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• 1. [Article] An Experimental Investigation of Nest Reuse and Nest Site Selection in an Open-Cup Nesting Passerine

  Eastern Kingbirds (Tyrannus tyrannus) breed from coast to coast in North America and build open-cup nests in trees. They have been extensively studied across most of their range and have only on occasion ...

  Citation × Citation Citation Abstract Copy Title: An Experimental Investigation of Nest Reuse and Nest Site Selection in an Open-Cup Nesting Passerine Author: Cancellieri, Sarah A. Year: 2013 Eastern Kingbirds (Tyrannus tyrannus) breed from coast to coast in North America and build open-cup nests in trees. They have been extensively studied across most of their range and have only on occasion been documented to reuse a nest from a previous season. However, at Malheur National Wildlife Refuge (MNWR), located in southeastern Oregon, ~10 % of female Eastern Kingbirds reuse old nests of mainly American Robins (Turdus migratorius). In an attempt to address why nest reuse is so common at MNWR, I used artificial nests to evaluate two hypotheses as to why nest reuse is common in this breeding population. The first hypothesis states that Eastern Kingbirds reuse nests to save time and/or energy (TES) and the second one states nest reuse occurs because there is a shortage of suitable nest sites (NSS). I was able to reject the TES hypothesis because artificial nests provided no apparent reproductive benefits to Eastern Kingbirds, except that if a nest had failed it took less time to lay a replacement clutch after an initial failure if an artificial nest was used instead of building a new nest. A more reasonable explanation is that Eastern Kingbirds face a limited availability of suitable nest sites. With this in mind, I took vegetation measurements to address the hypothesis that Eastern Kingbirds make adaptive choices when selecting a nest site, in which case they would choose sites that increase their probability of breeding successfully. Successful nests, both natural and artificial, were placed higher in a tree and on a steeper angled nest branch than their failed counterparts. Those findings suggest that Eastern Kingbirds make adaptive choices when selecting a nest site. Title: An Experimental Investigation of Nest Reuse and Nest Site Selection in an Open-Cup Nesting Passerine Author: Cancellieri, Sarah A. Year: 2013 Eastern Kingbirds (Tyrannus tyrannus) breed from coast to coast in North America and build open-cup nests in trees. They have been extensively studied across most of their range and have only on occasion been documented to reuse a nest from a previous season. However, at Malheur National Wildlife Refuge (MNWR), located in southeastern Oregon, ~10 % of female Eastern Kingbirds reuse old nests of mainly American Robins (Turdus migratorius). In an attempt to address why nest reuse is so common at MNWR, I used artificial nests to evaluate two hypotheses as to why nest reuse is common in this breeding population. The first hypothesis states that Eastern Kingbirds reuse nests to save time and/or energy (TES) and the second one states nest reuse occurs because there is a shortage of suitable nest sites (NSS). I was able to reject the TES hypothesis because artificial nests provided no apparent reproductive benefits to Eastern Kingbirds, except that if a nest had failed it took less time to lay a replacement clutch after an initial failure if an artificial nest was used instead of building a new nest. A more reasonable explanation is that Eastern Kingbirds face a limited availability of suitable nest sites. With this in mind, I took vegetation measurements to address the hypothesis that Eastern Kingbirds make adaptive choices when selecting a nest site, in which case they would choose sites that increase their probability of breeding successfully. Successful nests, both natural and artificial, were placed higher in a tree and on a steeper angled nest branch than their failed counterparts. Those findings suggest that Eastern Kingbirds make adaptive choices when selecting a nest site. Close

• 2. [Article] Behavioral Observation and Paternal Investment of Eastern Kingbirds at Malheur National Wildlife Refuge

  We have been studying the population of Eastern Kingbirds breeding in riparian habitats in Malheur National Wildlife Refuge (eastern Oregon) since 2002. These efforts have created an ideal research environment ...

  Citation × Citation Citation Abstract Copy Title: Behavioral Observation and Paternal Investment of Eastern Kingbirds at Malheur National Wildlife Refuge Author: Chutter, Christopher Michael Year: 2014 We have been studying the population of Eastern Kingbirds breeding in riparian habitats in Malheur National Wildlife Refuge (eastern Oregon) since 2002. These efforts have created an ideal research environment wherein most adults in the population have been color banded and DNA sampled and, as part of other research projects, nearly all broods since 2003 have been paternity tested. I decided to use behavioral video recordings of parental nest behavior undertaken between 2003 and 2010 for two unrelated projects. First, I tested the effectiveness of video sampling nesting behavior (see below and chapter 2). Second, I tested whether male kingbirds were able to affect their level of paternal investment in accordance with their level or realized paternity (see below and chapter 3). Chapter 2 was split into three distinct questions: 1) are parental nesting behaviors repeatable, 2) is a one hour sample sufficient to capture variability in these behaviors, and 3) is the first hour of recording sufficient to capture variability in these behaviors. I found overwhelming evidence that the behaviors I measured were repeatable. This is truly important, for if repeatability was disproven, it would call into question the use of sampling throughout the field of animal behavior. I similarly found strong evidence that a one hour sample was sufficient to capture variability in parental behaviors. From this I was able to suggest that further sampling effort would be better spent increasing sample size rather than observation length. Testing whether the first hour of recording was sufficient to capture variability in parental behavior found more muddled results. While there generally was correlation between behavioral values in the first hour and those over a longer observation period, most behaviors were found to have significantly lower values in the first hour. I tested whether this was the result of a lingering observer effect or a natural effect of time of day and concluded that an observer effect was the more likely explanation. In chapter 3, I ran one of the more in depth and complicated tests for a relationship between paternal investment and realized paternity that I was able to find in the literature. I used the standard male feeding rate as a measure of male investment as well as a far more nuanced measure derived from the first Eigenvector of an analysis of six different paternal behaviors. These were both tested using Akaike's Information Criterion against a number of variables likely to affect parental behavior, including realized paternity. Ultimately, I found no evidence that kingbirds were able to affect paternal investment in response to lost paternity, and conclude that they likely had no means to assess realized paternity within the nest of their social mate. However, from the other parameters tested I was able to find that males increased their investment as brood size increased and as the female spent more time attending, but not feeding, the nestlings. Also, male investment decreased as territory density increased and flight feather length increased. Theory surrounding the variability of extrapair paternity in birds can be used to account for these results. In a denser population males are predicted to reduce paternal effort for one of two reasons: increased likelihood of paternity loss in his own nest and an increased investment in pursuing extrapair copulations of his own. Longer flight feathers are a sign of age and quality in kingbirds, so males with this trait are expected to spend more time pursuing extrapair copulations as they are more likely to be successful in this endeavor. Title: Behavioral Observation and Paternal Investment of Eastern Kingbirds at Malheur National Wildlife Refuge Author: Chutter, Christopher Michael Year: 2014 We have been studying the population of Eastern Kingbirds breeding in riparian habitats in Malheur National Wildlife Refuge (eastern Oregon) since 2002. These efforts have created an ideal research environment wherein most adults in the population have been color banded and DNA sampled and, as part of other research projects, nearly all broods since 2003 have been paternity tested. I decided to use behavioral video recordings of parental nest behavior undertaken between 2003 and 2010 for two unrelated projects. First, I tested the effectiveness of video sampling nesting behavior (see below and chapter 2). Second, I tested whether male kingbirds were able to affect their level of paternal investment in accordance with their level or realized paternity (see below and chapter 3). Chapter 2 was split into three distinct questions: 1) are parental nesting behaviors repeatable, 2) is a one hour sample sufficient to capture variability in these behaviors, and 3) is the first hour of recording sufficient to capture variability in these behaviors. I found overwhelming evidence that the behaviors I measured were repeatable. This is truly important, for if repeatability was disproven, it would call into question the use of sampling throughout the field of animal behavior. I similarly found strong evidence that a one hour sample was sufficient to capture variability in parental behaviors. From this I was able to suggest that further sampling effort would be better spent increasing sample size rather than observation length. Testing whether the first hour of recording was sufficient to capture variability in parental behavior found more muddled results. While there generally was correlation between behavioral values in the first hour and those over a longer observation period, most behaviors were found to have significantly lower values in the first hour. I tested whether this was the result of a lingering observer effect or a natural effect of time of day and concluded that an observer effect was the more likely explanation. In chapter 3, I ran one of the more in depth and complicated tests for a relationship between paternal investment and realized paternity that I was able to find in the literature. I used the standard male feeding rate as a measure of male investment as well as a far more nuanced measure derived from the first Eigenvector of an analysis of six different paternal behaviors. These were both tested using Akaike's Information Criterion against a number of variables likely to affect parental behavior, including realized paternity. Ultimately, I found no evidence that kingbirds were able to affect paternal investment in response to lost paternity, and conclude that they likely had no means to assess realized paternity within the nest of their social mate. However, from the other parameters tested I was able to find that males increased their investment as brood size increased and as the female spent more time attending, but not feeding, the nestlings. Also, male investment decreased as territory density increased and flight feather length increased. Theory surrounding the variability of extrapair paternity in birds can be used to account for these results. In a denser population males are predicted to reduce paternal effort for one of two reasons: increased likelihood of paternity loss in his own nest and an increased investment in pursuing extrapair copulations of his own. Longer flight feathers are a sign of age and quality in kingbirds, so males with this trait are expected to spend more time pursuing extrapair copulations as they are more likely to be successful in this endeavor. Close

• 3. [Article] The Biology of Eastern Kingbirds at Malheur National Wildlife Refuge: Survival, Reproduction, and Testosterone Secretion

  This dissertation presents the results of a study that I undertook to better understand the breeding biology of Eastern Kingbirds (hereafter, kingbirds) at Malheur National Wildlife Refuge in southeastern ...

  Citation × Citation Citation Abstract Copy Title: The Biology of Eastern Kingbirds at Malheur National Wildlife Refuge: Survival, Reproduction, and Testosterone Secretion Author: Redmond, Lucas J. Year: 2015 This dissertation presents the results of a study that I undertook to better understand the breeding biology of Eastern Kingbirds (hereafter, kingbirds) at Malheur National Wildlife Refuge in southeastern Oregon from 2003 to 2009. Kingbirds are long-distance migratory songbirds that breed across much of North America. This species is socially monogamous but, via frequent extra-pair copulations, is genetically polygamous. Kingbirds exhibit relatively high breeding site fidelity, often returning to the same tree to nest in subsequent years. Both members of a pair provide parental care, but there are often specific duties performed by both male and female kingbirds. For example, males typically perform much of the vigilant nest defense that this species is well known for and contribute, to an extent, to the feeding of nestlings. Females, on the other hand, are entirely responsible for incubating and brooding. Beginning in 2002, most adults within the population of kingbirds at Malheur had been banded with a unique combination of three colored plastic leg bands and an aluminum USFWS band. Also, as many nests were located in each year as possible, and any young that survived to fledging age were banded with a unique combination of leg bands. Considerable effort was then spent each year to locate as many banded individuals as possible, which allowed me to document adult and juvenile survival. Annual survival rates of adult male and female kingbirds in the population at Malheur did not differ and were relatively high at approximately 0.65. Juvenile survival rate was approximately 0.29, indicating that slightly less than one-third of nestlings survive the interval between fledging and their first potential breeding season. Resighting probability was high for both sexes, although higher for adult males (0.94) than for adult female (0.84). The latter finding is consistent with the higher site fidelity of males than of females. Resighting probabilities for juveniles were much lower at 0.68 and 0.40 for males and females, respectively. Again, this was expected because natal site fidelity is typically much lower than breeding site fidelity. Compared to most other reports, resighting probability and return rates of juvenile kingbirds was high, presumably because the riparian habitats where kingbirds breed at Malheur function as an ecological island surrounded by, for kingbirds, unsuitable high desert habitat. Thus, unless they disperse very long distances, the only option for juvenile kingbirds is to begin nesting on the refuge. The collection of blood samples from birds has become an increasingly common practice in ornithology. The data that can be gained from these samples allow a number of interesting questions to be asked such as understanding the genetic mating system of a species, patterns of hormone secretion, and discerning migratory pathways via stable isotope analysis. The volume of blood collected is usually small and was assumed to cause no long-term negative effects on sampled individuals. However, few studies have rigorously examined the effect of blood sampling on survival. I used a multistate mark-recapture analysis to assess the effect blood sampling on annual survival of kingbirds by combining the annual survival data described above with whether or not individuals were subjected to blood sample collection. The results of this analysis indicated that blood sampling had no effect on annual survival rates of kingbirds. Whether or not this is the case for other species remains to be seen. However, my results support the assumption that when done correctly, blood sampling has little to no long-term negative effects on birds. Comparative analyses of many bird species show that testosterone secretion exhibits fairly predictable patterns among breeding birds of different mating systems. Monogamous species reach a peak during mate attraction and the period of female fertility which is then followed by a sharp decline when young are in the nest. By contrast, males of polygynous species tend to maintain higher levels of testosterone throughout the breeding season to, presumably, maximize opportunities for acquiring additional mates. Kingbirds are socially monogamous but cryptically polygamous. However, because of high rates of extra-pair paternity, variation in reproductive success among males is much higher than what is expected for a monogamous species, and, instead, is similar to what has been reported for polygynous species. Therefore from 2005 to 2009 I measured testosterone concentrations from the blood samples collected from male kingbirds to determine the breeding season profile of testosterone secretion in kingbirds and to understand the factors that influence testosterone variation among individuals. Contrary to expected for a monogamous species, the testosterone profile of kingbirds did not exhibit the brief peak in testosterone followed by a precipitous decline. Testosterone peaked early in the season, but declined very slowly as the nest cycle progressed. I attributed this gradual decline in testosterone to the cryptically polygynous nature of the kingbird extra-pair mating system. I also found substantial variation in testosterone concentration among male kingbirds and was able to identify several factors contributing to this variation. As expected, testosterone declined as the breeding season progressed (independent of stage in nest cycle) and nest density increased, while increases in testosterone were correlated with the number of fertile females within the population. This suggests that male kingbirds were capable of modulating testosterone concentrations to both cope with an increase in conspecific density, but also to ready themselves for times when extra-pair copulations were more likely. Title: The Biology of Eastern Kingbirds at Malheur National Wildlife Refuge: Survival, Reproduction, and Testosterone Secretion Author: Redmond, Lucas J. Year: 2015 This dissertation presents the results of a study that I undertook to better understand the breeding biology of Eastern Kingbirds (hereafter, kingbirds) at Malheur National Wildlife Refuge in southeastern Oregon from 2003 to 2009. Kingbirds are long-distance migratory songbirds that breed across much of North America. This species is socially monogamous but, via frequent extra-pair copulations, is genetically polygamous. Kingbirds exhibit relatively high breeding site fidelity, often returning to the same tree to nest in subsequent years. Both members of a pair provide parental care, but there are often specific duties performed by both male and female kingbirds. For example, males typically perform much of the vigilant nest defense that this species is well known for and contribute, to an extent, to the feeding of nestlings. Females, on the other hand, are entirely responsible for incubating and brooding. Beginning in 2002, most adults within the population of kingbirds at Malheur had been banded with a unique combination of three colored plastic leg bands and an aluminum USFWS band. Also, as many nests were located in each year as possible, and any young that survived to fledging age were banded with a unique combination of leg bands. Considerable effort was then spent each year to locate as many banded individuals as possible, which allowed me to document adult and juvenile survival. Annual survival rates of adult male and female kingbirds in the population at Malheur did not differ and were relatively high at approximately 0.65. Juvenile survival rate was approximately 0.29, indicating that slightly less than one-third of nestlings survive the interval between fledging and their first potential breeding season. Resighting probability was high for both sexes, although higher for adult males (0.94) than for adult female (0.84). The latter finding is consistent with the higher site fidelity of males than of females. Resighting probabilities for juveniles were much lower at 0.68 and 0.40 for males and females, respectively. Again, this was expected because natal site fidelity is typically much lower than breeding site fidelity. Compared to most other reports, resighting probability and return rates of juvenile kingbirds was high, presumably because the riparian habitats where kingbirds breed at Malheur function as an ecological island surrounded by, for kingbirds, unsuitable high desert habitat. Thus, unless they disperse very long distances, the only option for juvenile kingbirds is to begin nesting on the refuge. The collection of blood samples from birds has become an increasingly common practice in ornithology. The data that can be gained from these samples allow a number of interesting questions to be asked such as understanding the genetic mating system of a species, patterns of hormone secretion, and discerning migratory pathways via stable isotope analysis. The volume of blood collected is usually small and was assumed to cause no long-term negative effects on sampled individuals. However, few studies have rigorously examined the effect of blood sampling on survival. I used a multistate mark-recapture analysis to assess the effect blood sampling on annual survival of kingbirds by combining the annual survival data described above with whether or not individuals were subjected to blood sample collection. The results of this analysis indicated that blood sampling had no effect on annual survival rates of kingbirds. Whether or not this is the case for other species remains to be seen. However, my results support the assumption that when done correctly, blood sampling has little to no long-term negative effects on birds. Comparative analyses of many bird species show that testosterone secretion exhibits fairly predictable patterns among breeding birds of different mating systems. Monogamous species reach a peak during mate attraction and the period of female fertility which is then followed by a sharp decline when young are in the nest. By contrast, males of polygynous species tend to maintain higher levels of testosterone throughout the breeding season to, presumably, maximize opportunities for acquiring additional mates. Kingbirds are socially monogamous but cryptically polygamous. However, because of high rates of extra-pair paternity, variation in reproductive success among males is much higher than what is expected for a monogamous species, and, instead, is similar to what has been reported for polygynous species. Therefore from 2005 to 2009 I measured testosterone concentrations from the blood samples collected from male kingbirds to determine the breeding season profile of testosterone secretion in kingbirds and to understand the factors that influence testosterone variation among individuals. Contrary to expected for a monogamous species, the testosterone profile of kingbirds did not exhibit the brief peak in testosterone followed by a precipitous decline. Testosterone peaked early in the season, but declined very slowly as the nest cycle progressed. I attributed this gradual decline in testosterone to the cryptically polygynous nature of the kingbird extra-pair mating system. I also found substantial variation in testosterone concentration among male kingbirds and was able to identify several factors contributing to this variation. As expected, testosterone declined as the breeding season progressed (independent of stage in nest cycle) and nest density increased, while increases in testosterone were correlated with the number of fertile females within the population. This suggests that male kingbirds were capable of modulating testosterone concentrations to both cope with an increase in conspecific density, but also to ready themselves for times when extra-pair copulations were more likely. Close

• 4. [Article] Archaeological Investigations at 45CL1 Cathlapotle (1991-1996) , Ridgefield National Wildlife Refuge Clark County, Washington A Preliminary Report

  Archaeological investigations at site 45CL1, Clark County, Washington, demonstrate that the locality is a very large (c 1.5ha), deeply stratified (2-4m) town site with an occupation spanning at least 1000 ...

  Citation × Citation Citation Abstract Copy Title: Archaeological Investigations at 45CL1 Cathlapotle (1991-1996) , Ridgefield National Wildlife Refuge Clark County, Washington A Preliminary Report Author: Ames, Kenneth M., Smith, Cameron M., Cornett, William L., Sobel, Elizabeth A., Hamilton, Stephen C., Wolf, John, Raetz, Doria, U.S. Fish and Wildlife Service. Region 1, Portland State University. Department of Anthropology Year: 1999 Archaeological investigations at site 45CL1, Clark County, Washington, demonstrate that the locality is a very large (c 1.5ha), deeply stratified (2-4m) town site with an occupation spanning at least 1000 years (c. AD 1000 to 1840). Six large, complex depressions have been mapped. Test excavations show that these depressions represent the semisubterranean portions of residential structures, probably large plankhouses of the type common on the Lower Columbia River and the Northwest Coast in aboriginal times. The depressions may represent as many as 11 such dwellings. A seventh depression is deeply buried beneath midden deposits. The cultural deposits contain very high densities of artifacts, ecofacts (including both faunal and floral remains), debris and features. The site is near the Columbia River on a very active flood plain, resulting in site stratigraphy produced by a combination of active cultural and alluvial depositional processes. Site 45CL1, given its location and size, is the best candidate to be the site of Cathlapotle, a Middle Chinookan town visited by Lewis and Clark in 1806, as well as by other early Europeans in the area. The site is extraordinarily well preserved, having undergone only minor alterations since its abandonment, probably in the third or fourth decade of the 19th century AD. Title: Archaeological Investigations at 45CL1 Cathlapotle (1991-1996) , Ridgefield National Wildlife Refuge Clark County, Washington A Preliminary Report Author: Ames, Kenneth M., Smith, Cameron M., Cornett, William L., Sobel, Elizabeth A., Hamilton, Stephen C., Wolf, John, Raetz, Doria, U.S. Fish and Wildlife Service. Region 1, Portland State University. Department of Anthropology Year: 1999 Archaeological investigations at site 45CL1, Clark County, Washington, demonstrate that the locality is a very large (c 1.5ha), deeply stratified (2-4m) town site with an occupation spanning at least 1000 years (c. AD 1000 to 1840). Six large, complex depressions have been mapped. Test excavations show that these depressions represent the semisubterranean portions of residential structures, probably large plankhouses of the type common on the Lower Columbia River and the Northwest Coast in aboriginal times. The depressions may represent as many as 11 such dwellings. A seventh depression is deeply buried beneath midden deposits. The cultural deposits contain very high densities of artifacts, ecofacts (including both faunal and floral remains), debris and features. The site is near the Columbia River on a very active flood plain, resulting in site stratigraphy produced by a combination of active cultural and alluvial depositional processes. Site 45CL1, given its location and size, is the best candidate to be the site of Cathlapotle, a Middle Chinookan town visited by Lewis and Clark in 1806, as well as by other early Europeans in the area. The site is extraordinarily well preserved, having undergone only minor alterations since its abandonment, probably in the third or fourth decade of the 19th century AD. Close

• 5. [Article] Finding and Dating Cathlapotle

  The people of the Cathlapotle town played a significant role in the fur trade era history of the Lower Columbia River, including Lewis and Clark’s visit on March 29th, 1806. Archaeologists and others have ...

  Citation × Citation Citation Abstract Copy Title: Finding and Dating Cathlapotle Author: Ames, Kenneth M., Sobel, Elizabeth A. Year: 2009 The people of the Cathlapotle town played a significant role in the fur trade era history of the Lower Columbia River, including Lewis and Clark’s visit on March 29th, 1806. Archaeologists and others have sought the town’s location for years. Long-term research has established that archaeological site 45CL1 on the US Fish and Wildlife Refuge near Ridgefield, Washington is Cathlapotle. This determination is based on the close match between site details with various ethnohistoric accounts of Cathlapotle. The site was occupied by ca. AD 1450 and probably moved there from another nearby location. It was abandoned sometime in the 1830s or 1840s. This chronology is based on 54 radiocarbon dates, historic trade goods including glass beads and ceramics, and documentary accounts. Title: Finding and Dating Cathlapotle Author: Ames, Kenneth M., Sobel, Elizabeth A. Year: 2009 The people of the Cathlapotle town played a significant role in the fur trade era history of the Lower Columbia River, including Lewis and Clark’s visit on March 29th, 1806. Archaeologists and others have sought the town’s location for years. Long-term research has established that archaeological site 45CL1 on the US Fish and Wildlife Refuge near Ridgefield, Washington is Cathlapotle. This determination is based on the close match between site details with various ethnohistoric accounts of Cathlapotle. The site was occupied by ca. AD 1450 and probably moved there from another nearby location. It was abandoned sometime in the 1830s or 1840s. This chronology is based on 54 radiocarbon dates, historic trade goods including glass beads and ceramics, and documentary accounts. Close

• 6. [Article] Experimental Analysis of Nest-Site Choice and its Relationship to Nest Success in an Open-Cup–Nesting Passerine

  Nest placement presumably reflects selection for secure sites to minimize failure. Most tests of this hypothesis, however, have failed to support it. We used artificial nests (ARTs) to experimentally evaluate ...

  Citation × Citation Citation Abstract Copy Title: Experimental Analysis of Nest-Site Choice and its Relationship to Nest Success in an Open-Cup–Nesting Passerine Author: Cancellieri, Sarah, Murphy, Michael T. Year: 2014 Nest placement presumably reflects selection for secure sites to minimize failure. Most tests of this hypothesis, however, have failed to support it. We used artificial nests (ARTs) to experimentally evaluate nest-site-choice behavior by an open-cup–nesting bird, the Eastern Kingbird (Tyrannus tyrannus). In 2010 and 2011, we placed ARTs in trees in the riparian zone at Malheur National Wildlife Refuge, Oregon, USA, to test whether (1) characteristics describing the physical location in trees of used and unused ARTs differed, (2) used ART sites more closely resembled naturally chosen sites, (3) successful natural nests (NATs) and successful ARTs were similarly located along the major axis describing nest placement, and (4) unused ARTs resembled failed NATs. Used and unused ART sites differed, but unused ART sites were more similar to NAT sites. The latter unexpected result occurred because (1) unused ARTs were located at sites between more heavily used higher and lower locations and (2) most kingbirds nesting at lower locations used ARTs instead of building their own nest. In both ARTs and NATs, differences between successful and failed nests exhibited the same pattern for most nest-site variables, and the major gradient describing nest location was the same; successful nests tended to be placed on more vertically oriented branches that were placed closer to the top of the tree. Kingbird nest placement was thus selective. However, extensive overlap in the locations in trees of failed NATs and both successful and unused ARTs suggests that other factors, such as macrohabitat characteristics or prior experience of individual birds with particular nest sites, may have influenced success and/or decisions to use or reject nests in particular locations. Thus, consideration of phenomena beyond the nest site itself may be required to fully understand the process of nest-site choice in birds. Title: Experimental Analysis of Nest-Site Choice and its Relationship to Nest Success in an Open-Cup–Nesting Passerine Author: Cancellieri, Sarah, Murphy, Michael T. Year: 2014 Nest placement presumably reflects selection for secure sites to minimize failure. Most tests of this hypothesis, however, have failed to support it. We used artificial nests (ARTs) to experimentally evaluate nest-site-choice behavior by an open-cup–nesting bird, the Eastern Kingbird (Tyrannus tyrannus). In 2010 and 2011, we placed ARTs in trees in the riparian zone at Malheur National Wildlife Refuge, Oregon, USA, to test whether (1) characteristics describing the physical location in trees of used and unused ARTs differed, (2) used ART sites more closely resembled naturally chosen sites, (3) successful natural nests (NATs) and successful ARTs were similarly located along the major axis describing nest placement, and (4) unused ARTs resembled failed NATs. Used and unused ART sites differed, but unused ART sites were more similar to NAT sites. The latter unexpected result occurred because (1) unused ARTs were located at sites between more heavily used higher and lower locations and (2) most kingbirds nesting at lower locations used ARTs instead of building their own nest. In both ARTs and NATs, differences between successful and failed nests exhibited the same pattern for most nest-site variables, and the major gradient describing nest location was the same; successful nests tended to be placed on more vertically oriented branches that were placed closer to the top of the tree. Kingbird nest placement was thus selective. However, extensive overlap in the locations in trees of failed NATs and both successful and unused ARTs suggests that other factors, such as macrohabitat characteristics or prior experience of individual birds with particular nest sites, may have influenced success and/or decisions to use or reject nests in particular locations. Thus, consideration of phenomena beyond the nest site itself may be required to fully understand the process of nest-site choice in birds. Close

• 7. [Article] Characterization of the Shallow Subsurface Geohydrology of the Ni-les'tun Unit on the Bandon Marsh National Wildlife Refuge

  The Bandon Marsh is a large marsh restoration project located in southwest Oregon. The land has been previously used for dairy farms and much of the marsh was used for cattle grazing. The goal of the restoration ...

  Citation × Citation Citation Abstract Copy Title: Characterization of the Shallow Subsurface Geohydrology of the Ni-les'tun Unit on the Bandon Marsh National Wildlife Refuge Author: Beard, Christopher Madison Year: 2013 The Bandon Marsh is a large marsh restoration project located in southwest Oregon. The land has been previously used for dairy farms and much of the marsh was used for cattle grazing. The goal of the restoration is to recreate a natural habitat for shorebirds and fish. The purpose of this thesis is to gather and analyze data on the geology of the marsh and both ground and surface water quality to evaluate its ability to support biology. The US Fish and Wildlife Service began restoration on the Bandon marsh in 2005. Research on the Ni-les'tun Unit began in that same year with Geoprobe coring, Ground Penetrating Radar (GPR), piezometer well installation, and surface water analysis. Twelve cores were taken with the Geoprobe in 2006 and later analyzed for this thesis. A total of 50 GPR lines were run across the marsh ranging from ~25 m to 1,200 m in length for a total of about 9,700 meters (9.7 km). The ten piezometer wells were sampled to collect groundwater pH, oxidation-reduction potential, salinity, temperature, and well water elevation. Twelve channel sampling sites were chosen to be sampled in three field runs taking place in winter 2011, summer 2011, and winter 2012. The summer 2011 and winter 2012 sampling runs were done after tide gates had been removed from the marsh. The Geoprobe core samples were retrieved for extensive lab analysis. Analyses included; bulk density, porosity, permeability, and grain size studies. Lab results revealed a low permeability fine grained upper sedimentary unit ranging from ~0.5 to ~2.5 meters depth with a coarser grained higher permeability lower unit. GPR lines were analyzed for groundwater surface depths and buried channel cut and fill features (Peterson et. al., 2004). GPR results showed a fairly shallow groundwater surface around 0.5 meters in the north down to 2.0 meters in the south. The piezometer wells showed a seasonal variation in well water elevation with higher elevations in the winter and lower in the summer. Well water chemistry showed both seasonal and spatial variations. Values for pH and dissolved oxygen were lower in the north side of the marsh and higher in the south side, and pH was higher in the summer than in the winter. Wells that were more proximal (within ~400 m) to the Coquille River Estuary showed higher conductivity (salinity) values than those that were more distal (greater than ~400 m). Channel sampling results showed similar trends as the groundwater results. The lower pH values tended to be in the north side of the marsh as well as lower dissolved oxygen values. Again, there was a spatial variability in conductivity with the higher values found closer to the Coquille River. Conductivity in the channel water showed a great seasonal variability with the highest values occurring in the summer time and much lower values in the winter. As expected, channel water increased about 10 °C on average from winter to summer. The results of all of these observations and analyses are combined to put together a generalized flow model showing the different water inputs into the channel waters on the Ni-les'tun Unit. Once these inputs were derived, conclusions can be made on the quality of the water and its ability to support small aquatic life. Title: Characterization of the Shallow Subsurface Geohydrology of the Ni-les'tun Unit on the Bandon Marsh National Wildlife Refuge Author: Beard, Christopher Madison Year: 2013 The Bandon Marsh is a large marsh restoration project located in southwest Oregon. The land has been previously used for dairy farms and much of the marsh was used for cattle grazing. The goal of the restoration is to recreate a natural habitat for shorebirds and fish. The purpose of this thesis is to gather and analyze data on the geology of the marsh and both ground and surface water quality to evaluate its ability to support biology. The US Fish and Wildlife Service began restoration on the Bandon marsh in 2005. Research on the Ni-les'tun Unit began in that same year with Geoprobe coring, Ground Penetrating Radar (GPR), piezometer well installation, and surface water analysis. Twelve cores were taken with the Geoprobe in 2006 and later analyzed for this thesis. A total of 50 GPR lines were run across the marsh ranging from ~25 m to 1,200 m in length for a total of about 9,700 meters (9.7 km). The ten piezometer wells were sampled to collect groundwater pH, oxidation-reduction potential, salinity, temperature, and well water elevation. Twelve channel sampling sites were chosen to be sampled in three field runs taking place in winter 2011, summer 2011, and winter 2012. The summer 2011 and winter 2012 sampling runs were done after tide gates had been removed from the marsh. The Geoprobe core samples were retrieved for extensive lab analysis. Analyses included; bulk density, porosity, permeability, and grain size studies. Lab results revealed a low permeability fine grained upper sedimentary unit ranging from ~0.5 to ~2.5 meters depth with a coarser grained higher permeability lower unit. GPR lines were analyzed for groundwater surface depths and buried channel cut and fill features (Peterson et. al., 2004). GPR results showed a fairly shallow groundwater surface around 0.5 meters in the north down to 2.0 meters in the south. The piezometer wells showed a seasonal variation in well water elevation with higher elevations in the winter and lower in the summer. Well water chemistry showed both seasonal and spatial variations. Values for pH and dissolved oxygen were lower in the north side of the marsh and higher in the south side, and pH was higher in the summer than in the winter. Wells that were more proximal (within ~400 m) to the Coquille River Estuary showed higher conductivity (salinity) values than those that were more distal (greater than ~400 m). Channel sampling results showed similar trends as the groundwater results. The lower pH values tended to be in the north side of the marsh as well as lower dissolved oxygen values. Again, there was a spatial variability in conductivity with the higher values found closer to the Coquille River. Conductivity in the channel water showed a great seasonal variability with the highest values occurring in the summer time and much lower values in the winter. As expected, channel water increased about 10 °C on average from winter to summer. The results of all of these observations and analyses are combined to put together a generalized flow model showing the different water inputs into the channel waters on the Ni-les'tun Unit. Once these inputs were derived, conclusions can be made on the quality of the water and its ability to support small aquatic life. Close

• 8. [Article] A Survey for Leucocytozoon Simondi Mathis and Leger (1910) in Canada Geese of the Malheur Wildlife Refuge

  Blood smears of blood collected in 1971 and 1972 from 291 Canada geese at Malheur National Wildlife Refuge were examined to determine the incidence of the avian malarial parasite, Leucocytozoon simondi. ...

  Citation × Citation Citation Abstract Copy Title: A Survey for Leucocytozoon Simondi Mathis and Leger (1910) in Canada Geese of the Malheur Wildlife Refuge Author: Snively, John Val Year: 1973 Blood smears of blood collected in 1971 and 1972 from 291 Canada geese at Malheur National Wildlife Refuge were examined to determine the incidence of the avian malarial parasite, Leucocytozoon simondi. No parasites were found. Absence of blood parasites suggests that birds using migration routes to the north and northeast of Malheur are free of the disease. The Leucocytozoon infections known in California waterfowl may be attributable to sources within the flyway east or northeast of California, to infected birds crossing over from other flyways, or abnormal conditions in the molting grounds of sub-adult and non-nesting birds where major flyways converge in northcentral Canada. Title: A Survey for Leucocytozoon Simondi Mathis and Leger (1910) in Canada Geese of the Malheur Wildlife Refuge Author: Snively, John Val Year: 1973 Blood smears of blood collected in 1971 and 1972 from 291 Canada geese at Malheur National Wildlife Refuge were examined to determine the incidence of the avian malarial parasite, Leucocytozoon simondi. No parasites were found. Absence of blood parasites suggests that birds using migration routes to the north and northeast of Malheur are free of the disease. The Leucocytozoon infections known in California waterfowl may be attributable to sources within the flyway east or northeast of California, to infected birds crossing over from other flyways, or abnormal conditions in the molting grounds of sub-adult and non-nesting birds where major flyways converge in northcentral Canada. Close

• 9. [Article] Functional analysis of Great Basin projectile points

  This paper presents a refined methodology for distinguishing the stone points of arrows from the stone points of spear thrower darts in archaeological assemblages from the Great Basin. The methodology ...

  Citation × Citation Citation Abstract Copy Title: Functional analysis of Great Basin projectile points Author: Wiggin, Roger Year: 1979 This paper presents a refined methodology for distinguishing the stone points of arrows from the stone points of spear thrower darts in archaeological assemblages from the Great Basin. The methodology was developed from a sample of 111 complete stone points collected during the archaeological reconnaissance and testing of the Malheur National Wildlife Refuge, Harney County, Oregon, 1971-1974. The points were measured for five metric variables and frequency histograms of each variable were generated by computer. The effectiveness of each variable as an index of functional class (i.e., arrow point vs. dart point) was judged on the basis of the apparent bimodality of its curve and the t-test. The variables of weight and neck width showed significant bimodality, confirming the work of previous researchers. A new variable, stem thickness, also showed significant bimodality. All three can be considered useful indices of functional class. They also appear to be functionally independent. An attempt was made at developing a multivariable index of point function, using a multivariable clustering analysis, the K-means test. Results of the K-means test were inconclusive but showed promise for further development. Title: Functional analysis of Great Basin projectile points Author: Wiggin, Roger Year: 1979 This paper presents a refined methodology for distinguishing the stone points of arrows from the stone points of spear thrower darts in archaeological assemblages from the Great Basin. The methodology was developed from a sample of 111 complete stone points collected during the archaeological reconnaissance and testing of the Malheur National Wildlife Refuge, Harney County, Oregon, 1971-1974. The points were measured for five metric variables and frequency histograms of each variable were generated by computer. The effectiveness of each variable as an index of functional class (i.e., arrow point vs. dart point) was judged on the basis of the apparent bimodality of its curve and the t-test. The variables of weight and neck width showed significant bimodality, confirming the work of previous researchers. A new variable, stem thickness, also showed significant bimodality. All three can be considered useful indices of functional class. They also appear to be functionally independent. An attempt was made at developing a multivariable index of point function, using a multivariable clustering analysis, the K-means test. Results of the K-means test were inconclusive but showed promise for further development. Close

• 10. [Article] Investigating the Link Between Surface Water and Groundwater in the Tule Lake Subbasin, Oregon and California

  Water allocation in the upper Klamath Basin of Oregon and California has been challenging. Irrigators have increasingly turned to groundwater to make up for surface water shortages because of shifts in ...

  Citation × Citation Citation Abstract Copy Title: Investigating the Link Between Surface Water and Groundwater in the Tule Lake Subbasin, Oregon and California Author: Pischel, Esther Maria Year: 2014 Water allocation in the upper Klamath Basin of Oregon and California has been challenging. Irrigators have increasingly turned to groundwater to make up for surface water shortages because of shifts in allocation toward in-stream flows for Endangered Species Act listed fishes. The largest increase in groundwater pumping has been in and around the Bureau of Reclamation's Klamath Irrigation Project, which includes the Tule Lake subbasin in the southern part of the upper Klamath Basin. Previous groundwater flow model simulations indicate that water level declines from pumping may result in decreased flow to agricultural drains in the Tule Lake subbasin. Agricultural drains on the Klamath Project are an important source of water for downstream irrigators and for the Tule Lake and Lower Klamath Lake National Wildlife Refuges. To better assess the impact of increased pumping on drain flow and on the water balance of the groundwater system, flow data from agricultural drains were evaluated to investigate the changes that have taken place in groundwater discharge to drains since pumping volumes increased. Additionally, a fine-grid groundwater model of the Tule Lake subbasin was developed based on the existing regional flow model. The fine-grid model has sufficient vertical and horizontal resolution to simulate vertical head gradients, takes advantage of time-series data from 38 observation wells for model calibration, and allows agricultural drains to be more explicitly represented. Results of the drain flow analysis show that the groundwater discharge to agricultural drains has decreased by approximately 4000 hectare-meters from the 1997-2000 average discharge. Most of this decrease takes place in the northern and southeastern portions of the subbasin. Results of the groundwater model show that the initial source of water to wells is groundwater storage. By 2006, approximately 56% of the water from wells is sourced from agricultural drains. Title: Investigating the Link Between Surface Water and Groundwater in the Tule Lake Subbasin, Oregon and California Author: Pischel, Esther Maria Year: 2014 Water allocation in the upper Klamath Basin of Oregon and California has been challenging. Irrigators have increasingly turned to groundwater to make up for surface water shortages because of shifts in allocation toward in-stream flows for Endangered Species Act listed fishes. The largest increase in groundwater pumping has been in and around the Bureau of Reclamation's Klamath Irrigation Project, which includes the Tule Lake subbasin in the southern part of the upper Klamath Basin. Previous groundwater flow model simulations indicate that water level declines from pumping may result in decreased flow to agricultural drains in the Tule Lake subbasin. Agricultural drains on the Klamath Project are an important source of water for downstream irrigators and for the Tule Lake and Lower Klamath Lake National Wildlife Refuges. To better assess the impact of increased pumping on drain flow and on the water balance of the groundwater system, flow data from agricultural drains were evaluated to investigate the changes that have taken place in groundwater discharge to drains since pumping volumes increased. Additionally, a fine-grid groundwater model of the Tule Lake subbasin was developed based on the existing regional flow model. The fine-grid model has sufficient vertical and horizontal resolution to simulate vertical head gradients, takes advantage of time-series data from 38 observation wells for model calibration, and allows agricultural drains to be more explicitly represented. Results of the drain flow analysis show that the groundwater discharge to agricultural drains has decreased by approximately 4000 hectare-meters from the 1997-2000 average discharge. Most of this decrease takes place in the northern and southeastern portions of the subbasin. Results of the groundwater model show that the initial source of water to wells is groundwater storage. By 2006, approximately 56% of the water from wells is sourced from agricultural drains. Close