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• 1. [Article] The Landscape: Sauvie Island

  A brief overview of the geography, history and outlook for Sauvie Island, an area situated at the confluence of the Willamette and Columbia Rivers that is important as a wildlife refuge, farming areas, ...

  Citation × Citation Citation Abstract Copy Title: The Landscape: Sauvie Island Author: Roth, Rick Year: 2001 A brief overview of the geography, history and outlook for Sauvie Island, an area situated at the confluence of the Willamette and Columbia Rivers that is important as a wildlife refuge, farming areas, and for recreation. Title: The Landscape: Sauvie Island Author: Roth, Rick Year: 2001 A brief overview of the geography, history and outlook for Sauvie Island, an area situated at the confluence of the Willamette and Columbia Rivers that is important as a wildlife refuge, farming areas, and for recreation. Close

• 2. [Article] Wet Meadow Plant Associations, Double O Unit, Malheur National Wildlife Refuge, Harney County, Oregon

  This report summarizes vegetation data collected in July 2015 in wet meadow and marshy habitats on the Double O Unit of Malheur National Wildlife Refuge (MNWR). Because vegetation sampled at the Double ...

  Citation × Citation Citation Abstract Copy Title: Wet Meadow Plant Associations, Double O Unit, Malheur National Wildlife Refuge, Harney County, Oregon Author: Christy, John A. Year: 2016 This report summarizes vegetation data collected in July 2015 in wet meadow and marshy habitats on the Double O Unit of Malheur National Wildlife Refuge (MNWR). Because vegetation sampled at the Double O was wetter and more alkaline than wet meadows sampled at the south end of the refuge in 2012 and 2013 (Christy 2014), data from the Double O Unit were analyzed and summarized separately. A total of 83 plots were sampled in 2015, and analysis of the data identified 14 plant associations: Alopecurus aequalis - Juncus balticus, Alopecurus pratensis - Potentilla anserina, Carex praegracilis - Juncus balticus, Cicuta douglasii - Carex nebrascensis, Distichlis spicata - Amphiscirpus nevadensis, Distichlis spicata - Nitrophila occidentalis, Eleocharis palustris - Juncus balticus, Eleocharis rostellata, Juncus balticus - Glaux maritima, Hippuris vulgaris - Triglochin maritima, Leymus triticoides - Juncus balticus, Schoenoplectus americanus, Spartina gracilis, and Triglochin maritima. Plant associations spanned a wetland gradient from seasonally moist to seasonally or perennially flooded, but surface water had left most stands at time of sampling. Mean Wetland Indicator Status scores and species composition help to place the plant associations within gradients in soil moisture and alkalinity. Seven of the 14 plant associations are listed in the International Vegetation Classification, and the remaining types are provisional Title: Wet Meadow Plant Associations, Double O Unit, Malheur National Wildlife Refuge, Harney County, Oregon Author: Christy, John A. Year: 2016 This report summarizes vegetation data collected in July 2015 in wet meadow and marshy habitats on the Double O Unit of Malheur National Wildlife Refuge (MNWR). Because vegetation sampled at the Double O was wetter and more alkaline than wet meadows sampled at the south end of the refuge in 2012 and 2013 (Christy 2014), data from the Double O Unit were analyzed and summarized separately. A total of 83 plots were sampled in 2015, and analysis of the data identified 14 plant associations: Alopecurus aequalis - Juncus balticus, Alopecurus pratensis - Potentilla anserina, Carex praegracilis - Juncus balticus, Cicuta douglasii - Carex nebrascensis, Distichlis spicata - Amphiscirpus nevadensis, Distichlis spicata - Nitrophila occidentalis, Eleocharis palustris - Juncus balticus, Eleocharis rostellata, Juncus balticus - Glaux maritima, Hippuris vulgaris - Triglochin maritima, Leymus triticoides - Juncus balticus, Schoenoplectus americanus, Spartina gracilis, and Triglochin maritima. Plant associations spanned a wetland gradient from seasonally moist to seasonally or perennially flooded, but surface water had left most stands at time of sampling. Mean Wetland Indicator Status scores and species composition help to place the plant associations within gradients in soil moisture and alkalinity. Seven of the 14 plant associations are listed in the International Vegetation Classification, and the remaining types are provisional Close

• 3. [Article] An Experimental Investigation of Nest Reuse and Nest Site Selection in an Open-Cup Nesting Passerine

  Eastern Kingbirds (Tyrannus tyrannus) breed from coast to coast in North America and build open-cup nests in trees. They have been extensively studied across most of their range and have only on occasion ...

  Citation × Citation Citation Abstract Copy Title: An Experimental Investigation of Nest Reuse and Nest Site Selection in an Open-Cup Nesting Passerine Author: Cancellieri, Sarah A. Year: 2013 Eastern Kingbirds (Tyrannus tyrannus) breed from coast to coast in North America and build open-cup nests in trees. They have been extensively studied across most of their range and have only on occasion been documented to reuse a nest from a previous season. However, at Malheur National Wildlife Refuge (MNWR), located in southeastern Oregon, ~10 % of female Eastern Kingbirds reuse old nests of mainly American Robins (Turdus migratorius). In an attempt to address why nest reuse is so common at MNWR, I used artificial nests to evaluate two hypotheses as to why nest reuse is common in this breeding population. The first hypothesis states that Eastern Kingbirds reuse nests to save time and/or energy (TES) and the second one states nest reuse occurs because there is a shortage of suitable nest sites (NSS). I was able to reject the TES hypothesis because artificial nests provided no apparent reproductive benefits to Eastern Kingbirds, except that if a nest had failed it took less time to lay a replacement clutch after an initial failure if an artificial nest was used instead of building a new nest. A more reasonable explanation is that Eastern Kingbirds face a limited availability of suitable nest sites. With this in mind, I took vegetation measurements to address the hypothesis that Eastern Kingbirds make adaptive choices when selecting a nest site, in which case they would choose sites that increase their probability of breeding successfully. Successful nests, both natural and artificial, were placed higher in a tree and on a steeper angled nest branch than their failed counterparts. Those findings suggest that Eastern Kingbirds make adaptive choices when selecting a nest site. Title: An Experimental Investigation of Nest Reuse and Nest Site Selection in an Open-Cup Nesting Passerine Author: Cancellieri, Sarah A. Year: 2013 Eastern Kingbirds (Tyrannus tyrannus) breed from coast to coast in North America and build open-cup nests in trees. They have been extensively studied across most of their range and have only on occasion been documented to reuse a nest from a previous season. However, at Malheur National Wildlife Refuge (MNWR), located in southeastern Oregon, ~10 % of female Eastern Kingbirds reuse old nests of mainly American Robins (Turdus migratorius). In an attempt to address why nest reuse is so common at MNWR, I used artificial nests to evaluate two hypotheses as to why nest reuse is common in this breeding population. The first hypothesis states that Eastern Kingbirds reuse nests to save time and/or energy (TES) and the second one states nest reuse occurs because there is a shortage of suitable nest sites (NSS). I was able to reject the TES hypothesis because artificial nests provided no apparent reproductive benefits to Eastern Kingbirds, except that if a nest had failed it took less time to lay a replacement clutch after an initial failure if an artificial nest was used instead of building a new nest. A more reasonable explanation is that Eastern Kingbirds face a limited availability of suitable nest sites. With this in mind, I took vegetation measurements to address the hypothesis that Eastern Kingbirds make adaptive choices when selecting a nest site, in which case they would choose sites that increase their probability of breeding successfully. Successful nests, both natural and artificial, were placed higher in a tree and on a steeper angled nest branch than their failed counterparts. Those findings suggest that Eastern Kingbirds make adaptive choices when selecting a nest site. Close

• 4. [Article] Wet Meadow Plant Associations, Malheur National Wildlife Refuge, Harney County, Oregon

  In July 2012, we sampled 131 plots in wet meadow habitat at the southern end of the Malheur National Wildlife Refuge. Analysis of the data identified eleven different plant associations: Alopecurus pratensis, ...

  Citation × Citation Citation Abstract Copy Title: Wet Meadow Plant Associations, Malheur National Wildlife Refuge, Harney County, Oregon Author: Christy, John A. Year: 2013 In July 2012, we sampled 131 plots in wet meadow habitat at the southern end of the Malheur National Wildlife Refuge. Analysis of the data identified eleven different plant associations: Alopecurus pratensis, Carex aquatilis var. aquatilis, Carex nebrascensis, Carex pellita, Carex praegracilis, Carex sheldonii, Distichlis spicata, Juncus balticus, Leymus triticoides, Phalaris arundinacea, and Sparganium eurycarpum. Plant associations spanned a wetland gradient from seasonally moist to seasonally or perennially flooded, but surface water had left most stands at time of sampling. Mean Wetland Indicator Status scores help to place the plant associations within gradients in soil moisture and alkalinity. The Alopecurus pratensis and Phalaris arundinacea associations are dominated by exotic, invasive species and appear to be outcompeting native plant associations. Phases of the Carex praegracilis association appear to be replaced by the Alopecurus pratensis association, while the Carex pellita, Juncus balticus, and Leymus triticoides associations may be replaced by the Phalaris arundinacea association. The Carex aquatilis and Sparganium eurycarpum associations were undersampled and provide only an estimate of composition. Most associations are consistent with what has been included in the National Vegetation Classification. Title: Wet Meadow Plant Associations, Malheur National Wildlife Refuge, Harney County, Oregon Author: Christy, John A. Year: 2013 In July 2012, we sampled 131 plots in wet meadow habitat at the southern end of the Malheur National Wildlife Refuge. Analysis of the data identified eleven different plant associations: Alopecurus pratensis, Carex aquatilis var. aquatilis, Carex nebrascensis, Carex pellita, Carex praegracilis, Carex sheldonii, Distichlis spicata, Juncus balticus, Leymus triticoides, Phalaris arundinacea, and Sparganium eurycarpum. Plant associations spanned a wetland gradient from seasonally moist to seasonally or perennially flooded, but surface water had left most stands at time of sampling. Mean Wetland Indicator Status scores help to place the plant associations within gradients in soil moisture and alkalinity. The Alopecurus pratensis and Phalaris arundinacea associations are dominated by exotic, invasive species and appear to be outcompeting native plant associations. Phases of the Carex praegracilis association appear to be replaced by the Alopecurus pratensis association, while the Carex pellita, Juncus balticus, and Leymus triticoides associations may be replaced by the Phalaris arundinacea association. The Carex aquatilis and Sparganium eurycarpum associations were undersampled and provide only an estimate of composition. Most associations are consistent with what has been included in the National Vegetation Classification. Close

• 5. [Article] Influence of Wetland Landscape Structure on Duck Nest Success at Malheur National Wildlife Refuge, Oregon

  The decline of waterfowl populations and their requisite wetland habitats remains a concern. Because migratory bird refuges are often artificial landscapes of actively managed wetlands, and wildlife populations ...

  Citation × Citation Citation Abstract Copy Title: Influence of Wetland Landscape Structure on Duck Nest Success at Malheur National Wildlife Refuge, Oregon Author: Craver, Daniel Robert Year: 2010 The decline of waterfowl populations and their requisite wetland habitats remains a concern. Because migratory bird refuges are often artificial landscapes of actively managed wetlands, and wildlife populations experience their greatest change during the breeding season, refuges should be designed to maximize breeding habitat. While past nest success studies have focused on at-nest variables, new approaches are needed to evaluate the effect of composition and configuration of plant communities at the landscape scale. This study aims to quantify landscape patterns within individual refuge management units to determine influence upon historical nesting success averages of ducks at Malheur National Wildlife Refuge in southeastern Oregon. The Mayfield estimate of nest success for 8 duck species yields a survival rate of 25% for the years 1987-1998 with a range of 0-74% across 48 management units. FRAGSTATS was used to calculate a suite of 9 landscape pattern metrics per unit at the class-level for each of 3 wetland habitat classes, and at the landscape-level using all habitat types. These 36 variables were tested for association with survival rate using Pearson R correlation. Results suggest fragmentation of dry meadow habitat; patch size, complexity, and extensiveness of wet meadow habitat; and diversity of habitats across the landscape positively influence duck nest success at the scale of the individual management unit. Title: Influence of Wetland Landscape Structure on Duck Nest Success at Malheur National Wildlife Refuge, Oregon Author: Craver, Daniel Robert Year: 2010 The decline of waterfowl populations and their requisite wetland habitats remains a concern. Because migratory bird refuges are often artificial landscapes of actively managed wetlands, and wildlife populations experience their greatest change during the breeding season, refuges should be designed to maximize breeding habitat. While past nest success studies have focused on at-nest variables, new approaches are needed to evaluate the effect of composition and configuration of plant communities at the landscape scale. This study aims to quantify landscape patterns within individual refuge management units to determine influence upon historical nesting success averages of ducks at Malheur National Wildlife Refuge in southeastern Oregon. The Mayfield estimate of nest success for 8 duck species yields a survival rate of 25% for the years 1987-1998 with a range of 0-74% across 48 management units. FRAGSTATS was used to calculate a suite of 9 landscape pattern metrics per unit at the class-level for each of 3 wetland habitat classes, and at the landscape-level using all habitat types. These 36 variables were tested for association with survival rate using Pearson R correlation. Results suggest fragmentation of dry meadow habitat; patch size, complexity, and extensiveness of wet meadow habitat; and diversity of habitats across the landscape positively influence duck nest success at the scale of the individual management unit. Close

• 6. [Article] A Survey for Leucocytozoon Simondi Mathis and Leger (1910) in Canada Geese of the Malheur Wildlife Refuge

  Blood smears of blood collected in 1971 and 1972 from 291 Canada geese at Malheur National Wildlife Refuge were examined to determine the incidence of the avian malarial parasite, Leucocytozoon simondi. ...

  Citation × Citation Citation Abstract Copy Title: A Survey for Leucocytozoon Simondi Mathis and Leger (1910) in Canada Geese of the Malheur Wildlife Refuge Author: Snively, John Val Year: 1973 Blood smears of blood collected in 1971 and 1972 from 291 Canada geese at Malheur National Wildlife Refuge were examined to determine the incidence of the avian malarial parasite, Leucocytozoon simondi. No parasites were found. Absence of blood parasites suggests that birds using migration routes to the north and northeast of Malheur are free of the disease. The Leucocytozoon infections known in California waterfowl may be attributable to sources within the flyway east or northeast of California, to infected birds crossing over from other flyways, or abnormal conditions in the molting grounds of sub-adult and non-nesting birds where major flyways converge in northcentral Canada. Title: A Survey for Leucocytozoon Simondi Mathis and Leger (1910) in Canada Geese of the Malheur Wildlife Refuge Author: Snively, John Val Year: 1973 Blood smears of blood collected in 1971 and 1972 from 291 Canada geese at Malheur National Wildlife Refuge were examined to determine the incidence of the avian malarial parasite, Leucocytozoon simondi. No parasites were found. Absence of blood parasites suggests that birds using migration routes to the north and northeast of Malheur are free of the disease. The Leucocytozoon infections known in California waterfowl may be attributable to sources within the flyway east or northeast of California, to infected birds crossing over from other flyways, or abnormal conditions in the molting grounds of sub-adult and non-nesting birds where major flyways converge in northcentral Canada. Close

• 7. [Article] Aspects of the ecology of the common raven in Harney Basin, Oregon

  Common Ravens (Corvus corax L.) have been implicated as significant predators on the eggs of waterfowl and shorebirds on Malheur National Wildlife Refuge. Malheur Refuge, located in southeastern Oregon ...

  Citation × Citation Citation Abstract Copy Title: Aspects of the ecology of the common raven in Harney Basin, Oregon Author: Stiehl, Richard B. Year: 1978 Common Ravens (Corvus corax L.) have been implicated as significant predators on the eggs of waterfowl and shorebirds on Malheur National Wildlife Refuge. Malheur Refuge, located in southeastern Oregon on the northern edge of the Great Basin, is one of the largest waterfowl refuges in the United States and is an important breeding area for waterfowl. In order to provide fundamental information on which a sound raven management plan could be based, research was conducted from 1975-1977 on aspects of population density, brood phenology, nesting success, seasonal use of the study area, roosting behavior, and food habits of ravens on and near the refuge. Nesting density was determined to be one pair per approximately 25 km2 . Most nests occurred in rimrocks, but trees and abandoned human structures were also used. Investigation of 87 nests revealed that the incubation period was 21 + 1 days. Incubation began with the laying of the first egg; hatching was asynchronous. Nesting period was 41 + 3 days. An original method of age-classing ravens is described. A total of 266 ravens was marked with patagial tags. Observation of marked individuals as far as 480 km from the study area suggests considerable mobility in the population. Population numbers vary seasonally, peaking in the winter. The Harney Basin is the location of an exceptionally large winter roost for ravens. Analysis of food remains, collected from 34 nests, indicates that ravens have varied diets and that there are significant differences in the diets of ravens nesting in different habitats. A correlation exists between the proportion of the diet that is avian material and the proximity of the raven nest to waterfowl production areas. Based on these findings, suggestions are offered for a management plan for Common Ravens on Malheur National Wildlife Refuge. Title: Aspects of the ecology of the common raven in Harney Basin, Oregon Author: Stiehl, Richard B. Year: 1978 Common Ravens (Corvus corax L.) have been implicated as significant predators on the eggs of waterfowl and shorebirds on Malheur National Wildlife Refuge. Malheur Refuge, located in southeastern Oregon on the northern edge of the Great Basin, is one of the largest waterfowl refuges in the United States and is an important breeding area for waterfowl. In order to provide fundamental information on which a sound raven management plan could be based, research was conducted from 1975-1977 on aspects of population density, brood phenology, nesting success, seasonal use of the study area, roosting behavior, and food habits of ravens on and near the refuge. Nesting density was determined to be one pair per approximately 25 km2 . Most nests occurred in rimrocks, but trees and abandoned human structures were also used. Investigation of 87 nests revealed that the incubation period was 21 + 1 days. Incubation began with the laying of the first egg; hatching was asynchronous. Nesting period was 41 + 3 days. An original method of age-classing ravens is described. A total of 266 ravens was marked with patagial tags. Observation of marked individuals as far as 480 km from the study area suggests considerable mobility in the population. Population numbers vary seasonally, peaking in the winter. The Harney Basin is the location of an exceptionally large winter roost for ravens. Analysis of food remains, collected from 34 nests, indicates that ravens have varied diets and that there are significant differences in the diets of ravens nesting in different habitats. A correlation exists between the proportion of the diet that is avian material and the proximity of the raven nest to waterfowl production areas. Based on these findings, suggestions are offered for a management plan for Common Ravens on Malheur National Wildlife Refuge. Close

• 8. [Article] Behavioral Observation and Paternal Investment of Eastern Kingbirds at Malheur National Wildlife Refuge

  We have been studying the population of Eastern Kingbirds breeding in riparian habitats in Malheur National Wildlife Refuge (eastern Oregon) since 2002. These efforts have created an ideal research environment ...

  Citation × Citation Citation Abstract Copy Title: Behavioral Observation and Paternal Investment of Eastern Kingbirds at Malheur National Wildlife Refuge Author: Chutter, Christopher Michael Year: 2014 We have been studying the population of Eastern Kingbirds breeding in riparian habitats in Malheur National Wildlife Refuge (eastern Oregon) since 2002. These efforts have created an ideal research environment wherein most adults in the population have been color banded and DNA sampled and, as part of other research projects, nearly all broods since 2003 have been paternity tested. I decided to use behavioral video recordings of parental nest behavior undertaken between 2003 and 2010 for two unrelated projects. First, I tested the effectiveness of video sampling nesting behavior (see below and chapter 2). Second, I tested whether male kingbirds were able to affect their level of paternal investment in accordance with their level or realized paternity (see below and chapter 3). Chapter 2 was split into three distinct questions: 1) are parental nesting behaviors repeatable, 2) is a one hour sample sufficient to capture variability in these behaviors, and 3) is the first hour of recording sufficient to capture variability in these behaviors. I found overwhelming evidence that the behaviors I measured were repeatable. This is truly important, for if repeatability was disproven, it would call into question the use of sampling throughout the field of animal behavior. I similarly found strong evidence that a one hour sample was sufficient to capture variability in parental behaviors. From this I was able to suggest that further sampling effort would be better spent increasing sample size rather than observation length. Testing whether the first hour of recording was sufficient to capture variability in parental behavior found more muddled results. While there generally was correlation between behavioral values in the first hour and those over a longer observation period, most behaviors were found to have significantly lower values in the first hour. I tested whether this was the result of a lingering observer effect or a natural effect of time of day and concluded that an observer effect was the more likely explanation. In chapter 3, I ran one of the more in depth and complicated tests for a relationship between paternal investment and realized paternity that I was able to find in the literature. I used the standard male feeding rate as a measure of male investment as well as a far more nuanced measure derived from the first Eigenvector of an analysis of six different paternal behaviors. These were both tested using Akaike's Information Criterion against a number of variables likely to affect parental behavior, including realized paternity. Ultimately, I found no evidence that kingbirds were able to affect paternal investment in response to lost paternity, and conclude that they likely had no means to assess realized paternity within the nest of their social mate. However, from the other parameters tested I was able to find that males increased their investment as brood size increased and as the female spent more time attending, but not feeding, the nestlings. Also, male investment decreased as territory density increased and flight feather length increased. Theory surrounding the variability of extrapair paternity in birds can be used to account for these results. In a denser population males are predicted to reduce paternal effort for one of two reasons: increased likelihood of paternity loss in his own nest and an increased investment in pursuing extrapair copulations of his own. Longer flight feathers are a sign of age and quality in kingbirds, so males with this trait are expected to spend more time pursuing extrapair copulations as they are more likely to be successful in this endeavor. Title: Behavioral Observation and Paternal Investment of Eastern Kingbirds at Malheur National Wildlife Refuge Author: Chutter, Christopher Michael Year: 2014 We have been studying the population of Eastern Kingbirds breeding in riparian habitats in Malheur National Wildlife Refuge (eastern Oregon) since 2002. These efforts have created an ideal research environment wherein most adults in the population have been color banded and DNA sampled and, as part of other research projects, nearly all broods since 2003 have been paternity tested. I decided to use behavioral video recordings of parental nest behavior undertaken between 2003 and 2010 for two unrelated projects. First, I tested the effectiveness of video sampling nesting behavior (see below and chapter 2). Second, I tested whether male kingbirds were able to affect their level of paternal investment in accordance with their level or realized paternity (see below and chapter 3). Chapter 2 was split into three distinct questions: 1) are parental nesting behaviors repeatable, 2) is a one hour sample sufficient to capture variability in these behaviors, and 3) is the first hour of recording sufficient to capture variability in these behaviors. I found overwhelming evidence that the behaviors I measured were repeatable. This is truly important, for if repeatability was disproven, it would call into question the use of sampling throughout the field of animal behavior. I similarly found strong evidence that a one hour sample was sufficient to capture variability in parental behaviors. From this I was able to suggest that further sampling effort would be better spent increasing sample size rather than observation length. Testing whether the first hour of recording was sufficient to capture variability in parental behavior found more muddled results. While there generally was correlation between behavioral values in the first hour and those over a longer observation period, most behaviors were found to have significantly lower values in the first hour. I tested whether this was the result of a lingering observer effect or a natural effect of time of day and concluded that an observer effect was the more likely explanation. In chapter 3, I ran one of the more in depth and complicated tests for a relationship between paternal investment and realized paternity that I was able to find in the literature. I used the standard male feeding rate as a measure of male investment as well as a far more nuanced measure derived from the first Eigenvector of an analysis of six different paternal behaviors. These were both tested using Akaike's Information Criterion against a number of variables likely to affect parental behavior, including realized paternity. Ultimately, I found no evidence that kingbirds were able to affect paternal investment in response to lost paternity, and conclude that they likely had no means to assess realized paternity within the nest of their social mate. However, from the other parameters tested I was able to find that males increased their investment as brood size increased and as the female spent more time attending, but not feeding, the nestlings. Also, male investment decreased as territory density increased and flight feather length increased. Theory surrounding the variability of extrapair paternity in birds can be used to account for these results. In a denser population males are predicted to reduce paternal effort for one of two reasons: increased likelihood of paternity loss in his own nest and an increased investment in pursuing extrapair copulations of his own. Longer flight feathers are a sign of age and quality in kingbirds, so males with this trait are expected to spend more time pursuing extrapair copulations as they are more likely to be successful in this endeavor. Close

• 9. [Article] Archaeological Investigations at 45CL1 Cathlapotle (1991-1996) , Ridgefield National Wildlife Refuge Clark County, Washington A Preliminary Report

  Archaeological investigations at site 45CL1, Clark County, Washington, demonstrate that the locality is a very large (c 1.5ha), deeply stratified (2-4m) town site with an occupation spanning at least 1000 ...

  Citation × Citation Citation Abstract Copy Title: Archaeological Investigations at 45CL1 Cathlapotle (1991-1996) , Ridgefield National Wildlife Refuge Clark County, Washington A Preliminary Report Author: Ames, Kenneth M., Smith, Cameron M., Cornett, William L., Sobel, Elizabeth A., Hamilton, Stephen C., Wolf, John, Raetz, Doria, U.S. Fish and Wildlife Service. Region 1, Portland State University. Department of Anthropology Year: 1999 Archaeological investigations at site 45CL1, Clark County, Washington, demonstrate that the locality is a very large (c 1.5ha), deeply stratified (2-4m) town site with an occupation spanning at least 1000 years (c. AD 1000 to 1840). Six large, complex depressions have been mapped. Test excavations show that these depressions represent the semisubterranean portions of residential structures, probably large plankhouses of the type common on the Lower Columbia River and the Northwest Coast in aboriginal times. The depressions may represent as many as 11 such dwellings. A seventh depression is deeply buried beneath midden deposits. The cultural deposits contain very high densities of artifacts, ecofacts (including both faunal and floral remains), debris and features. The site is near the Columbia River on a very active flood plain, resulting in site stratigraphy produced by a combination of active cultural and alluvial depositional processes. Site 45CL1, given its location and size, is the best candidate to be the site of Cathlapotle, a Middle Chinookan town visited by Lewis and Clark in 1806, as well as by other early Europeans in the area. The site is extraordinarily well preserved, having undergone only minor alterations since its abandonment, probably in the third or fourth decade of the 19th century AD. Title: Archaeological Investigations at 45CL1 Cathlapotle (1991-1996) , Ridgefield National Wildlife Refuge Clark County, Washington A Preliminary Report Author: Ames, Kenneth M., Smith, Cameron M., Cornett, William L., Sobel, Elizabeth A., Hamilton, Stephen C., Wolf, John, Raetz, Doria, U.S. Fish and Wildlife Service. Region 1, Portland State University. Department of Anthropology Year: 1999 Archaeological investigations at site 45CL1, Clark County, Washington, demonstrate that the locality is a very large (c 1.5ha), deeply stratified (2-4m) town site with an occupation spanning at least 1000 years (c. AD 1000 to 1840). Six large, complex depressions have been mapped. Test excavations show that these depressions represent the semisubterranean portions of residential structures, probably large plankhouses of the type common on the Lower Columbia River and the Northwest Coast in aboriginal times. The depressions may represent as many as 11 such dwellings. A seventh depression is deeply buried beneath midden deposits. The cultural deposits contain very high densities of artifacts, ecofacts (including both faunal and floral remains), debris and features. The site is near the Columbia River on a very active flood plain, resulting in site stratigraphy produced by a combination of active cultural and alluvial depositional processes. Site 45CL1, given its location and size, is the best candidate to be the site of Cathlapotle, a Middle Chinookan town visited by Lewis and Clark in 1806, as well as by other early Europeans in the area. The site is extraordinarily well preserved, having undergone only minor alterations since its abandonment, probably in the third or fourth decade of the 19th century AD. Close

• 10. [Article] Nest Reuse by Eastern Kingbirds: Adaptive Behavior or Ecological Constraint?

  The reuse of old nests by open-cup nesting passerines is a seemingly rare but potentially adaptive behavior if, as a consequence, females begin to breed earlier, lay larger clutches, or fledge more young. ...

  Citation × Citation Citation Abstract Copy Title: Nest Reuse by Eastern Kingbirds: Adaptive Behavior or Ecological Constraint? Author: Redmond, Lucas J., Murphy, Michael T., Dolan, Amy C. Year: 2007 The reuse of old nests by open-cup nesting passerines is a seemingly rare but potentially adaptive behavior if, as a consequence, females begin to breed earlier, lay larger clutches, or fledge more young. We report an unusually high rate of nest reuse (~10% of 341 nests) for Eastern Kingbirds (Tyrannus tyrannus) breeding at Malheur National Wildlife Refuge, Oregon. We found no difference in availability of nesting habitat or food abundance in territories in which nests were and were not reused. We also found no support for the hypotheses that kingbirds benefited from nest reuse by breeding earlier, laying larger clutches, or fledging more young, and, contrary to expectations, females that reused nests laid significantly smaller eggs than females who built their own nests. Nest reuse was independent of age: a roughly equal number of females for which we had multiple years of data both reused nests and built new nests, but at different points in their lives. Competition for nest sites seems high at Malheur National Wildlife Refuge because many open-cup nesting species utilize similar nest sites in the limited zone of riparian vegetation. A shortage of high-quality nest sites, coupled with interspecific competition, may underlie the high frequency of nest reuse in this kingbird population. Title: Nest Reuse by Eastern Kingbirds: Adaptive Behavior or Ecological Constraint? Author: Redmond, Lucas J., Murphy, Michael T., Dolan, Amy C. Year: 2007 The reuse of old nests by open-cup nesting passerines is a seemingly rare but potentially adaptive behavior if, as a consequence, females begin to breed earlier, lay larger clutches, or fledge more young. We report an unusually high rate of nest reuse (~10% of 341 nests) for Eastern Kingbirds (Tyrannus tyrannus) breeding at Malheur National Wildlife Refuge, Oregon. We found no difference in availability of nesting habitat or food abundance in territories in which nests were and were not reused. We also found no support for the hypotheses that kingbirds benefited from nest reuse by breeding earlier, laying larger clutches, or fledging more young, and, contrary to expectations, females that reused nests laid significantly smaller eggs than females who built their own nests. Nest reuse was independent of age: a roughly equal number of females for which we had multiple years of data both reused nests and built new nests, but at different points in their lives. Competition for nest sites seems high at Malheur National Wildlife Refuge because many open-cup nesting species utilize similar nest sites in the limited zone of riparian vegetation. A shortage of high-quality nest sites, coupled with interspecific competition, may underlie the high frequency of nest reuse in this kingbird population. Close

• 11. [Article] Use of Water Indices Derived from Landsat OLI Imagery and GIS to Estimate the Hydrologic Connectivity of Wetlands in the Tualatin River National Wildlife Refuge

  This study compared two remote sensing water indices: the Normalized Difference Water Index (NDWI) and the Modified NDWI (MNDWI). Both indices were calculated using publically-available data from the Landsat ...

  Citation × Citation Citation Abstract Copy Title: Use of Water Indices Derived from Landsat OLI Imagery and GIS to Estimate the Hydrologic Connectivity of Wetlands in the Tualatin River National Wildlife Refuge Author: Blackmore, Debra Sue Year: 2016 This study compared two remote sensing water indices: the Normalized Difference Water Index (NDWI) and the Modified NDWI (MNDWI). Both indices were calculated using publically-available data from the Landsat 8 Operational Land Imager (OLI). The research goal was to determine whether the indices are effective in locating open water and measuring surface soil moisture. To demonstrate the application of water indices, analysis was conducted for freshwater wetlands in the Tualatin River Basin in northwestern Oregon to estimate hydrologic connectivity and hydrological permanence between these wetlands and nearby water bodies. Remote sensing techniques have been used to study wetlands in recent decades; however, scientific studies have rarely addressed hydrologic connectivity and hydrologic permanence, in spite of the documented importance of these properties. Research steps were designed to be straightforward for easy repeatability: 1) locate sample sites, 2) predict wetness with water indices, 3) estimate wetness with soil samples from the field, 4) validate the index predictions against the soil samples from the field, and 5) in the demonstration step, estimate hydrologic connectivity and hydrological permanence. Results indicate that both indices predicted the presence of large, open water features with clarity; that dry conditions were predicted by MNDWI with more subtle differentiation; and that NDWI results seem more sensitive to sites with vegetation. Use of this low-cost method to discover patterns of surface moisture in the landscape could directly improve the ability to manage wetland environments. Title: Use of Water Indices Derived from Landsat OLI Imagery and GIS to Estimate the Hydrologic Connectivity of Wetlands in the Tualatin River National Wildlife Refuge Author: Blackmore, Debra Sue Year: 2016 This study compared two remote sensing water indices: the Normalized Difference Water Index (NDWI) and the Modified NDWI (MNDWI). Both indices were calculated using publically-available data from the Landsat 8 Operational Land Imager (OLI). The research goal was to determine whether the indices are effective in locating open water and measuring surface soil moisture. To demonstrate the application of water indices, analysis was conducted for freshwater wetlands in the Tualatin River Basin in northwestern Oregon to estimate hydrologic connectivity and hydrological permanence between these wetlands and nearby water bodies. Remote sensing techniques have been used to study wetlands in recent decades; however, scientific studies have rarely addressed hydrologic connectivity and hydrologic permanence, in spite of the documented importance of these properties. Research steps were designed to be straightforward for easy repeatability: 1) locate sample sites, 2) predict wetness with water indices, 3) estimate wetness with soil samples from the field, 4) validate the index predictions against the soil samples from the field, and 5) in the demonstration step, estimate hydrologic connectivity and hydrological permanence. Results indicate that both indices predicted the presence of large, open water features with clarity; that dry conditions were predicted by MNDWI with more subtle differentiation; and that NDWI results seem more sensitive to sites with vegetation. Use of this low-cost method to discover patterns of surface moisture in the landscape could directly improve the ability to manage wetland environments. Close

• 12. [Article] The Biology of Eastern Kingbirds at Malheur National Wildlife Refuge: Survival, Reproduction, and Testosterone Secretion

  This dissertation presents the results of a study that I undertook to better understand the breeding biology of Eastern Kingbirds (hereafter, kingbirds) at Malheur National Wildlife Refuge in southeastern ...

  Citation × Citation Citation Abstract Copy Title: The Biology of Eastern Kingbirds at Malheur National Wildlife Refuge: Survival, Reproduction, and Testosterone Secretion Author: Redmond, Lucas J. Year: 2015 This dissertation presents the results of a study that I undertook to better understand the breeding biology of Eastern Kingbirds (hereafter, kingbirds) at Malheur National Wildlife Refuge in southeastern Oregon from 2003 to 2009. Kingbirds are long-distance migratory songbirds that breed across much of North America. This species is socially monogamous but, via frequent extra-pair copulations, is genetically polygamous. Kingbirds exhibit relatively high breeding site fidelity, often returning to the same tree to nest in subsequent years. Both members of a pair provide parental care, but there are often specific duties performed by both male and female kingbirds. For example, males typically perform much of the vigilant nest defense that this species is well known for and contribute, to an extent, to the feeding of nestlings. Females, on the other hand, are entirely responsible for incubating and brooding. Beginning in 2002, most adults within the population of kingbirds at Malheur had been banded with a unique combination of three colored plastic leg bands and an aluminum USFWS band. Also, as many nests were located in each year as possible, and any young that survived to fledging age were banded with a unique combination of leg bands. Considerable effort was then spent each year to locate as many banded individuals as possible, which allowed me to document adult and juvenile survival. Annual survival rates of adult male and female kingbirds in the population at Malheur did not differ and were relatively high at approximately 0.65. Juvenile survival rate was approximately 0.29, indicating that slightly less than one-third of nestlings survive the interval between fledging and their first potential breeding season. Resighting probability was high for both sexes, although higher for adult males (0.94) than for adult female (0.84). The latter finding is consistent with the higher site fidelity of males than of females. Resighting probabilities for juveniles were much lower at 0.68 and 0.40 for males and females, respectively. Again, this was expected because natal site fidelity is typically much lower than breeding site fidelity. Compared to most other reports, resighting probability and return rates of juvenile kingbirds was high, presumably because the riparian habitats where kingbirds breed at Malheur function as an ecological island surrounded by, for kingbirds, unsuitable high desert habitat. Thus, unless they disperse very long distances, the only option for juvenile kingbirds is to begin nesting on the refuge. The collection of blood samples from birds has become an increasingly common practice in ornithology. The data that can be gained from these samples allow a number of interesting questions to be asked such as understanding the genetic mating system of a species, patterns of hormone secretion, and discerning migratory pathways via stable isotope analysis. The volume of blood collected is usually small and was assumed to cause no long-term negative effects on sampled individuals. However, few studies have rigorously examined the effect of blood sampling on survival. I used a multistate mark-recapture analysis to assess the effect blood sampling on annual survival of kingbirds by combining the annual survival data described above with whether or not individuals were subjected to blood sample collection. The results of this analysis indicated that blood sampling had no effect on annual survival rates of kingbirds. Whether or not this is the case for other species remains to be seen. However, my results support the assumption that when done correctly, blood sampling has little to no long-term negative effects on birds. Comparative analyses of many bird species show that testosterone secretion exhibits fairly predictable patterns among breeding birds of different mating systems. Monogamous species reach a peak during mate attraction and the period of female fertility which is then followed by a sharp decline when young are in the nest. By contrast, males of polygynous species tend to maintain higher levels of testosterone throughout the breeding season to, presumably, maximize opportunities for acquiring additional mates. Kingbirds are socially monogamous but cryptically polygamous. However, because of high rates of extra-pair paternity, variation in reproductive success among males is much higher than what is expected for a monogamous species, and, instead, is similar to what has been reported for polygynous species. Therefore from 2005 to 2009 I measured testosterone concentrations from the blood samples collected from male kingbirds to determine the breeding season profile of testosterone secretion in kingbirds and to understand the factors that influence testosterone variation among individuals. Contrary to expected for a monogamous species, the testosterone profile of kingbirds did not exhibit the brief peak in testosterone followed by a precipitous decline. Testosterone peaked early in the season, but declined very slowly as the nest cycle progressed. I attributed this gradual decline in testosterone to the cryptically polygynous nature of the kingbird extra-pair mating system. I also found substantial variation in testosterone concentration among male kingbirds and was able to identify several factors contributing to this variation. As expected, testosterone declined as the breeding season progressed (independent of stage in nest cycle) and nest density increased, while increases in testosterone were correlated with the number of fertile females within the population. This suggests that male kingbirds were capable of modulating testosterone concentrations to both cope with an increase in conspecific density, but also to ready themselves for times when extra-pair copulations were more likely. Title: The Biology of Eastern Kingbirds at Malheur National Wildlife Refuge: Survival, Reproduction, and Testosterone Secretion Author: Redmond, Lucas J. Year: 2015 This dissertation presents the results of a study that I undertook to better understand the breeding biology of Eastern Kingbirds (hereafter, kingbirds) at Malheur National Wildlife Refuge in southeastern Oregon from 2003 to 2009. Kingbirds are long-distance migratory songbirds that breed across much of North America. This species is socially monogamous but, via frequent extra-pair copulations, is genetically polygamous. Kingbirds exhibit relatively high breeding site fidelity, often returning to the same tree to nest in subsequent years. Both members of a pair provide parental care, but there are often specific duties performed by both male and female kingbirds. For example, males typically perform much of the vigilant nest defense that this species is well known for and contribute, to an extent, to the feeding of nestlings. Females, on the other hand, are entirely responsible for incubating and brooding. Beginning in 2002, most adults within the population of kingbirds at Malheur had been banded with a unique combination of three colored plastic leg bands and an aluminum USFWS band. Also, as many nests were located in each year as possible, and any young that survived to fledging age were banded with a unique combination of leg bands. Considerable effort was then spent each year to locate as many banded individuals as possible, which allowed me to document adult and juvenile survival. Annual survival rates of adult male and female kingbirds in the population at Malheur did not differ and were relatively high at approximately 0.65. Juvenile survival rate was approximately 0.29, indicating that slightly less than one-third of nestlings survive the interval between fledging and their first potential breeding season. Resighting probability was high for both sexes, although higher for adult males (0.94) than for adult female (0.84). The latter finding is consistent with the higher site fidelity of males than of females. Resighting probabilities for juveniles were much lower at 0.68 and 0.40 for males and females, respectively. Again, this was expected because natal site fidelity is typically much lower than breeding site fidelity. Compared to most other reports, resighting probability and return rates of juvenile kingbirds was high, presumably because the riparian habitats where kingbirds breed at Malheur function as an ecological island surrounded by, for kingbirds, unsuitable high desert habitat. Thus, unless they disperse very long distances, the only option for juvenile kingbirds is to begin nesting on the refuge. The collection of blood samples from birds has become an increasingly common practice in ornithology. The data that can be gained from these samples allow a number of interesting questions to be asked such as understanding the genetic mating system of a species, patterns of hormone secretion, and discerning migratory pathways via stable isotope analysis. The volume of blood collected is usually small and was assumed to cause no long-term negative effects on sampled individuals. However, few studies have rigorously examined the effect of blood sampling on survival. I used a multistate mark-recapture analysis to assess the effect blood sampling on annual survival of kingbirds by combining the annual survival data described above with whether or not individuals were subjected to blood sample collection. The results of this analysis indicated that blood sampling had no effect on annual survival rates of kingbirds. Whether or not this is the case for other species remains to be seen. However, my results support the assumption that when done correctly, blood sampling has little to no long-term negative effects on birds. Comparative analyses of many bird species show that testosterone secretion exhibits fairly predictable patterns among breeding birds of different mating systems. Monogamous species reach a peak during mate attraction and the period of female fertility which is then followed by a sharp decline when young are in the nest. By contrast, males of polygynous species tend to maintain higher levels of testosterone throughout the breeding season to, presumably, maximize opportunities for acquiring additional mates. Kingbirds are socially monogamous but cryptically polygamous. However, because of high rates of extra-pair paternity, variation in reproductive success among males is much higher than what is expected for a monogamous species, and, instead, is similar to what has been reported for polygynous species. Therefore from 2005 to 2009 I measured testosterone concentrations from the blood samples collected from male kingbirds to determine the breeding season profile of testosterone secretion in kingbirds and to understand the factors that influence testosterone variation among individuals. Contrary to expected for a monogamous species, the testosterone profile of kingbirds did not exhibit the brief peak in testosterone followed by a precipitous decline. Testosterone peaked early in the season, but declined very slowly as the nest cycle progressed. I attributed this gradual decline in testosterone to the cryptically polygynous nature of the kingbird extra-pair mating system. I also found substantial variation in testosterone concentration among male kingbirds and was able to identify several factors contributing to this variation. As expected, testosterone declined as the breeding season progressed (independent of stage in nest cycle) and nest density increased, while increases in testosterone were correlated with the number of fertile females within the population. This suggests that male kingbirds were capable of modulating testosterone concentrations to both cope with an increase in conspecific density, but also to ready themselves for times when extra-pair copulations were more likely. Close

• 13. [Article] Finding and Dating Cathlapotle

  The people of the Cathlapotle town played a significant role in the fur trade era history of the Lower Columbia River, including Lewis and Clark’s visit on March 29th, 1806. Archaeologists and others have ...

  Citation × Citation Citation Abstract Copy Title: Finding and Dating Cathlapotle Author: Ames, Kenneth M., Sobel, Elizabeth A. Year: 2009 The people of the Cathlapotle town played a significant role in the fur trade era history of the Lower Columbia River, including Lewis and Clark’s visit on March 29th, 1806. Archaeologists and others have sought the town’s location for years. Long-term research has established that archaeological site 45CL1 on the US Fish and Wildlife Refuge near Ridgefield, Washington is Cathlapotle. This determination is based on the close match between site details with various ethnohistoric accounts of Cathlapotle. The site was occupied by ca. AD 1450 and probably moved there from another nearby location. It was abandoned sometime in the 1830s or 1840s. This chronology is based on 54 radiocarbon dates, historic trade goods including glass beads and ceramics, and documentary accounts. Title: Finding and Dating Cathlapotle Author: Ames, Kenneth M., Sobel, Elizabeth A. Year: 2009 The people of the Cathlapotle town played a significant role in the fur trade era history of the Lower Columbia River, including Lewis and Clark’s visit on March 29th, 1806. Archaeologists and others have sought the town’s location for years. Long-term research has established that archaeological site 45CL1 on the US Fish and Wildlife Refuge near Ridgefield, Washington is Cathlapotle. This determination is based on the close match between site details with various ethnohistoric accounts of Cathlapotle. The site was occupied by ca. AD 1450 and probably moved there from another nearby location. It was abandoned sometime in the 1830s or 1840s. This chronology is based on 54 radiocarbon dates, historic trade goods including glass beads and ceramics, and documentary accounts. Close

• 14. [Article] Functional analysis of Great Basin projectile points

  This paper presents a refined methodology for distinguishing the stone points of arrows from the stone points of spear thrower darts in archaeological assemblages from the Great Basin. The methodology ...

  Citation × Citation Citation Abstract Copy Title: Functional analysis of Great Basin projectile points Author: Wiggin, Roger Year: 1979 This paper presents a refined methodology for distinguishing the stone points of arrows from the stone points of spear thrower darts in archaeological assemblages from the Great Basin. The methodology was developed from a sample of 111 complete stone points collected during the archaeological reconnaissance and testing of the Malheur National Wildlife Refuge, Harney County, Oregon, 1971-1974. The points were measured for five metric variables and frequency histograms of each variable were generated by computer. The effectiveness of each variable as an index of functional class (i.e., arrow point vs. dart point) was judged on the basis of the apparent bimodality of its curve and the t-test. The variables of weight and neck width showed significant bimodality, confirming the work of previous researchers. A new variable, stem thickness, also showed significant bimodality. All three can be considered useful indices of functional class. They also appear to be functionally independent. An attempt was made at developing a multivariable index of point function, using a multivariable clustering analysis, the K-means test. Results of the K-means test were inconclusive but showed promise for further development. Title: Functional analysis of Great Basin projectile points Author: Wiggin, Roger Year: 1979 This paper presents a refined methodology for distinguishing the stone points of arrows from the stone points of spear thrower darts in archaeological assemblages from the Great Basin. The methodology was developed from a sample of 111 complete stone points collected during the archaeological reconnaissance and testing of the Malheur National Wildlife Refuge, Harney County, Oregon, 1971-1974. The points were measured for five metric variables and frequency histograms of each variable were generated by computer. The effectiveness of each variable as an index of functional class (i.e., arrow point vs. dart point) was judged on the basis of the apparent bimodality of its curve and the t-test. The variables of weight and neck width showed significant bimodality, confirming the work of previous researchers. A new variable, stem thickness, also showed significant bimodality. All three can be considered useful indices of functional class. They also appear to be functionally independent. An attempt was made at developing a multivariable index of point function, using a multivariable clustering analysis, the K-means test. Results of the K-means test were inconclusive but showed promise for further development. Close

• 15. [Article] Avian Affairs: A Study of the Occurrence and Benefits of Extra-pair Breeding Behavior in Eastern Kingbirds (Tyrannus Tyrannus)

  Over three consecutive breeding seasons I examined the breeding system of Eastern Kingbirds in Malheur National Wildlife Refuge in SE, Oregon. To assess genetic paternity, I optimized seven microsatellite ...

  Citation × Citation Citation Abstract Copy Title: Avian Affairs: A Study of the Occurrence and Benefits of Extra-pair Breeding Behavior in Eastern Kingbirds (Tyrannus Tyrannus) Author: Dolan, Amy Catherine Year: 2006 Over three consecutive breeding seasons I examined the breeding system of Eastern Kingbirds in Malheur National Wildlife Refuge in SE, Oregon. To assess genetic paternity, I optimized seven microsatellite markers and determined that extra pair paternity was common. Extra-pair young were present in approximately 60% of nests, representing 47% of young. I examined implications of extra-pair paternity from three perspectives. First, I examined the impact of extra-pair paternity on the opportunity for sexual selection within a socially monogamous system and identified correlates of male reproductive success. Some males were successful at the expense of others, and 24% of males did not sire any young. Thus, variance in male reproductive success was more than nine times greater than female variance, indicating that sexual selection operates strongly in this system. Extra-pair success was positively related to tarsus length and inversely dependent upon dawn song start time. Within-pair success was positively related to nest defense intensity during the incubation period. Second, I assessed the four genetic hypotheses proposed to explain female cooperation in extra-pair copulations. Extra-pair young were not distributed randomly among broods. Cuckolded within-pair males were not more closely related to females than were extra-pair sires. Pair wise comparisons indicated males with specific song and morphological traits were chosen as sires. Brood success (proportion of brood to fledge and recruit) was unrelated to thenumber of males who sired young in the brood. Extra-pair young were larger and heavier than their within-pair half-siblings. These results unambiguously support the 'good genes' hypothesis of female extra-pair mating. Last, I examined the influence of breeding date and parental condition (mother, social father and genetic sire) on offspring sex ratio. Given that male variance in reproductive success is much greater than that of females, I predicted the characters of genetic sires would be the most important factor influencing offspring sex ratio. Contrary to my prediction, male traits did not influence offspring sex ratio. However, more males hatched early in the season and females in better condition produced more sons. I also found that male nestling survival to the next breeding season was inversely related to date. Title: Avian Affairs: A Study of the Occurrence and Benefits of Extra-pair Breeding Behavior in Eastern Kingbirds (Tyrannus Tyrannus) Author: Dolan, Amy Catherine Year: 2006 Over three consecutive breeding seasons I examined the breeding system of Eastern Kingbirds in Malheur National Wildlife Refuge in SE, Oregon. To assess genetic paternity, I optimized seven microsatellite markers and determined that extra pair paternity was common. Extra-pair young were present in approximately 60% of nests, representing 47% of young. I examined implications of extra-pair paternity from three perspectives. First, I examined the impact of extra-pair paternity on the opportunity for sexual selection within a socially monogamous system and identified correlates of male reproductive success. Some males were successful at the expense of others, and 24% of males did not sire any young. Thus, variance in male reproductive success was more than nine times greater than female variance, indicating that sexual selection operates strongly in this system. Extra-pair success was positively related to tarsus length and inversely dependent upon dawn song start time. Within-pair success was positively related to nest defense intensity during the incubation period. Second, I assessed the four genetic hypotheses proposed to explain female cooperation in extra-pair copulations. Extra-pair young were not distributed randomly among broods. Cuckolded within-pair males were not more closely related to females than were extra-pair sires. Pair wise comparisons indicated males with specific song and morphological traits were chosen as sires. Brood success (proportion of brood to fledge and recruit) was unrelated to thenumber of males who sired young in the brood. Extra-pair young were larger and heavier than their within-pair half-siblings. These results unambiguously support the 'good genes' hypothesis of female extra-pair mating. Last, I examined the influence of breeding date and parental condition (mother, social father and genetic sire) on offspring sex ratio. Given that male variance in reproductive success is much greater than that of females, I predicted the characters of genetic sires would be the most important factor influencing offspring sex ratio. Contrary to my prediction, male traits did not influence offspring sex ratio. However, more males hatched early in the season and females in better condition produced more sons. I also found that male nestling survival to the next breeding season was inversely related to date. Close

• 16. [Article] Lambda Alpha Anthropology Honors Society (LAB): Bridging The Gap Between Coursework And Career

  Many anthropology majors gain an academic familiarity with the subject through coursework, but lack an understanding of how to utilize their anthropological skills outside of academia in future careers. ...

  Citation × Citation Citation Abstract Copy Title: Lambda Alpha Anthropology Honors Society (LAB): Bridging The Gap Between Coursework And Career Author: Stein, Adriana, Berganini, Stefanie Year: 2015 Many anthropology majors gain an academic familiarity with the subject through coursework, but lack an understanding of how to utilize their anthropological skills outside of academia in future careers. To remedy this issue, the Lambda Alpha Beta Honor Society (LAB), a student-led anthropology organization at Portland State University, provides opportunities for participation in events that get students involved with our local community, such as non-profit organizations and local cultural institutions. Some of these organizations include the Oregon Museum of Science and Industry (OMSI), Clark Public Utilities, The Ridgefield Wildlife Refuge, Potluck in the Park, and Habitat for Humanity. As an unfunded student group, LAB provides opportunities for students at no cost that simultaneously boost their career while also giving back to the community. We also have spearheaded the first crowdsourcing fundraiser in the anthropology department in cooperation with the PSU Foundation to raise money to update our PSU anthropological research photography exhibit. Furthermore, we helped apply for funding to send PSU graduate students to present their research at the Society for Applied Anthropology Conference (SfAA) in Pittsburgh, PA. This conference provides a major networking opportunity for our students and assures our relationship within the applied anthropology subfield in general. Therefore, LAB provides an example of a student group that bridges the gap between studying anthropology in the classroom and practicing anthropology as a career. Many anthropology majors gain an academic familiarity with the subject through coursework, but lack an understanding of how to utilize their anthropological skills outside of academia in future careers. To remedy this issue, the Lambda Alpha Beta Honor Society (LAB), a student-led anthropology organization at Portland State University, provides opportunities for participation in events that get students involved with our local community, such as non-profit organizations and local cultural institutions. Some of these organizations include the Oregon Museum of Science and Industry (OMSI), Clark Public Utilities, The Ridgefield Wildlife Refuge, Potluck in the Park, and Habitat for Humanity. As an unfunded student group, LAB provides opportunities for students at no cost that simultaneously boost their career while also giving back to the community. We also have spearheaded the first crowdsourcing fundraiser in the anthropology department in cooperation with the PSU Foundation to raise money to update our PSU anthropological research photography exhibit. Furthermore, we helped apply for funding to send PSU graduate students to present their research at the Society for Applied Anthropology Conference (SfAA) in Pittsburgh, PA. This conference provides a major networking opportunity for our students and assures our relationship within the applied anthropology subfield in general. Therefore, LAB provides an example of a student group that bridges the gap between studying anthropology in the classroom and practicing anthropology as a career. Title: Lambda Alpha Anthropology Honors Society (LAB): Bridging The Gap Between Coursework And Career Author: Stein, Adriana, Berganini, Stefanie Year: 2015 Many anthropology majors gain an academic familiarity with the subject through coursework, but lack an understanding of how to utilize their anthropological skills outside of academia in future careers. To remedy this issue, the Lambda Alpha Beta Honor Society (LAB), a student-led anthropology organization at Portland State University, provides opportunities for participation in events that get students involved with our local community, such as non-profit organizations and local cultural institutions. Some of these organizations include the Oregon Museum of Science and Industry (OMSI), Clark Public Utilities, The Ridgefield Wildlife Refuge, Potluck in the Park, and Habitat for Humanity. As an unfunded student group, LAB provides opportunities for students at no cost that simultaneously boost their career while also giving back to the community. We also have spearheaded the first crowdsourcing fundraiser in the anthropology department in cooperation with the PSU Foundation to raise money to update our PSU anthropological research photography exhibit. Furthermore, we helped apply for funding to send PSU graduate students to present their research at the Society for Applied Anthropology Conference (SfAA) in Pittsburgh, PA. This conference provides a major networking opportunity for our students and assures our relationship within the applied anthropology subfield in general. Therefore, LAB provides an example of a student group that bridges the gap between studying anthropology in the classroom and practicing anthropology as a career. Many anthropology majors gain an academic familiarity with the subject through coursework, but lack an understanding of how to utilize their anthropological skills outside of academia in future careers. To remedy this issue, the Lambda Alpha Beta Honor Society (LAB), a student-led anthropology organization at Portland State University, provides opportunities for participation in events that get students involved with our local community, such as non-profit organizations and local cultural institutions. Some of these organizations include the Oregon Museum of Science and Industry (OMSI), Clark Public Utilities, The Ridgefield Wildlife Refuge, Potluck in the Park, and Habitat for Humanity. As an unfunded student group, LAB provides opportunities for students at no cost that simultaneously boost their career while also giving back to the community. We also have spearheaded the first crowdsourcing fundraiser in the anthropology department in cooperation with the PSU Foundation to raise money to update our PSU anthropological research photography exhibit. Furthermore, we helped apply for funding to send PSU graduate students to present their research at the Society for Applied Anthropology Conference (SfAA) in Pittsburgh, PA. This conference provides a major networking opportunity for our students and assures our relationship within the applied anthropology subfield in general. Therefore, LAB provides an example of a student group that bridges the gap between studying anthropology in the classroom and practicing anthropology as a career. Close

• 17. [Article] Experimental Analysis of Nest-Site Choice and its Relationship to Nest Success in an Open-Cup–Nesting Passerine

  Nest placement presumably reflects selection for secure sites to minimize failure. Most tests of this hypothesis, however, have failed to support it. We used artificial nests (ARTs) to experimentally evaluate ...

  Citation × Citation Citation Abstract Copy Title: Experimental Analysis of Nest-Site Choice and its Relationship to Nest Success in an Open-Cup–Nesting Passerine Author: Cancellieri, Sarah, Murphy, Michael T. Year: 2014 Nest placement presumably reflects selection for secure sites to minimize failure. Most tests of this hypothesis, however, have failed to support it. We used artificial nests (ARTs) to experimentally evaluate nest-site-choice behavior by an open-cup–nesting bird, the Eastern Kingbird (Tyrannus tyrannus). In 2010 and 2011, we placed ARTs in trees in the riparian zone at Malheur National Wildlife Refuge, Oregon, USA, to test whether (1) characteristics describing the physical location in trees of used and unused ARTs differed, (2) used ART sites more closely resembled naturally chosen sites, (3) successful natural nests (NATs) and successful ARTs were similarly located along the major axis describing nest placement, and (4) unused ARTs resembled failed NATs. Used and unused ART sites differed, but unused ART sites were more similar to NAT sites. The latter unexpected result occurred because (1) unused ARTs were located at sites between more heavily used higher and lower locations and (2) most kingbirds nesting at lower locations used ARTs instead of building their own nest. In both ARTs and NATs, differences between successful and failed nests exhibited the same pattern for most nest-site variables, and the major gradient describing nest location was the same; successful nests tended to be placed on more vertically oriented branches that were placed closer to the top of the tree. Kingbird nest placement was thus selective. However, extensive overlap in the locations in trees of failed NATs and both successful and unused ARTs suggests that other factors, such as macrohabitat characteristics or prior experience of individual birds with particular nest sites, may have influenced success and/or decisions to use or reject nests in particular locations. Thus, consideration of phenomena beyond the nest site itself may be required to fully understand the process of nest-site choice in birds. Title: Experimental Analysis of Nest-Site Choice and its Relationship to Nest Success in an Open-Cup–Nesting Passerine Author: Cancellieri, Sarah, Murphy, Michael T. Year: 2014 Nest placement presumably reflects selection for secure sites to minimize failure. Most tests of this hypothesis, however, have failed to support it. We used artificial nests (ARTs) to experimentally evaluate nest-site-choice behavior by an open-cup–nesting bird, the Eastern Kingbird (Tyrannus tyrannus). In 2010 and 2011, we placed ARTs in trees in the riparian zone at Malheur National Wildlife Refuge, Oregon, USA, to test whether (1) characteristics describing the physical location in trees of used and unused ARTs differed, (2) used ART sites more closely resembled naturally chosen sites, (3) successful natural nests (NATs) and successful ARTs were similarly located along the major axis describing nest placement, and (4) unused ARTs resembled failed NATs. Used and unused ART sites differed, but unused ART sites were more similar to NAT sites. The latter unexpected result occurred because (1) unused ARTs were located at sites between more heavily used higher and lower locations and (2) most kingbirds nesting at lower locations used ARTs instead of building their own nest. In both ARTs and NATs, differences between successful and failed nests exhibited the same pattern for most nest-site variables, and the major gradient describing nest location was the same; successful nests tended to be placed on more vertically oriented branches that were placed closer to the top of the tree. Kingbird nest placement was thus selective. However, extensive overlap in the locations in trees of failed NATs and both successful and unused ARTs suggests that other factors, such as macrohabitat characteristics or prior experience of individual birds with particular nest sites, may have influenced success and/or decisions to use or reject nests in particular locations. Thus, consideration of phenomena beyond the nest site itself may be required to fully understand the process of nest-site choice in birds. Close

• 18. [Article] Small Tidal Channels Improve Foraging Opportunities for Calidris Shorebirds

  Estuarine intertidal habitats are heterogeneous, therefore migratory shorebirds are expected to forage in microhabitats where they can maximize their energy intake. Identifying proximate factors that migratory ...

  Citation × Citation Citation Abstract Copy Title: Small Tidal Channels Improve Foraging Opportunities for Calidris Shorebirds Author: Miller, Aileen K., de Rivera, Catherine E. Year: 2014 Estuarine intertidal habitats are heterogeneous, therefore migratory shorebirds are expected to forage in microhabitats where they can maximize their energy intake. Identifying proximate factors that migratory shorebirds use to accept or reject a particular habitat patch will help land managers make conservation and restoration decisions that provide the greatest benefits to shorebird populations during migration, a period of intense energy usage. We examined whether small semipermanent tidal channels were preferentially used by foraging Western Sandpipers (Calidris mauri) and Dunlins (C. alpina) during a spring migratory stopover in Bandon Marsh, an Oregon, USA, estuary. Further, we tested alternative hypotheses about how channels might be beneficial to shorebirds by examining infauna abundance, composition, and depth alongside channels compared with that in open mudflats. The densities of Western Sandpipers and Dunlins were at least 4 times higher along channels than in open mudflat habitats. Infauna along channels were twice as abundant as those in the surrounding mudflats, and infauna were generally found closer to the surface. Furthermore, sediment alongside channels required less than half the force to probe, probably increasing shorebirds' overall access to prey. We conclude that shorebirds selected microhabitats near channels because of the greater food abundance and availability. Additional evidence suggested that another microhabitat, patches of red algae (Gracilaria sp.), may also have conferred advantages on foraging shorebirds. Channels may be a valuable microhabitat to consider when selecting conservation areas for shorebirds, and maintaining channels (e.g., through maintaining natural hydrology and sedimentation patterns) should be considered in management practices. Title: Small Tidal Channels Improve Foraging Opportunities for Calidris Shorebirds Author: Miller, Aileen K., de Rivera, Catherine E. Year: 2014 Estuarine intertidal habitats are heterogeneous, therefore migratory shorebirds are expected to forage in microhabitats where they can maximize their energy intake. Identifying proximate factors that migratory shorebirds use to accept or reject a particular habitat patch will help land managers make conservation and restoration decisions that provide the greatest benefits to shorebird populations during migration, a period of intense energy usage. We examined whether small semipermanent tidal channels were preferentially used by foraging Western Sandpipers (Calidris mauri) and Dunlins (C. alpina) during a spring migratory stopover in Bandon Marsh, an Oregon, USA, estuary. Further, we tested alternative hypotheses about how channels might be beneficial to shorebirds by examining infauna abundance, composition, and depth alongside channels compared with that in open mudflats. The densities of Western Sandpipers and Dunlins were at least 4 times higher along channels than in open mudflat habitats. Infauna along channels were twice as abundant as those in the surrounding mudflats, and infauna were generally found closer to the surface. Furthermore, sediment alongside channels required less than half the force to probe, probably increasing shorebirds' overall access to prey. We conclude that shorebirds selected microhabitats near channels because of the greater food abundance and availability. Additional evidence suggested that another microhabitat, patches of red algae (Gracilaria sp.), may also have conferred advantages on foraging shorebirds. Channels may be a valuable microhabitat to consider when selecting conservation areas for shorebirds, and maintaining channels (e.g., through maintaining natural hydrology and sedimentation patterns) should be considered in management practices. Close

• 19. [Article] Canada’s New Role in North American Energy Security

  Energy analysts have given renewed attention to Canada's position in the North American energy market since the September 11th attacks, because of fear that conflict might interrupt the flow of oil from ...

  Citation × Citation Citation Abstract Copy Title: Canada’s New Role in North American Energy Security Author: Smallman, Shawn Year: 2003 Energy analysts have given renewed attention to Canada's position in the North American energy market since the September 11th attacks, because of fear that conflict might interrupt the flow of oil from the Middle East. There are currently $30 billion (U.S.) in projects to develop the Alberta oil sands, in addition to new petroleum projects in Newfoundland, and major natural gas finds off the Atlantic coast. While Canada is already the single major oil exporter to the United States (ahead of both Saudi Arabia and Venezuela), its production could double by 2010. Canada’s rapidly increasing energy production has major implications both for hemispheric relations and for the United States’ strategic position. In the aftermath of September 11th energy security has become a key concern for the United States. After the attacks many commentators argued that the United States relied too heavily on Middle Eastern sources of supply, and that this factor not only limited U.S. actions, but also obliged the U.S. to engage with regimes that it found to be distasteful if not dangerous. Newspaper columnists decried America’s reliance on Saudi Arabia for energy, and some called for the United States to increase its energy self-sufficiency by opening the Arctic National Wildlife Refuge (ANWR) in Alaska to drilling. Richard Holbrook, the former U.S. ambassador to the United Nations, stated that Americans' "greatest single failure over the last twenty five years was our failure to reduce our dependence on foreign oil . . . which would have reduced the leverage of Saudi Arabia." Yet in fact the United States' energy position is now stronger than it appears, and there is reason to believe that it will improve in the future, because of the growing importance of Canada as a source of petroleum. Title: Canada’s New Role in North American Energy Security Author: Smallman, Shawn Year: 2003 Energy analysts have given renewed attention to Canada's position in the North American energy market since the September 11th attacks, because of fear that conflict might interrupt the flow of oil from the Middle East. There are currently $30 billion (U.S.) in projects to develop the Alberta oil sands, in addition to new petroleum projects in Newfoundland, and major natural gas finds off the Atlantic coast. While Canada is already the single major oil exporter to the United States (ahead of both Saudi Arabia and Venezuela), its production could double by 2010. Canada’s rapidly increasing energy production has major implications both for hemispheric relations and for the United States’ strategic position. In the aftermath of September 11th energy security has become a key concern for the United States. After the attacks many commentators argued that the United States relied too heavily on Middle Eastern sources of supply, and that this factor not only limited U.S. actions, but also obliged the U.S. to engage with regimes that it found to be distasteful if not dangerous. Newspaper columnists decried America’s reliance on Saudi Arabia for energy, and some called for the United States to increase its energy self-sufficiency by opening the Arctic National Wildlife Refuge (ANWR) in Alaska to drilling. Richard Holbrook, the former U.S. ambassador to the United Nations, stated that Americans' "greatest single failure over the last twenty five years was our failure to reduce our dependence on foreign oil . . . which would have reduced the leverage of Saudi Arabia." Yet in fact the United States' energy position is now stronger than it appears, and there is reason to believe that it will improve in the future, because of the growing importance of Canada as a source of petroleum. Close

• 20. [Article] A Geospatial Tool for Wetland Prioritization at the Watershed Scale

  There is an increasing demand for assessing ecosystem functions for freshwater wetlands, especially when comparing or prioritizing among wetlands at the watershed scale. We estimated the relative potential ...

  Citation × Citation Citation Abstract Copy Title: A Geospatial Tool for Wetland Prioritization at the Watershed Scale Author: Blackmore, Debra Sue, Chang, Heejun Year: 2015 There is an increasing demand for assessing ecosystem functions for freshwater wetlands, especially when comparing or prioritizing among wetlands at the watershed scale. We estimated the relative potential of selected ecosystem functions for freshwater wetlands within a watershed using widely available geospatial data. We developed four functions to estimate 1) flood storage, 2) late season flow, 3) sediment retention and 4) temperature control in four pilot watersheds in Oregon (Tualatin, Coquille, Upper Grande Ronde and Sprague). These watersheds are geographically separated from each other representing diverse ecoregion environments. Spatial analysis and geographic information system (GIS) were designed for maximum re-use, based on publicly-available data, commonly-used software, semi-automated techniques and wetland characterizations that attempt to capture fundamental wetland processes. Our data sources include 30-meter digital elevation models, NRCS soil survey extracts, USGS National Land Cover Data, USGS HUC8 boundaries (polygons) and statewide wetland delineations (polygons) processed within ArcGIS 10.2 and Python 2.7.5 software. Model parameters were compiled using multiple proxy values for size, slope, aspect, proximity, flow path distance, hydrologic gradient, shade, and soil characteristics. WPT characterizations emphasize the multi-faceted value of freshwater wetlands, relating potential within a watershed as well as providing model-based characterizations between watersheds. Our wetland prioritization tool (WPT) provides useful information to estimate and compare the relative potential for selected wetland functions, thereby improving success in wetland conservation, restoration, and mitigation efforts. Title: A Geospatial Tool for Wetland Prioritization at the Watershed Scale Author: Blackmore, Debra Sue, Chang, Heejun Year: 2015 There is an increasing demand for assessing ecosystem functions for freshwater wetlands, especially when comparing or prioritizing among wetlands at the watershed scale. We estimated the relative potential of selected ecosystem functions for freshwater wetlands within a watershed using widely available geospatial data. We developed four functions to estimate 1) flood storage, 2) late season flow, 3) sediment retention and 4) temperature control in four pilot watersheds in Oregon (Tualatin, Coquille, Upper Grande Ronde and Sprague). These watersheds are geographically separated from each other representing diverse ecoregion environments. Spatial analysis and geographic information system (GIS) were designed for maximum re-use, based on publicly-available data, commonly-used software, semi-automated techniques and wetland characterizations that attempt to capture fundamental wetland processes. Our data sources include 30-meter digital elevation models, NRCS soil survey extracts, USGS National Land Cover Data, USGS HUC8 boundaries (polygons) and statewide wetland delineations (polygons) processed within ArcGIS 10.2 and Python 2.7.5 software. Model parameters were compiled using multiple proxy values for size, slope, aspect, proximity, flow path distance, hydrologic gradient, shade, and soil characteristics. WPT characterizations emphasize the multi-faceted value of freshwater wetlands, relating potential within a watershed as well as providing model-based characterizations between watersheds. Our wetland prioritization tool (WPT) provides useful information to estimate and compare the relative potential for selected wetland functions, thereby improving success in wetland conservation, restoration, and mitigation efforts. Close