on the Elwha were lower than on the Duckabush but not statistically different from those on the Hoh and Soleduck Rivers. Spotted Sandpipers had similar densities on all rivers except the Soleduck River where their numbers were lower. The piscivorous foraging guild was represented by the Common Merganser (Mergus merganser) and Belted Kingfisher (Ceryle alcyon). Common Mergansers were relatively more abundant on the Elwha River than on the other rivers due to their high numbers below the lowest dam where wild and hatchery anadromous fish are present. Belted Kingfisher relative densities showed no significant differences between rivers as they were found in low numbers on all rivers. Using relative abundances of some river dependent bird species as indicators of ecosystem recovery after dam removal may be useful tool surveys are continued throughout the process. In spite of lack of anadromy, the Elwha River ecosystem supports a significant population of Harlequin Ducks when compared to other Olympic rivers. American Dippers are also well represented on the Elwha River, specifically above the dams. Because of their relative abundance, these benthivore populations may have a measurable, upriver response to any nutrient enrichment as a result of dam removal and both species should be monitored. The Duckabush River, with a similar Harlequin Duck relative abundance in the lower reaches, should be monitored concurrently to account for region-wide population shifts of that species over time. The Common Merganser may be the best indicator of below dam effects on the Elwha River and numbers could be compared with the lower Soleduck River, however alternative survey methods, specifically drift boat surveys, should be considered to improve count accuracy. During June and July of 1997, time-activity budget data were gathered on NB/FB Harlequin Ducks on the Elwha, Duckabush and Dosewallips Rivers and at their mouths and at the mouth of Salt Creek. Time spent foraging by NBIFB Harlequin Ducks was similar among the three rivers examined. NB/FB Harlequin Ducks on the spent 33.1%, 33.2%, 36.4% of their time feeding, on the Elwha, Duckabush Dosewallips Rivers respectively. The Elwha River system was the only system where time activity budgets could be compared between river and adjacent coastal habitats. Time spent foraging at the mouth of the Elwha was significantly higher (52.5%) than on the Elwha River proper. Food availability on river habitat may be a factor in NB/FB females' decision to remain on river habitat during the breeding season or migrate to the ocean. Time-activity budgets provide information on Harlequin Duck life history but a better understanding of their foraging behavior and the role of food availability in habitat selection is needed before time-activity budgets alone can be used as a tool for assessing ecosystem response to dam removal. Time-activity budget data collected concurrently with food availability data from field studies, along with energetics and food preference data from field or laboratory studies may establish a clearer link between time spent feeding and habitat quality and preferences. Close

• 47. [Article] Life history and evolutionary adaptation of Pacific salmon and its application in management

  An approach to understanding and managing anadromous salmon, steelhead, and sea-run cutthroat trout (Oncorhynchus spp.) based on life history and evolutionary adaptive capacities of species and stocks ...

  Citation × Citation Citation Abstract Copy Title: Life history and evolutionary adaptation of Pacific salmon and its application in management Author: Wevers, Mary Jo An approach to understanding and managing anadromous salmon, steelhead, and sea-run cutthroat trout (Oncorhynchus spp.) based on life history and evolutionary adaptive capacities of species and stocks is presented. Species, stocks, and local populations are viewed as systems that are continuously adapting to changing environmental conditions. They have the potential capacity to evolve in different ways in different environments through both life history and evolutionary adaptation. Habitat organization forms a template for genus, species, stock, and local population life history organization. Harvesting, habitat alteration resulting from land use practices and other human activities can alter the organization and adaptive capacities of species and stocks, and thus their long term persistence. The adaptive capacity of Oncorhynchus relative to its habitat and management environment is examined at the species, stock, and local population levels. Life history characteristics of representative stocks and local populations are analyzed using Detrended Correspondence Analysis (DECORANA). Fresh water migration distance and latitude are used to "explain" ordination patterns of Oncorhynchus species in the North Pacific Basin. Fresh water migration difficulty and mean annual runoff are used to interpret life history patterns of Columbia Basin chinook salmon stocks. Upstream migration difficulty and fall water temperatures are used to explain the ordination patterns of local populations of Willamette spring chinook salmon. Fishery management practices are examined in terms of their impacts on the organization and adaptive capacity of species, stocks, and local populations of Oncorhynchus. Management generalizations and guidelines derived from the life history theory are applied to management of Willamette spring chinook salmon. Maintaining habitat changes in the Willamette Basin within the historic range of fluctuations will tend to maximize co-organization of local populations. Management activities should provide relatively constant habitat and fishery conditions for natural selection processes to "organize" life history traits over a period of at least a few generations. Fisheries should selectively harvest local populations that show a high degree of realization of their adaptive capacity. By focusing instead on maximizing the co-organization of stocks, their habitat and fishery environments, and protecting the adaptive capacities of stocks, we will go a long way toward providing long-term sustainability for social communities dependent on fisheries. Title: Life history and evolutionary adaptation of Pacific salmon and its application in management Author: Wevers, Mary Jo An approach to understanding and managing anadromous salmon, steelhead, and sea-run cutthroat trout (Oncorhynchus spp.) based on life history and evolutionary adaptive capacities of species and stocks is presented. Species, stocks, and local populations are viewed as systems that are continuously adapting to changing environmental conditions. They have the potential capacity to evolve in different ways in different environments through both life history and evolutionary adaptation. Habitat organization forms a template for genus, species, stock, and local population life history organization. Harvesting, habitat alteration resulting from land use practices and other human activities can alter the organization and adaptive capacities of species and stocks, and thus their long term persistence. The adaptive capacity of Oncorhynchus relative to its habitat and management environment is examined at the species, stock, and local population levels. Life history characteristics of representative stocks and local populations are analyzed using Detrended Correspondence Analysis (DECORANA). Fresh water migration distance and latitude are used to "explain" ordination patterns of Oncorhynchus species in the North Pacific Basin. Fresh water migration difficulty and mean annual runoff are used to interpret life history patterns of Columbia Basin chinook salmon stocks. Upstream migration difficulty and fall water temperatures are used to explain the ordination patterns of local populations of Willamette spring chinook salmon. Fishery management practices are examined in terms of their impacts on the organization and adaptive capacity of species, stocks, and local populations of Oncorhynchus. Management generalizations and guidelines derived from the life history theory are applied to management of Willamette spring chinook salmon. Maintaining habitat changes in the Willamette Basin within the historic range of fluctuations will tend to maximize co-organization of local populations. Management activities should provide relatively constant habitat and fishery conditions for natural selection processes to "organize" life history traits over a period of at least a few generations. Fisheries should selectively harvest local populations that show a high degree of realization of their adaptive capacity. By focusing instead on maximizing the co-organization of stocks, their habitat and fishery environments, and protecting the adaptive capacities of stocks, we will go a long way toward providing long-term sustainability for social communities dependent on fisheries. Close

• 48. [Article] Relationships among juvenile anadromous salmonids, their freshwater habitat, and landscape characteristics over multiple years and spatial scales in the Elk River, Oregon

  Research on the distribution of juvenile salmonids in streams has been dominated by studies examining small areas over short periods. However, information relevant to freshwater influences on population ...

  Citation × Citation Citation Abstract Copy Title: Relationships among juvenile anadromous salmonids, their freshwater habitat, and landscape characteristics over multiple years and spatial scales in the Elk River, Oregon Author: Burnett, Kelly M. Research on the distribution of juvenile salmonids in streams has been dominated by studies examining small areas over short periods. However, information relevant to freshwater influences on population persistence is likely to derive from longer-term, multi-scale studies. Relationships were examined among juvenile anadromous salmonids, their freshwater habitat, and landscape characteristics throughout the Elk River, Oregon over 7 years at multiple spatial scales. Ocean-type chinook salmon (Oncorhynchus tshawytscha), coho salmon (0. kisutch), coastal cutthroat trout (0. clarki) and winter-run steelhead (0. mykiss) comprised the salmonid assemblage. Habitat selection was quantified at stream system, valley segment, and channel unit scales by selection ratios estimated with bootstrapping methods. Unconstrained valleys in tributaries and pools in the mainstem were typically selected by each species except steelhead, which often avoided these. Valley segment types generally did not differ for characteristics routinely assessed in stream surveys. Thus, fish probably perceived other biotic or abiotic differences among valley segment types. Evidence suggested competition may have influenced selection by coho and chinook salmon. Discriminant analysis indicated that level of use by juvenile chinook salmon appeared related to valley segment type and spatial position. Unconstrained valleys, nearby valley segments, and valley segments with larger, deeper pools, containing more wood were most highly used by chinook salmon. Mean volume and maximum depth of pools were each directly related to catchment area, which explained more variation than landscape characteristics summarized at any of five spatial scales. At each scale except the most spatially extensive, wood density in valley segments was negatively related to the percent area in resistant rock types and positively related to the percent area in mature to old forests. The most variation was explained with these landscape variables summarized at an intermediate spatial scale (i.e., sub-catchment). Although spatial scales appeared similar in processes affecting wood density, finer scales omitted key source areas for wood delivery, and coarser scales included source areas less tightly coupled to wood dynamics in surveyed channels. If only 1 or 2 years of data or one spatial scale had been examined, as commonly occurs, conclusions may have differed substantially from those in this study. Title: Relationships among juvenile anadromous salmonids, their freshwater habitat, and landscape characteristics over multiple years and spatial scales in the Elk River, Oregon Author: Burnett, Kelly M. Research on the distribution of juvenile salmonids in streams has been dominated by studies examining small areas over short periods. However, information relevant to freshwater influences on population persistence is likely to derive from longer-term, multi-scale studies. Relationships were examined among juvenile anadromous salmonids, their freshwater habitat, and landscape characteristics throughout the Elk River, Oregon over 7 years at multiple spatial scales. Ocean-type chinook salmon (Oncorhynchus tshawytscha), coho salmon (0. kisutch), coastal cutthroat trout (0. clarki) and winter-run steelhead (0. mykiss) comprised the salmonid assemblage. Habitat selection was quantified at stream system, valley segment, and channel unit scales by selection ratios estimated with bootstrapping methods. Unconstrained valleys in tributaries and pools in the mainstem were typically selected by each species except steelhead, which often avoided these. Valley segment types generally did not differ for characteristics routinely assessed in stream surveys. Thus, fish probably perceived other biotic or abiotic differences among valley segment types. Evidence suggested competition may have influenced selection by coho and chinook salmon. Discriminant analysis indicated that level of use by juvenile chinook salmon appeared related to valley segment type and spatial position. Unconstrained valleys, nearby valley segments, and valley segments with larger, deeper pools, containing more wood were most highly used by chinook salmon. Mean volume and maximum depth of pools were each directly related to catchment area, which explained more variation than landscape characteristics summarized at any of five spatial scales. At each scale except the most spatially extensive, wood density in valley segments was negatively related to the percent area in resistant rock types and positively related to the percent area in mature to old forests. The most variation was explained with these landscape variables summarized at an intermediate spatial scale (i.e., sub-catchment). Although spatial scales appeared similar in processes affecting wood density, finer scales omitted key source areas for wood delivery, and coarser scales included source areas less tightly coupled to wood dynamics in surveyed channels. If only 1 or 2 years of data or one spatial scale had been examined, as commonly occurs, conclusions may have differed substantially from those in this study. Close

• 49. [Article] Age, length and weight studies of three species of Columbia River salmon (Oncorhynchus keta, O. gorbuscha and O. kisutch)

  Samples, consisting of scales, length and weight measurements, and sex determinations of chum, pink, and silver salmons, were taken from the commercial catch in the Columbia River in 1914. Five hundred ...

  Citation × Citation Citation Abstract Copy Title: Age, length and weight studies of three species of Columbia River salmon (Oncorhynchus keta, O. gorbuscha and O. kisutch) Author: Marr, John C., Oregon. Fish Commission Samples, consisting of scales, length and weight measurements, and sex determinations of chum, pink, and silver salmons, were taken from the commercial catch in the Columbia River in 1914. Five hundred eighteen chum scales were examined. All fish had gone to sea early in their first year; and 70.5 percent had returned in their third year, 28.7 percent in their fourth, and 0.8 percent in their fifth. Growth curves, based on scale and fish measurements, are given for all sex-age groups. The length-weight relationship may be expressed by the equation Weight = 0.0001378 x Length [superscript 2.6004]. Tendencies were noted, during the course of the run, for older fish and for males to be relatively more abundant in the earlier portions of the run. In comparisons with data from other localities, it was demonstrated that, from south to north: (1) there is a decrease in mean length at the same age; (2) older fish are progressively more abundant; and (3) the runs are progressively earlier. Pink salmon are uncommon in the Columbia River, and only six were examined. All had gone to sea early in their first year, and all had returned in their second year. Eight hundred eighty-five silver salmon scales were examined. Of the total, 6.1 percent had gone to sea early in their second year, and returned late in their second year (2 [subscript 2]); 0.3 percent had gone to sea early in their third, and returned late in their third (3 [subscript 3]); 83.9 percent had gone to sea in their second, and returned in their third (3 [subscript 2]); and 9.7 percent had gone to sea in their third year, and returned during their fourth year (4 [subscript 3]). Growth curves are given for all, age groups. The length-weight relationship may be expressed by the equation Weight = 0.0000373 x Length [superscript 2.8945]. As in the case of the chums, changes were noted during the run in the sex ratio and the percentage of age classes. Comparison with data from other localities revealed south-to-north changes in size and in abundance of different age groups, similar to those demonstrated for the chums. It is felt that the differences between races of a single river system will prove to be of the same nature as those here demonstrated between populations of different river systems. Knowledge of the differences between the races of salmons entering the Columbia River may be of extreme importance in the management of the fisheries. Title: Age, length and weight studies of three species of Columbia River salmon (Oncorhynchus keta, O. gorbuscha and O. kisutch) Author: Marr, John C., Oregon. Fish Commission Samples, consisting of scales, length and weight measurements, and sex determinations of chum, pink, and silver salmons, were taken from the commercial catch in the Columbia River in 1914. Five hundred eighteen chum scales were examined. All fish had gone to sea early in their first year; and 70.5 percent had returned in their third year, 28.7 percent in their fourth, and 0.8 percent in their fifth. Growth curves, based on scale and fish measurements, are given for all sex-age groups. The length-weight relationship may be expressed by the equation Weight = 0.0001378 x Length [superscript 2.6004]. Tendencies were noted, during the course of the run, for older fish and for males to be relatively more abundant in the earlier portions of the run. In comparisons with data from other localities, it was demonstrated that, from south to north: (1) there is a decrease in mean length at the same age; (2) older fish are progressively more abundant; and (3) the runs are progressively earlier. Pink salmon are uncommon in the Columbia River, and only six were examined. All had gone to sea early in their first year, and all had returned in their second year. Eight hundred eighty-five silver salmon scales were examined. Of the total, 6.1 percent had gone to sea early in their second year, and returned late in their second year (2 [subscript 2]); 0.3 percent had gone to sea early in their third, and returned late in their third (3 [subscript 3]); 83.9 percent had gone to sea in their second, and returned in their third (3 [subscript 2]); and 9.7 percent had gone to sea in their third year, and returned during their fourth year (4 [subscript 3]). Growth curves are given for all, age groups. The length-weight relationship may be expressed by the equation Weight = 0.0000373 x Length [superscript 2.8945]. As in the case of the chums, changes were noted during the run in the sex ratio and the percentage of age classes. Comparison with data from other localities revealed south-to-north changes in size and in abundance of different age groups, similar to those demonstrated for the chums. It is felt that the differences between races of a single river system will prove to be of the same nature as those here demonstrated between populations of different river systems. Knowledge of the differences between the races of salmons entering the Columbia River may be of extreme importance in the management of the fisheries. Close

• 50. [Article] Joint Production of Hydropower and Salmon - Optimal Bioeconomic Measure Combination in a Finnish Case River

  Suggested Bibliographic Reference: Challenging New Frontiers in the Global Seafood Sector: Proceedings of the Eighteenth Biennial Conference of the International Institute of Fisheries Economics and Trade, ...

  Citation × Citation Citation Abstract Copy Title: Joint Production of Hydropower and Salmon - Optimal Bioeconomic Measure Combination in a Finnish Case River Author: Nieminen, Emmi Year: 2016 Suggested Bibliographic Reference: Challenging New Frontiers in the Global Seafood Sector: Proceedings of the Eighteenth Biennial Conference of the International Institute of Fisheries Economics and Trade, July 11-15, 2016. Compiled by Stefani J. Evers and Ann L. Shriver. International Institute of Fisheries Economics and Trade (IIFET), Corvallis, 2016. Title: Joint Production of Hydropower and Salmon - Optimal Bioeconomic Measure Combination in a Finnish Case River Author: Nieminen, Emmi Year: 2016 Suggested Bibliographic Reference: Challenging New Frontiers in the Global Seafood Sector: Proceedings of the Eighteenth Biennial Conference of the International Institute of Fisheries Economics and Trade, July 11-15, 2016. Compiled by Stefani J. Evers and Ann L. Shriver. International Institute of Fisheries Economics and Trade (IIFET), Corvallis, 2016. Close