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• 4871. [Article] Unraveling the Diagenetic Cycling and Sequestration of Rare Earth Elements (REEs) in Marine Sediments to Understand REE Accumulation in Ancient Shales

  The rare earth elements (REEs) have been established as powerful tracers for a range of physiochemical processes occurring in the natural environment. They also hold significant economic importance as ...

  Citation × Citation Citation Abstract Copy Title: Unraveling the Diagenetic Cycling and Sequestration of Rare Earth Elements (REEs) in Marine Sediments to Understand REE Accumulation in Ancient Shales Author: Yang, Jon The rare earth elements (REEs) have been established as powerful tracers for a range of physiochemical processes occurring in the natural environment. They also hold significant economic importance as many technological advancements are reliant upon the REEs for their unique magnetic, luminescent, and electrochemical characteristics. In sedimentary settings, understanding the cycling of the REEs is necessary to fully evaluate the application of the REEs as geochemical tracers and to understand the individual factors that may influence the magnitude and signature of REE accumulations. The association of the REEs with the major element cycles of iron (Fe) and phosphorus (P) have been extensively studied in modern sedimentary environments, but there remain uncertainties on the long-term accumulation and signature of the REEs. In particular, our knowledge of how the preserved record of the REEs in ancient marine black shales is derived from the cycling of REEs observed within modern sediments remains unclear. For these marine black shales, this gap in our knowledge has important ramifications for the potential use of the REEs as geochemical tracers of processes such as groundwater flow or hydraulic fracturing activity as well as the potential sourcing of REEs for economic extraction. In this dissertation, I attempt to refine our understanding of REE accumulation in ancient black shales, and how the original signals of the diagenetic cycling of REEs may be preserved, by using sequential extraction techniques to isolate individual reactive components in the sediments. Based on documented associations of the REEs and authigenic phosphate minerals in sedimentary systems, I first investigated the preserved record of P in two ancient black shales to unravel the environmental parameters that lead to the sequestration of P as carbonate fluorapatite (CFA) (Chapter 2). This authigenic mineral form is considered the main sink term of reactive P from the marine system, and the preserved record of CFA and other components of the P system may help understand the cycling of P during the deposition of these ancient black shales. Using the preserved signals of P cycling in the individual components of the P system of the ancient black shales, I estimated paleo-benthic P fluxes to evaluate how the sedimentary cycling of P may have impacted the nutrient delivery to the depositional basin. During the deposition of the Marcellus black shale formation in the Devonian Appalachian Basin, I estimated paleo-benthic inputs of P from the depositional area of the formation that were within the ranges of P delivered from large river systems such as the pre-anthropogenic Mississippi and Mackenzie rivers. These estimates suggest that a return flux of P from the sediments could have been a significant source of nutrients to the basin and provide support for a positive feedback mechanism suggested for epeiric, anoxic basins where a benthic flux of P could help stimulate primary productivity in the surface, leading to a greater drawdown of oxygen in deeper waters, and helping to maintain anoxic conditions in the basin that may enhance organic matter preservation. In the next chapter, I add in the signals of the sedimentary REE cycle preserved in ancient black shales in association with the preserved components of the P cycle (Chapter 3). In addition to the two blacks shales analyzed in Chapter 2 for the individual components of the P system, I analyzed the whole-rock REE content of black shales ranging in age from Jurassic to Devonian and representing a range of depositional environments. In these black shales, authigenic carbonates and fluorapatites hosted the majority of the REEs in sediment intervals with active authigenic mineral formation. Based on published relationships between REEs in diagenetic coatings and pore waters in modern sediments, I inferred from the preserved REE patterns of authigenic phases in black shales that the recorded signature reflects diagenetic zonation, where middle REE-enrichments might record the signal of Fe-(oxyhydr)oxide dissolution and heavy REE- enrichments might record the later diagenetic signal of methanogenesis. In sedimentary intervals where authigenic phases were not prevalent, which represents the majority of the black shale samples investigated, the REE signal was determined by remnant organic matter hosting the light REEs and siliciclastic material hosting the heavy REEs. Although these samples resembled typical globally average shale patterns of the REEs, a closer inspection of the components of the REE system suggest that average values of shale may be an amalgamation of different REEs phases. While the connections between the cycling of P and the REEs have been investigated in Chapters 2 and 3, the connections with the cycling of Fe, which have also been demonstrated to strongly influence the REEs, are absent due to the labile nature of the reactive Fe components that are not preserved in the black shales. Thus, in the final chapter of the dissertation, I describe the REE content measured in reactive Fe and P phases from modern sediments near the island of South Georgia (sub-Antarctic) that receive a large input of reactive Fe through glacial output (Chapter 4). Despite this large input of reactive amorphous Fe-(oxyhydr)oxide phases, the majority of the REEs measured in the sediments were hosted in refractory siliciclastics, with less than 15 % of the REEs in labile (extractable) phases. The low sedimentary REE pool associated with the labile amorphous Fe phase likely reflects deposition under a very shallow (< 300 m) water column that precludes scavenging of REEs from seawater. This inference is consistent with measurements I performed on the dissolved REEs in the water column at South Georgia, that show relatively lower concentrations compared to deeper waters in the Southern Ocean that experience greater degrees of particulate scavenging. For the REEs that were measured within the labile amorphous Fe and P phases, however, I found that the fidelity of the preserved REE record is also impacted by the ability of the REEs to transfer from labile phases to more permanently stable mineral phases. With increasing sediment depth, the REEs demonstrated a transfer from relatively amorphous phases of Fe-(oxyhydr)oxides to more crystalline carbonate components of the sediments, preserving a characteristic middle-REE-enrichment of Fe-(oxyhydr)oxides. Additionally, the REEs also showed a transfer from more labile, amorphous forms of Fe- (oxyhydr)oxides such as ferrihydrite/lepidocrocite into more crystalline forms such as goethite/hematite phases with depth. Collectively, the chapters of this dissertation describe the formation of REE signatures from the initial signals of diagenesis in marine sediments to the preserved record observable today in ancient marine black shales. The major element cycles of Fe and P strongly influence the cycling of REEs both in the origination of REE signatures (e.g., middle-REE-enrichments from Fe-(oxyhydr)oxides) and preservation of diagenetic signals (e.g., middle- to heavy-REE-enrichments in authigenic carbonates and fluorapatites). The insights learned about REE accumulation in marine black shales, which I present in this dissertation might potentially assist in evaluating which phases of the black shales are accessed during groundwater or hydraulic fracturing studies as well as guiding targets for potential REE extraction. Title: Unraveling the Diagenetic Cycling and Sequestration of Rare Earth Elements (REEs) in Marine Sediments to Understand REE Accumulation in Ancient Shales Author: Yang, Jon The rare earth elements (REEs) have been established as powerful tracers for a range of physiochemical processes occurring in the natural environment. They also hold significant economic importance as many technological advancements are reliant upon the REEs for their unique magnetic, luminescent, and electrochemical characteristics. In sedimentary settings, understanding the cycling of the REEs is necessary to fully evaluate the application of the REEs as geochemical tracers and to understand the individual factors that may influence the magnitude and signature of REE accumulations. The association of the REEs with the major element cycles of iron (Fe) and phosphorus (P) have been extensively studied in modern sedimentary environments, but there remain uncertainties on the long-term accumulation and signature of the REEs. In particular, our knowledge of how the preserved record of the REEs in ancient marine black shales is derived from the cycling of REEs observed within modern sediments remains unclear. For these marine black shales, this gap in our knowledge has important ramifications for the potential use of the REEs as geochemical tracers of processes such as groundwater flow or hydraulic fracturing activity as well as the potential sourcing of REEs for economic extraction. In this dissertation, I attempt to refine our understanding of REE accumulation in ancient black shales, and how the original signals of the diagenetic cycling of REEs may be preserved, by using sequential extraction techniques to isolate individual reactive components in the sediments. Based on documented associations of the REEs and authigenic phosphate minerals in sedimentary systems, I first investigated the preserved record of P in two ancient black shales to unravel the environmental parameters that lead to the sequestration of P as carbonate fluorapatite (CFA) (Chapter 2). This authigenic mineral form is considered the main sink term of reactive P from the marine system, and the preserved record of CFA and other components of the P system may help understand the cycling of P during the deposition of these ancient black shales. Using the preserved signals of P cycling in the individual components of the P system of the ancient black shales, I estimated paleo-benthic P fluxes to evaluate how the sedimentary cycling of P may have impacted the nutrient delivery to the depositional basin. During the deposition of the Marcellus black shale formation in the Devonian Appalachian Basin, I estimated paleo-benthic inputs of P from the depositional area of the formation that were within the ranges of P delivered from large river systems such as the pre-anthropogenic Mississippi and Mackenzie rivers. These estimates suggest that a return flux of P from the sediments could have been a significant source of nutrients to the basin and provide support for a positive feedback mechanism suggested for epeiric, anoxic basins where a benthic flux of P could help stimulate primary productivity in the surface, leading to a greater drawdown of oxygen in deeper waters, and helping to maintain anoxic conditions in the basin that may enhance organic matter preservation. In the next chapter, I add in the signals of the sedimentary REE cycle preserved in ancient black shales in association with the preserved components of the P cycle (Chapter 3). In addition to the two blacks shales analyzed in Chapter 2 for the individual components of the P system, I analyzed the whole-rock REE content of black shales ranging in age from Jurassic to Devonian and representing a range of depositional environments. In these black shales, authigenic carbonates and fluorapatites hosted the majority of the REEs in sediment intervals with active authigenic mineral formation. Based on published relationships between REEs in diagenetic coatings and pore waters in modern sediments, I inferred from the preserved REE patterns of authigenic phases in black shales that the recorded signature reflects diagenetic zonation, where middle REE-enrichments might record the signal of Fe-(oxyhydr)oxide dissolution and heavy REE- enrichments might record the later diagenetic signal of methanogenesis. In sedimentary intervals where authigenic phases were not prevalent, which represents the majority of the black shale samples investigated, the REE signal was determined by remnant organic matter hosting the light REEs and siliciclastic material hosting the heavy REEs. Although these samples resembled typical globally average shale patterns of the REEs, a closer inspection of the components of the REE system suggest that average values of shale may be an amalgamation of different REEs phases. While the connections between the cycling of P and the REEs have been investigated in Chapters 2 and 3, the connections with the cycling of Fe, which have also been demonstrated to strongly influence the REEs, are absent due to the labile nature of the reactive Fe components that are not preserved in the black shales. Thus, in the final chapter of the dissertation, I describe the REE content measured in reactive Fe and P phases from modern sediments near the island of South Georgia (sub-Antarctic) that receive a large input of reactive Fe through glacial output (Chapter 4). Despite this large input of reactive amorphous Fe-(oxyhydr)oxide phases, the majority of the REEs measured in the sediments were hosted in refractory siliciclastics, with less than 15 % of the REEs in labile (extractable) phases. The low sedimentary REE pool associated with the labile amorphous Fe phase likely reflects deposition under a very shallow (< 300 m) water column that precludes scavenging of REEs from seawater. This inference is consistent with measurements I performed on the dissolved REEs in the water column at South Georgia, that show relatively lower concentrations compared to deeper waters in the Southern Ocean that experience greater degrees of particulate scavenging. For the REEs that were measured within the labile amorphous Fe and P phases, however, I found that the fidelity of the preserved REE record is also impacted by the ability of the REEs to transfer from labile phases to more permanently stable mineral phases. With increasing sediment depth, the REEs demonstrated a transfer from relatively amorphous phases of Fe-(oxyhydr)oxides to more crystalline carbonate components of the sediments, preserving a characteristic middle-REE-enrichment of Fe-(oxyhydr)oxides. Additionally, the REEs also showed a transfer from more labile, amorphous forms of Fe- (oxyhydr)oxides such as ferrihydrite/lepidocrocite into more crystalline forms such as goethite/hematite phases with depth. Collectively, the chapters of this dissertation describe the formation of REE signatures from the initial signals of diagenesis in marine sediments to the preserved record observable today in ancient marine black shales. The major element cycles of Fe and P strongly influence the cycling of REEs both in the origination of REE signatures (e.g., middle-REE-enrichments from Fe-(oxyhydr)oxides) and preservation of diagenetic signals (e.g., middle- to heavy-REE-enrichments in authigenic carbonates and fluorapatites). The insights learned about REE accumulation in marine black shales, which I present in this dissertation might potentially assist in evaluating which phases of the black shales are accessed during groundwater or hydraulic fracturing studies as well as guiding targets for potential REE extraction. Close

• 4872. [Article] Two Studies Addressing Practical Needs of Wheat Farmers, Processors, and Breeders : Changes in Falling Number and Alpha-amylase During Grain Storage, and Improved Predictions of Wheat-flour Dough Properties

  Wheat (Triticum aestivum) is a globally traded staple food crop. The diverse and pleasing nature of wheat-derived products is a result of the complex interactions of the polymeric components from the wheat ...

  Citation × Citation Citation Abstract Copy Title: Two Studies Addressing Practical Needs of Wheat Farmers, Processors, and Breeders : Changes in Falling Number and Alpha-amylase During Grain Storage, and Improved Predictions of Wheat-flour Dough Properties Author: Adams, Mike R. Wheat (Triticum aestivum) is a globally traded staple food crop. The diverse and pleasing nature of wheat-derived products is a result of the complex interactions of the polymeric components from the wheat endosperm. Changes in the functionality of these polymeric components, as a result of changes in growing conditions or different genetics, impacts market price and end-product quality and directly affects farmers and processors. Wheat is of particular economic importance to the U.S. Pacific Northwest and, specifically, to the state of Oregon. Providing quality wheat for export is paramount to the survival of the Oregon wheat industry. This dissertation focuses on wheat quality from the perspective of serving the practical needs of farmers, processors, and wheat breeders. The first study, split into two portions, concerns pre-harvest sprouting (PHS) and grain storage. PHS increases alpha amylase (αA) activity in wheat, which, in excess, reduces wheat end-product quality. Falling Number (FN) is the primary test used by industry to gauge PHS damage in wheats. Direct measurement of αA activity is the fundamental frame of reference. The objective of these studies was to determine if FN and αA activity of wheat samples changed during storage and if changes were a function of storage time, storage temperature, and degree of PHS damage. Samples from three Idaho locations were used. These captured a wide range of PHS degree, and therefore, wide ranges of FN values and αA activities. Samples were subdivided and stored at -20°C, +20°C, and +40°C. Low FN values and high αA activities were observed in soft wheats from locations that had rain events prior to harvest. Overall, FN and αA activity had the curvilinear relationship expected from the literature, indicating the validity of the sample set with regard to the FN/αA relationship. Changes in FN and αA activity were observed over a 90 day period of grain storage. FN differed between growing environments, wheat varieties, and storage temperatures. αA activity also differed between growing environments and wheat varieties, but not between storage temperatures. Highest rates of increase in FN were observed in hard wheats with high initial (day 0) FN values. Lowest rates of increase in FN were observed in soft wheats with low day 0 FN values. This contrasted with the changes that occurred in αA activity. Decreases in αA activity over storage time were most prevalent in soft wheats, particularly sprouted soft wheats (i.e. those with day 0 αA activities > 0.1 Ceralpha Units: CU). There were small decreases in αA activity in hard wheats but the distinction between high and low αA activity samples was not as evident as in the soft wheats because the vast majority of hard wheat samples tested had αA activities < 0.1 CU. Decreases in αA activity were in general not associated with corresponding increases in FN values over grain storage time. Increases in FN values occurred at a higher rates as storage temperature increased, particularly in hard wheats with high day 0 FN. Grain storage was successful as a way to raise FN values to > 300 s in very few cases. Storage was not effective in decreasing αA activity from > 0.1 CU to < 0.1 CU. Increases in FN over storage time for the hard wheats significantly differed between locations. However, decreases in αA activities over storage time for the hard wheats were not significantly different between locations. This again highlights a lack of correspondence between increased FN and decreased αA activity, suggesting that these two factors are somewhat decoupled when looking at changes in stored grain. Temperature-induced gluten crosslinking was explored as possible explanation for drastic increases in unsprouted hard wheat FN observed in samples from one location. Total polymeric protein (TPP) content was assessed at the end of the study for unsprouted hard wheats stored at +40°C and -20°C as well as sprouted hard wheats from stored at -20°C. TPP content was assessed as % large unextractable polymeric proteins (%LUPP) and % total unextractable polymeric proteins (%TUPP) using size exclusion high performance liquid chromatography (HPSEC). TPP content was not significantly different between storage temperatures for wheat varieties from the same location. %TUPP, but not % LUPP, was significantly lower in wheat varieties affected by PHS. Changes in FN at high storage temperature were not likely due to increased protein crosslinking. The second study aimed to validate the use of a rapid method for predicting dough strength at early generations in hard wheat breeding programs. Early generation quality screening improves breeding program efficiency. Hard wheats are used to make leavened bread products. The gluten proteins, particularly high molecular weight glutenin subunits (HMW-GS), form large, ramifying networks called the glutenin macropolymer (GMP). High GMP content is associated with increased dough strength and bread quality. Genetic differences in HMW-GS, and by inference, GMP, are responsible for differing dough properties between varieties. The Mixograph is used to measure dough mixing properties and predict end-product quality in breeding programs. GMP can also be measured as total polymeric protein (TPP) via HPSEC. The Solvent Retention Capacity (SRC) test has been proposed to predict hard wheat quality, specifically lactic acid SRC (LASRC). The objectives of this research were to provide preliminary information on the usefulness of using LASRC, on its own, to predict dough mixing properties, specifically as applied to early generation screening in a wheat breeding program, and to assess the relationship between LASRC and TPP. Wheat samples were categorized by flour protein concentration (FPC). Mixograph analysis was used as the baseline for dough properties and was analyzed both by eye and by the proprietary Mixsmart software. TPP content was assessed as %LUPP and %%TUPP. As a result of redundancy between the two TPP measures, only %LUPP was used for statistical analysis. Dough mixing parameters were slightly better correlated with LASRC than %LUPP. Correlations between LASRC, %LUPP, and dough mixing parameters were different between FPC categories, particularly in low FPC samples. A strict cutoff of 115% LASRC effectively screened out the bottom 10% of low quality hard wheats but retained a nearly equal amount of low quality hard wheats that would have been screened out by mixograph analysis. LASRC and %LUPP are not likely to be effective predictors of dough properties, but may have some value to screen for hard wheat quality in the early generations of a wheat breeding cycle. Title: Two Studies Addressing Practical Needs of Wheat Farmers, Processors, and Breeders : Changes in Falling Number and Alpha-amylase During Grain Storage, and Improved Predictions of Wheat-flour Dough Properties Author: Adams, Mike R. Wheat (Triticum aestivum) is a globally traded staple food crop. The diverse and pleasing nature of wheat-derived products is a result of the complex interactions of the polymeric components from the wheat endosperm. Changes in the functionality of these polymeric components, as a result of changes in growing conditions or different genetics, impacts market price and end-product quality and directly affects farmers and processors. Wheat is of particular economic importance to the U.S. Pacific Northwest and, specifically, to the state of Oregon. Providing quality wheat for export is paramount to the survival of the Oregon wheat industry. This dissertation focuses on wheat quality from the perspective of serving the practical needs of farmers, processors, and wheat breeders. The first study, split into two portions, concerns pre-harvest sprouting (PHS) and grain storage. PHS increases alpha amylase (αA) activity in wheat, which, in excess, reduces wheat end-product quality. Falling Number (FN) is the primary test used by industry to gauge PHS damage in wheats. Direct measurement of αA activity is the fundamental frame of reference. The objective of these studies was to determine if FN and αA activity of wheat samples changed during storage and if changes were a function of storage time, storage temperature, and degree of PHS damage. Samples from three Idaho locations were used. These captured a wide range of PHS degree, and therefore, wide ranges of FN values and αA activities. Samples were subdivided and stored at -20°C, +20°C, and +40°C. Low FN values and high αA activities were observed in soft wheats from locations that had rain events prior to harvest. Overall, FN and αA activity had the curvilinear relationship expected from the literature, indicating the validity of the sample set with regard to the FN/αA relationship. Changes in FN and αA activity were observed over a 90 day period of grain storage. FN differed between growing environments, wheat varieties, and storage temperatures. αA activity also differed between growing environments and wheat varieties, but not between storage temperatures. Highest rates of increase in FN were observed in hard wheats with high initial (day 0) FN values. Lowest rates of increase in FN were observed in soft wheats with low day 0 FN values. This contrasted with the changes that occurred in αA activity. Decreases in αA activity over storage time were most prevalent in soft wheats, particularly sprouted soft wheats (i.e. those with day 0 αA activities > 0.1 Ceralpha Units: CU). There were small decreases in αA activity in hard wheats but the distinction between high and low αA activity samples was not as evident as in the soft wheats because the vast majority of hard wheat samples tested had αA activities < 0.1 CU. Decreases in αA activity were in general not associated with corresponding increases in FN values over grain storage time. Increases in FN values occurred at a higher rates as storage temperature increased, particularly in hard wheats with high day 0 FN. Grain storage was successful as a way to raise FN values to > 300 s in very few cases. Storage was not effective in decreasing αA activity from > 0.1 CU to < 0.1 CU. Increases in FN over storage time for the hard wheats significantly differed between locations. However, decreases in αA activities over storage time for the hard wheats were not significantly different between locations. This again highlights a lack of correspondence between increased FN and decreased αA activity, suggesting that these two factors are somewhat decoupled when looking at changes in stored grain. Temperature-induced gluten crosslinking was explored as possible explanation for drastic increases in unsprouted hard wheat FN observed in samples from one location. Total polymeric protein (TPP) content was assessed at the end of the study for unsprouted hard wheats stored at +40°C and -20°C as well as sprouted hard wheats from stored at -20°C. TPP content was assessed as % large unextractable polymeric proteins (%LUPP) and % total unextractable polymeric proteins (%TUPP) using size exclusion high performance liquid chromatography (HPSEC). TPP content was not significantly different between storage temperatures for wheat varieties from the same location. %TUPP, but not % LUPP, was significantly lower in wheat varieties affected by PHS. Changes in FN at high storage temperature were not likely due to increased protein crosslinking. The second study aimed to validate the use of a rapid method for predicting dough strength at early generations in hard wheat breeding programs. Early generation quality screening improves breeding program efficiency. Hard wheats are used to make leavened bread products. The gluten proteins, particularly high molecular weight glutenin subunits (HMW-GS), form large, ramifying networks called the glutenin macropolymer (GMP). High GMP content is associated with increased dough strength and bread quality. Genetic differences in HMW-GS, and by inference, GMP, are responsible for differing dough properties between varieties. The Mixograph is used to measure dough mixing properties and predict end-product quality in breeding programs. GMP can also be measured as total polymeric protein (TPP) via HPSEC. The Solvent Retention Capacity (SRC) test has been proposed to predict hard wheat quality, specifically lactic acid SRC (LASRC). The objectives of this research were to provide preliminary information on the usefulness of using LASRC, on its own, to predict dough mixing properties, specifically as applied to early generation screening in a wheat breeding program, and to assess the relationship between LASRC and TPP. Wheat samples were categorized by flour protein concentration (FPC). Mixograph analysis was used as the baseline for dough properties and was analyzed both by eye and by the proprietary Mixsmart software. TPP content was assessed as %LUPP and %%TUPP. As a result of redundancy between the two TPP measures, only %LUPP was used for statistical analysis. Dough mixing parameters were slightly better correlated with LASRC than %LUPP. Correlations between LASRC, %LUPP, and dough mixing parameters were different between FPC categories, particularly in low FPC samples. A strict cutoff of 115% LASRC effectively screened out the bottom 10% of low quality hard wheats but retained a nearly equal amount of low quality hard wheats that would have been screened out by mixograph analysis. LASRC and %LUPP are not likely to be effective predictors of dough properties, but may have some value to screen for hard wheat quality in the early generations of a wheat breeding cycle. Close

• 4873. [Article] The readability of selected fourth, fifth and sixth grade social studies textbooks as determined by the Fry Readability Graph and group informal reading inventories

  The purpose of this study was to determine the readability of three series of selected fourth, fifth and sixth grade social studies textbooks, a total of nine textbooks. Two measurements were utilized: ...

  Citation × Citation Citation Abstract Copy Title: The readability of selected fourth, fifth and sixth grade social studies textbooks as determined by the Fry Readability Graph and group informal reading inventories Author: Pruitt, Janet Louise McCracken, 1930- The purpose of this study was to determine the readability of three series of selected fourth, fifth and sixth grade social studies textbooks, a total of nine textbooks. Two measurements were utilized: a readability formula (Fry Readability Graph; Knapp, 1971), and group informal reading inventories. The Fry Graph was applied to ten randomly selected 100-word passages in each text; proper nouns were included in the computations. The group informal reading inventories were constructed from the same textbooks on passages not previously taught. The tests were administered in May to 1467 students in 70 randomly selected classrooms. Procedures Both descriptive and inferential statistics were used to analyze the data. Readability levels derived by the application of the Fry Graph were reported for each text:, as was the range of readability within each text and the deviation of each sample from the overall readability of the textbook. The test scores from the group informal reading inventories were first reported in terms of the per cent of subjects scoring at the independent level (90, 95, 100 per cent), the instructional level (65, 70, 75, 80, 85 per cent) and the frustration level (60 per cent or lower) for the sample. Mean test scores were also computed for the sample, the urban and rural subsets, for each grade level, and for each publisher. To determine if there were significant differences among the subsets, the following null hypotheses were formulated: H1 There are no significant differences in the mean scores of the urban and rural subjects. H2 There are no significant differences in the mean scores of the subjects in grades four, five and six. H3 There are no significant differences among the mean scores of the subjects tested on the D. C. Heath, the Silver Burdett or the Benefic Press social studies series. The differences among groups were statistically analyzed at the .05 and .01 levels of significance by the pooled variance t-test or by the analysis of variance. When the analysis of variance resulted in a significant F value, the multiple range test was applied to determine the exact location of the mean difference. Findings of the Study Fry Readability Graph 1. Two textbooks, the fourth grade textbooks published by D. C. Heath and by Silver Burdett, had readability levels in agreement with the publishers' designated grade level. The remaining seven textbooks had readability levels one to four years above the designated grade level. 2. The average range of readability within the textbooks was 6.2 years with little evidence of a gradation from less difficult to more difficult reading material. 3. When the three series were ranked according to their relative difficulty on the basis of the Fry readability levels, it was determined that: a. The D. C. Heath series is the least difficult. b. The Silver Burdett series occupies an intermediate position. c. The Benefic Press series is the most difficult. Group Informal Reading Inventories 1. For the total sample 9.41 per cent of the subjects scored at the independent level, 32.04 per cent scored at the instructional level, and 58.55 per cent scored at the frustration level. The mean test score was 54.82 per cent. 2. The mean score (56.64 per cent) of the urban subjects was significantly higher (.01 level) than the mean score (52.35 per cent) of the rural subjects with a t value of 3.30. H1 was rejected. 3. The mean score for grade four was 50.35 per cent; for grade five, 53.96 per cent; and for grade six, 58.09 per cent. The analysis of variance resulted in a significant (.01 level) F value of 11.25. H2 was rejected. 4. The mean scores by publisher were: D. C. Heath, 55.88 per cent; Silver Burdett, 55.77 per cent; and Benefic Press, 50.38 per cent. The analysis of variance resulted in a significant (.01 level) F value of 5.47. The multiple range test showed significant differences between the Benefic Press and D. C. Heath series (.01 level) and between the Benefic Press and Silver Burdett series (.01 level). There were no significant differences between the D. C. Heath and Silver Burdett series. H3 was rejected. Implications 1. The findings of this study did not differ in any substantial way from the results reported by earlier investigators. Further studies of the readability of social studies textbooks similar in content to the three series analyzed in this investigation is not warranted. 2. Additional research is needed to determine if systematic instruction resembling the procedures prescribed for a well-developed basal reading lesson will significantly affect the ability of students to comprehend social studies textbooks. Recommendations 1. Current knowledge regarding the nature of concept development should be a major consideration for curriculum decisions in the social studies. Piaget's findings regarding the cognitive functioning of elementary school students provide insight regarding the capabilities of this age group. 2. Instructional alternatives that diminish heavy reliance on the textbook should be afforded a fair trial in the classroom, e.g., simulation and gaming (Cuetzkow, 1962; Inbar, 1972), process analog (Fielder, 1967; Joyce, 1972), role playing (Shaftel, 1967), problem solving (Fenton, 1967; Shaftel, 1967), inquiry approaches (Suchman, 1964; Clements, Fielder, Tabachnick, 1966), and inductive development of concepts and generalizations (Taba, 1966; Fenton, 1966; Hanna, 1965). 3. A publicly financed agency staffed by expert analysts and equipped with the necessary resources should be established to provide analysis services to publishers and to determine the difficulty level of all instructional materials as they appear on the market (Bormuth, 1971). 4. Pilot editions of all new textbooks should be field tested with a cross section of students using procedures similar to those utilized by the publishers of standardized tests. 5. When classroom teachers make reading assignments in social studies textbooks, they should faithfully adhere to the same procedures that are prescribed for a well-developed basal reading lesson. Title: The readability of selected fourth, fifth and sixth grade social studies textbooks as determined by the Fry Readability Graph and group informal reading inventories Author: Pruitt, Janet Louise McCracken, 1930- The purpose of this study was to determine the readability of three series of selected fourth, fifth and sixth grade social studies textbooks, a total of nine textbooks. Two measurements were utilized: a readability formula (Fry Readability Graph; Knapp, 1971), and group informal reading inventories. The Fry Graph was applied to ten randomly selected 100-word passages in each text; proper nouns were included in the computations. The group informal reading inventories were constructed from the same textbooks on passages not previously taught. The tests were administered in May to 1467 students in 70 randomly selected classrooms. Procedures Both descriptive and inferential statistics were used to analyze the data. Readability levels derived by the application of the Fry Graph were reported for each text:, as was the range of readability within each text and the deviation of each sample from the overall readability of the textbook. The test scores from the group informal reading inventories were first reported in terms of the per cent of subjects scoring at the independent level (90, 95, 100 per cent), the instructional level (65, 70, 75, 80, 85 per cent) and the frustration level (60 per cent or lower) for the sample. Mean test scores were also computed for the sample, the urban and rural subsets, for each grade level, and for each publisher. To determine if there were significant differences among the subsets, the following null hypotheses were formulated: H1 There are no significant differences in the mean scores of the urban and rural subjects. H2 There are no significant differences in the mean scores of the subjects in grades four, five and six. H3 There are no significant differences among the mean scores of the subjects tested on the D. C. Heath, the Silver Burdett or the Benefic Press social studies series. The differences among groups were statistically analyzed at the .05 and .01 levels of significance by the pooled variance t-test or by the analysis of variance. When the analysis of variance resulted in a significant F value, the multiple range test was applied to determine the exact location of the mean difference. Findings of the Study Fry Readability Graph 1. Two textbooks, the fourth grade textbooks published by D. C. Heath and by Silver Burdett, had readability levels in agreement with the publishers' designated grade level. The remaining seven textbooks had readability levels one to four years above the designated grade level. 2. The average range of readability within the textbooks was 6.2 years with little evidence of a gradation from less difficult to more difficult reading material. 3. When the three series were ranked according to their relative difficulty on the basis of the Fry readability levels, it was determined that: a. The D. C. Heath series is the least difficult. b. The Silver Burdett series occupies an intermediate position. c. The Benefic Press series is the most difficult. Group Informal Reading Inventories 1. For the total sample 9.41 per cent of the subjects scored at the independent level, 32.04 per cent scored at the instructional level, and 58.55 per cent scored at the frustration level. The mean test score was 54.82 per cent. 2. The mean score (56.64 per cent) of the urban subjects was significantly higher (.01 level) than the mean score (52.35 per cent) of the rural subjects with a t value of 3.30. H1 was rejected. 3. The mean score for grade four was 50.35 per cent; for grade five, 53.96 per cent; and for grade six, 58.09 per cent. The analysis of variance resulted in a significant (.01 level) F value of 11.25. H2 was rejected. 4. The mean scores by publisher were: D. C. Heath, 55.88 per cent; Silver Burdett, 55.77 per cent; and Benefic Press, 50.38 per cent. The analysis of variance resulted in a significant (.01 level) F value of 5.47. The multiple range test showed significant differences between the Benefic Press and D. C. Heath series (.01 level) and between the Benefic Press and Silver Burdett series (.01 level). There were no significant differences between the D. C. Heath and Silver Burdett series. H3 was rejected. Implications 1. The findings of this study did not differ in any substantial way from the results reported by earlier investigators. Further studies of the readability of social studies textbooks similar in content to the three series analyzed in this investigation is not warranted. 2. Additional research is needed to determine if systematic instruction resembling the procedures prescribed for a well-developed basal reading lesson will significantly affect the ability of students to comprehend social studies textbooks. Recommendations 1. Current knowledge regarding the nature of concept development should be a major consideration for curriculum decisions in the social studies. Piaget's findings regarding the cognitive functioning of elementary school students provide insight regarding the capabilities of this age group. 2. Instructional alternatives that diminish heavy reliance on the textbook should be afforded a fair trial in the classroom, e.g., simulation and gaming (Cuetzkow, 1962; Inbar, 1972), process analog (Fielder, 1967; Joyce, 1972), role playing (Shaftel, 1967), problem solving (Fenton, 1967; Shaftel, 1967), inquiry approaches (Suchman, 1964; Clements, Fielder, Tabachnick, 1966), and inductive development of concepts and generalizations (Taba, 1966; Fenton, 1966; Hanna, 1965). 3. A publicly financed agency staffed by expert analysts and equipped with the necessary resources should be established to provide analysis services to publishers and to determine the difficulty level of all instructional materials as they appear on the market (Bormuth, 1971). 4. Pilot editions of all new textbooks should be field tested with a cross section of students using procedures similar to those utilized by the publishers of standardized tests. 5. When classroom teachers make reading assignments in social studies textbooks, they should faithfully adhere to the same procedures that are prescribed for a well-developed basal reading lesson. Close

• 4874. [Article] Assessing equity in health system finance and health care utilization : the case of Chile, and a model to measure health care access

  Chile has experienced great success in terms of economic growth in the last decades. This growing economy brings changes in the Chilean health care system. Its health care system was primarily funded by ...

  Citation × Citation Citation Abstract Copy Title: Assessing equity in health system finance and health care utilization : the case of Chile, and a model to measure health care access Author: Nunez Mondaca, Alicia Lorena Chile has experienced great success in terms of economic growth in the last decades. This growing economy brings changes in the Chilean health care system. Its health care system was primarily funded by state sources until 1981, when a major reform was introduced that established new rules for the health insurance market. Since then, Chile has a public-private mixed health care system, both in financing and delivery of services. Citizens can choose for coverage between the Public National Health Insurance and the Private Health Insurance system. However, these systems have a common funding source coming from the mandatory contribution of employees, equivalent to 7% of their taxable income with an approximate limit of US$2,800 dollars. One of the more important Chilean health reforms towards the establishment of social guarantees was effective on July 2005, when the Regime of Explicit Health Guarantees, also known as Plan AUGE became effective. Plan AUGE is a health program that benefits all Chileans without discrimination of age, gender, economic status, health care, or place of residence. This plan includes the 69 diseases with higher impact on Chilean population in its different stages, but with feasibility of effective treatments. Changes in the health care system and its last reform brought questions about their impact on the distribution of health care services throughout country. Is Chile moving towards a better and more equitable health care system? The main purpose of this thesis is to investigate equity in health system finance and health care utilization as well as to explore alternative measurement of access to health care in Chile. The first two manuscripts examine equity issues in Chile. The purpose of the first one is to assess equity in health system finance in Chile, accounting for all finance sources. While equity in health system finance has been well studied in OECD countries, there are still few published empirical studies on Latin American health care systems, where there tends to be a wider gap in income-wealth distribution among states. This gap may increase the financial burden for people in the lower spectrum of income groups, which is the main concern in the first manuscript. It will focus on identifying policy variables that may contribute to more equitable distribution of the financial burden in health care. The equity principle we adopt for this study is the ability to pay principle. Based on this, we explore factors that contribute to inequities in the health care system finance and issues about who bears the heavier burden of out-of pocket (OOP) payment, progressivity of OOP payment, and the redistributive effect of OOP payment for health care as a source of finance in the Chilean health care system. Our analysis is based on data from the National Socioeconomic Survey (CASEN), and the 2006 National Survey on Satisfaction and OOP payments. Results from this study provide comprehensive understanding of the financial burden of health care in Chile. This study identified evidence of inequity, in spite of the progressivity of the health care system. Furthermore, our assessment of equity in health system finance identified relevant policy variables such as education, insurance system, and method of payment that should be taken into consideration in the ongoing debates and research in improving the Chilean system. Such findings will also benefit other Latin American countries that are concerned about equity in health system finance. The purpose of the second manuscript was to assess equity in health care utilization in Chile. Secondary data analyses from the National Socioeconomic Survey (CASEN) were performed to estimate the impact of different factors including AUGE in the utilization of health care services. We used a two-part model for the analysis of frequency of health care use in the country. Four other separate two-part models were also specified to estimate the frequency of use of preventive services, general practitioner services, specialty care and emergency care. An assessment of horizontal equity was also included. Results suggest the presence of pro-rich inequities in the use of medical care. The estimation of the two-part model found key factors affecting utilization of health care services such as education and the implementation of the AUGE program. These findings provide timely evidence to policy-makers to understand the current distribution and equity of health care utilization, and to strengthen availability of health services accordingly. The third manuscript was motivated by the previous findings. Its purpose was to explore an alternative measurement for health care access. The majority of studies nowadays use a single proxy to estimate access: the use of health care services. However, we saw many limitations on this approach since it only considers people that are already using the system and ignores those that are not. The final manuscript proposed a model to estimate access to health care services based on communitarian claims. The model identified barriers to health care access as well as the preferences of the community for priority settings. Title: Assessing equity in health system finance and health care utilization : the case of Chile, and a model to measure health care access Author: Nunez Mondaca, Alicia Lorena Chile has experienced great success in terms of economic growth in the last decades. This growing economy brings changes in the Chilean health care system. Its health care system was primarily funded by state sources until 1981, when a major reform was introduced that established new rules for the health insurance market. Since then, Chile has a public-private mixed health care system, both in financing and delivery of services. Citizens can choose for coverage between the Public National Health Insurance and the Private Health Insurance system. However, these systems have a common funding source coming from the mandatory contribution of employees, equivalent to 7% of their taxable income with an approximate limit of US$2,800 dollars. One of the more important Chilean health reforms towards the establishment of social guarantees was effective on July 2005, when the Regime of Explicit Health Guarantees, also known as Plan AUGE became effective. Plan AUGE is a health program that benefits all Chileans without discrimination of age, gender, economic status, health care, or place of residence. This plan includes the 69 diseases with higher impact on Chilean population in its different stages, but with feasibility of effective treatments. Changes in the health care system and its last reform brought questions about their impact on the distribution of health care services throughout country. Is Chile moving towards a better and more equitable health care system? The main purpose of this thesis is to investigate equity in health system finance and health care utilization as well as to explore alternative measurement of access to health care in Chile. The first two manuscripts examine equity issues in Chile. The purpose of the first one is to assess equity in health system finance in Chile, accounting for all finance sources. While equity in health system finance has been well studied in OECD countries, there are still few published empirical studies on Latin American health care systems, where there tends to be a wider gap in income-wealth distribution among states. This gap may increase the financial burden for people in the lower spectrum of income groups, which is the main concern in the first manuscript. It will focus on identifying policy variables that may contribute to more equitable distribution of the financial burden in health care. The equity principle we adopt for this study is the ability to pay principle. Based on this, we explore factors that contribute to inequities in the health care system finance and issues about who bears the heavier burden of out-of pocket (OOP) payment, progressivity of OOP payment, and the redistributive effect of OOP payment for health care as a source of finance in the Chilean health care system. Our analysis is based on data from the National Socioeconomic Survey (CASEN), and the 2006 National Survey on Satisfaction and OOP payments. Results from this study provide comprehensive understanding of the financial burden of health care in Chile. This study identified evidence of inequity, in spite of the progressivity of the health care system. Furthermore, our assessment of equity in health system finance identified relevant policy variables such as education, insurance system, and method of payment that should be taken into consideration in the ongoing debates and research in improving the Chilean system. Such findings will also benefit other Latin American countries that are concerned about equity in health system finance. The purpose of the second manuscript was to assess equity in health care utilization in Chile. Secondary data analyses from the National Socioeconomic Survey (CASEN) were performed to estimate the impact of different factors including AUGE in the utilization of health care services. We used a two-part model for the analysis of frequency of health care use in the country. Four other separate two-part models were also specified to estimate the frequency of use of preventive services, general practitioner services, specialty care and emergency care. An assessment of horizontal equity was also included. Results suggest the presence of pro-rich inequities in the use of medical care. The estimation of the two-part model found key factors affecting utilization of health care services such as education and the implementation of the AUGE program. These findings provide timely evidence to policy-makers to understand the current distribution and equity of health care utilization, and to strengthen availability of health services accordingly. The third manuscript was motivated by the previous findings. Its purpose was to explore an alternative measurement for health care access. The majority of studies nowadays use a single proxy to estimate access: the use of health care services. However, we saw many limitations on this approach since it only considers people that are already using the system and ignores those that are not. The final manuscript proposed a model to estimate access to health care services based on communitarian claims. The model identified barriers to health care access as well as the preferences of the community for priority settings. Close

• 4875. [Article] All in a DNA's work : conservation genetics and monitoring of the New Zealand endemic Maui's and Hector's dolphins

  The critically endangered Maui's dolphin (Cephalorhynchus hectori maui) and the endangered Hector's dolphin (C. h. hectori) are endemic to the coastal waters of New Zealand, where their primary threat ...

  Citation × Citation Citation Abstract Copy Title: All in a DNA's work : conservation genetics and monitoring of the New Zealand endemic Maui's and Hector's dolphins Author: Hamner, Rebecca Marie The critically endangered Maui's dolphin (Cephalorhynchus hectori maui) and the endangered Hector's dolphin (C. h. hectori) are endemic to the coastal waters of New Zealand, where their primary threat is fisheries-related mortality. The Maui's dolphin is among the most critically endangered cetaceans in the world, with its remnant population primarily concentrated in approximately 140 km along the central west coast of New Zealand's North Island. Its closely related sister subspecies, the Hector's dolphin, is more abundant and offers a useful comparison for studying the Maui's dolphin. My work used genetic tools to examine demographic and genetic parameters relevant for conservation considerations regarding Maui's and Hector's dolphins, as well as to build upon past genetic baselines for the purpose of long-term genetic monitoring of these subspecies. Three genetic datasets formed the basis for most analyses: (1) Maui's 01-07, including 54 Maui's dolphin individuals sampled between 2001 and 2007 (n = 70 biopsies, 12 beachcast); (2) Maui's 10-11, including 40 Maui's dolphin individuals sampled in 2010 and 2011 (n = 69 biopsies, 1 beachcast); and (3) Hector's CB11-12, including 148 Hector's dolphin individuals sampled in Cloudy Bay in 2011 and 2012 (n = 263 biopsies). Microsatellite genotypes were used to identify individuals for a genotype recapture abundance estimate of individuals age 1⁺ (N₁₊) and for the estimation of effective population size (N[subscript e]). Both populations exhibited a high N[subscript e] relative to N₁₊, consistent with expectations given their life history characteristics and the limited data available for other dolphin species. The abundance of Maui's dolphins was confirmed to be very low, Maui's 10-11 N₁₊ = 55 (95% CL = 48 - 69), and as expected, it had much lower linkage disequilibrium N[subscript e] (61, 95% CL = 29 - 338) than Hector's CB11-12 (N[subscript e] = 207, 95% CL = 127 - 447; N₁₊ = 272, 95% CL = 236 - 323). The slightly higher Ne/N₁₊ ratio of the Maui's dolphin compared to the Hector's dolphin is consistent with a recent decline in the Maui's dolphin. Although the point estimates of both N[subscript e] and N₁₊ decreased between the two Maui's dolphin datasets (Maui's 01-07: N[subscript e] = 74, 95% CL = 37 - 318; N₁₊ = 69, 95% CL = 38 - 125), the confidence intervals widely overlapped. Maui's 10-11 had significantly fewer alleles (average 4 alleles/locus) and lower heterozygosity (H₀ = 0.316, H[subscript e] = 0.319) than Hector's CB11-12 (average 7 alleles/locus, H₀ = 0.500, H[subscript e] = 0.495; all P <0.001). Interestingly, one microsatellite locus (PPHO104) had anomalously high diversity (31 to 63 alleles) in both Hector's and Maui's dolphins and appears to be influenced by diversifying selection. The observed and expected heterozygosity, internal relatedness, and F[subscript IS] of Maui's dolphins all showed patterns consistent with a decline of the subspecies, although none differed significantly over the short time interval between the two datasets collected in 2001-07 and 2010-11. The lack of significant decline in any of the parameters analyzed for Maui's dolphins is not surprising given the low power to detect a low to moderate decline over the short interval (<1 generation) between the two sampling periods. Compared to minimum viable effective population sizes proposed to guide management decisions, the Maui's dolphin has declined below the recommended threshold of N[subscript e] = 50, recently increased to N[subscript e] ≥100, thought to be necessary to avoid inbreeding depression in the short term (5 generations, ~65.2 years for Maui's and Hector's dolphins). Additionally, both the Maui's dolphin and Cloudy Bay Hector's dolphin populations are below the recommended threshold of N[subscript e] = 500, recently increased to N[subscript e] ≥1000, thought to be necessary to preserve long-term evolutionary potential. This is less of a concern for the Cloudy Bay Hector's population, which is thought to maintain gene flow with neighboring populations. However, for the small, isolated Maui's dolphin population, inbreeding depression is likely to be an increasing concern. Furthermore, each Maui's dolphin individual holds a disproportionate amount of the total genetic variation of the subspecies and would represent a disproportionately large demographic and genetic loss if it died before realizing its reproductive potential in the population. There is, however, potential for genetic restoration by interbreeding with Hector's dolphins, as genetic monitoring of Maui's dolphins revealed the first contemporary dispersal of four (two living females, one dead female, one dead male) Hector's dolphins into the Maui's dolphin distribution. Two Hector's dolphins (one dead female neonate, one living male) were also sampled along the North Island's southwest coast, outside the presumed range of either subspecies. Together, these records provide evidence of long-distance dispersal by Hector's dolphins (≥400 km) and the possibility of an unsampled Hector's dolphin population along the southwest coast of the North Island or northern South Island. These results highlight the value of genetic monitoring for subspecies lacking distinctive physical appearances, as such discoveries are not detected by other means but have important conservation implications. Although the Maui's dolphin is critically endangered, it is not necessarily doomed to extinction. The subspecies appears to be maintaining an equal sex ratio and connectivity within its remnant range, and the highly diverse locus PPHO104 could potentially offer clues to an inbreeding avoidance mechanism. If Maui's dolphins interbreed with the recently identified Hector's dolphin immigrants, it could provide genetic restoration, enhancing chances of long-term survival of the Maui's dolphin. Continued genetic monitoring and examination of recovered carcasses for phenotypic signs of inbreeding are important for gauging genetic threats to the survival of Maui's dolphins, as well as determining if any Hector's dolphin populations appear to be declining toward the critically endangered state of the Maui's dolphin. The results of this work contributed to the decision by the New Zealand Department of Conservation and Ministry for Primary Industries to conduct an updated risk assessment for Maui's dolphins and accelerate the review of the Maui's Dolphin Threat Management Plan. Consequently, commercial and recreational set net restrictions were extended slightly to reduce entanglement risk to Maui's dolphins utilizing the southern part of their distribution, as well as any Hector's dolphins that disperse north into that area. The results related to the population of Hector's dolphins in Cloudy Bay provide information that will contribute to the upcoming review of the Hector's dolphin component of the Threat Management Plan. Title: All in a DNA's work : conservation genetics and monitoring of the New Zealand endemic Maui's and Hector's dolphins Author: Hamner, Rebecca Marie The critically endangered Maui's dolphin (Cephalorhynchus hectori maui) and the endangered Hector's dolphin (C. h. hectori) are endemic to the coastal waters of New Zealand, where their primary threat is fisheries-related mortality. The Maui's dolphin is among the most critically endangered cetaceans in the world, with its remnant population primarily concentrated in approximately 140 km along the central west coast of New Zealand's North Island. Its closely related sister subspecies, the Hector's dolphin, is more abundant and offers a useful comparison for studying the Maui's dolphin. My work used genetic tools to examine demographic and genetic parameters relevant for conservation considerations regarding Maui's and Hector's dolphins, as well as to build upon past genetic baselines for the purpose of long-term genetic monitoring of these subspecies. Three genetic datasets formed the basis for most analyses: (1) Maui's 01-07, including 54 Maui's dolphin individuals sampled between 2001 and 2007 (n = 70 biopsies, 12 beachcast); (2) Maui's 10-11, including 40 Maui's dolphin individuals sampled in 2010 and 2011 (n = 69 biopsies, 1 beachcast); and (3) Hector's CB11-12, including 148 Hector's dolphin individuals sampled in Cloudy Bay in 2011 and 2012 (n = 263 biopsies). Microsatellite genotypes were used to identify individuals for a genotype recapture abundance estimate of individuals age 1⁺ (N₁₊) and for the estimation of effective population size (N[subscript e]). Both populations exhibited a high N[subscript e] relative to N₁₊, consistent with expectations given their life history characteristics and the limited data available for other dolphin species. The abundance of Maui's dolphins was confirmed to be very low, Maui's 10-11 N₁₊ = 55 (95% CL = 48 - 69), and as expected, it had much lower linkage disequilibrium N[subscript e] (61, 95% CL = 29 - 338) than Hector's CB11-12 (N[subscript e] = 207, 95% CL = 127 - 447; N₁₊ = 272, 95% CL = 236 - 323). The slightly higher Ne/N₁₊ ratio of the Maui's dolphin compared to the Hector's dolphin is consistent with a recent decline in the Maui's dolphin. Although the point estimates of both N[subscript e] and N₁₊ decreased between the two Maui's dolphin datasets (Maui's 01-07: N[subscript e] = 74, 95% CL = 37 - 318; N₁₊ = 69, 95% CL = 38 - 125), the confidence intervals widely overlapped. Maui's 10-11 had significantly fewer alleles (average 4 alleles/locus) and lower heterozygosity (H₀ = 0.316, H[subscript e] = 0.319) than Hector's CB11-12 (average 7 alleles/locus, H₀ = 0.500, H[subscript e] = 0.495; all P <0.001). Interestingly, one microsatellite locus (PPHO104) had anomalously high diversity (31 to 63 alleles) in both Hector's and Maui's dolphins and appears to be influenced by diversifying selection. The observed and expected heterozygosity, internal relatedness, and F[subscript IS] of Maui's dolphins all showed patterns consistent with a decline of the subspecies, although none differed significantly over the short time interval between the two datasets collected in 2001-07 and 2010-11. The lack of significant decline in any of the parameters analyzed for Maui's dolphins is not surprising given the low power to detect a low to moderate decline over the short interval (<1 generation) between the two sampling periods. Compared to minimum viable effective population sizes proposed to guide management decisions, the Maui's dolphin has declined below the recommended threshold of N[subscript e] = 50, recently increased to N[subscript e] ≥100, thought to be necessary to avoid inbreeding depression in the short term (5 generations, ~65.2 years for Maui's and Hector's dolphins). Additionally, both the Maui's dolphin and Cloudy Bay Hector's dolphin populations are below the recommended threshold of N[subscript e] = 500, recently increased to N[subscript e] ≥1000, thought to be necessary to preserve long-term evolutionary potential. This is less of a concern for the Cloudy Bay Hector's population, which is thought to maintain gene flow with neighboring populations. However, for the small, isolated Maui's dolphin population, inbreeding depression is likely to be an increasing concern. Furthermore, each Maui's dolphin individual holds a disproportionate amount of the total genetic variation of the subspecies and would represent a disproportionately large demographic and genetic loss if it died before realizing its reproductive potential in the population. There is, however, potential for genetic restoration by interbreeding with Hector's dolphins, as genetic monitoring of Maui's dolphins revealed the first contemporary dispersal of four (two living females, one dead female, one dead male) Hector's dolphins into the Maui's dolphin distribution. Two Hector's dolphins (one dead female neonate, one living male) were also sampled along the North Island's southwest coast, outside the presumed range of either subspecies. Together, these records provide evidence of long-distance dispersal by Hector's dolphins (≥400 km) and the possibility of an unsampled Hector's dolphin population along the southwest coast of the North Island or northern South Island. These results highlight the value of genetic monitoring for subspecies lacking distinctive physical appearances, as such discoveries are not detected by other means but have important conservation implications. Although the Maui's dolphin is critically endangered, it is not necessarily doomed to extinction. The subspecies appears to be maintaining an equal sex ratio and connectivity within its remnant range, and the highly diverse locus PPHO104 could potentially offer clues to an inbreeding avoidance mechanism. If Maui's dolphins interbreed with the recently identified Hector's dolphin immigrants, it could provide genetic restoration, enhancing chances of long-term survival of the Maui's dolphin. Continued genetic monitoring and examination of recovered carcasses for phenotypic signs of inbreeding are important for gauging genetic threats to the survival of Maui's dolphins, as well as determining if any Hector's dolphin populations appear to be declining toward the critically endangered state of the Maui's dolphin. The results of this work contributed to the decision by the New Zealand Department of Conservation and Ministry for Primary Industries to conduct an updated risk assessment for Maui's dolphins and accelerate the review of the Maui's Dolphin Threat Management Plan. Consequently, commercial and recreational set net restrictions were extended slightly to reduce entanglement risk to Maui's dolphins utilizing the southern part of their distribution, as well as any Hector's dolphins that disperse north into that area. The results related to the population of Hector's dolphins in Cloudy Bay provide information that will contribute to the upcoming review of the Hector's dolphin component of the Threat Management Plan. Close

• 4876. [Article] Ecological effects of post-wildfire management activities (salvage-logging and grass-seeding) on vegetation composition, diversity, biomass, and growth and survival of Pinus ponderosa and Purshia tridentata

  Salvage-logging and artificial seeding of grass following wildfire are common practices in coniferous forests of the western United States, yet few studies have quantified the ecological effects of these ...

  Citation × Citation Citation Abstract Copy Title: Ecological effects of post-wildfire management activities (salvage-logging and grass-seeding) on vegetation composition, diversity, biomass, and growth and survival of Pinus ponderosa and Purshia tridentata Author: Sexton, Timothy Ogden Salvage-logging and artificial seeding of grass following wildfire are common practices in coniferous forests of the western United States, yet few studies have quantified the ecological effects of these post-fire activities. The effects of post-wildfire salvage-logging and grass-seeding on vegetation composition, aboveground biomass, and growth and survival of Pinus ponderosa and Purshia tridentata were quantified on the area burned by the 1992 Lone Pine Fire, Winema National Forest in the Klamath Basin, Oregon. Prior to the fire, the area was dominated by uneven-aged stands of Pinus ponderosa with Purshia tridentata and Stipa occidentalis in the understory. The fire was a stand-replacement disturbance, where the majority of trees, the herbaceous component, and crowns of understory shrubs were killed. Salvage logging resulted in a significant decrease in understory biomass, species richness, species diversity, and growth and survival of P. ponderosa and P. tridentata. In addition plant community composition was shifted from native forb dominance to grass dominance. In 1993, the understory biomass of salvage-logged sites was 38% of the aboveground biomass produced on nonsalvaged sites (322 kg ha^-1 vs 843 kg ha^-1). In 1994, salvage-logged sites produced only 27% of the biomass produced on nonsalvaged sites (402 kg ha^-1 vs 1468 kg ha^-1). Salvage-logging reduced species richness, species diversity, and altered species composition. The first and second years following logging, species richness was reduced by 13% (20 versus 23), and 30% (15 versus 22), respectively. In 1993 and 1994, native forb frequency on nonsalvaged sites was 80% and 77% respectively, while salvage-logged sites recorded 68% and 31% respectively. Conversely, graminoid frequency was significantly higher on salvage-logged sites. In 1994, native graminoid frequency was 35% in nonsalvaged sites and > 61% on salvage-logged sites. During the first two years following salvage-logging, mean height growth of naturally-regenerated Pinus ponderosa was significantly lower on salvaged sites (9.4 versus 7.8 cm yr^-1) as was density of natural Purshia tridentata seedlings (313 versus 530 seedlings ha^-1). Survival in salvage-logged treatments was 22% lower for planted Purshia tridentata seedlings (57% versus 45%). Height growth in salvage-logged treatments was 16% lower for planted Pinus ponderosa (4.4 versus 3.7 cm). Total aboveground biomass on nonsalvaged burned sites (controls) averaged 843 kg ha^-1 in 1993, and 1473 kg ha^-1 in 1994. In 1993, the first year following the fire, sites seeded to Secale cereale produced = 1995 kg ha^-1 total aboveground biomass; most of the biomass was S. cereale. Those seeded sites produced 89% less native forb biomass than controls (82 versus 780 kg ha^-1), and = 80% less native forb biomass than areas seeded to the native grasses Sitanion hystrix and Festuca idahoensis. In 1994, the second post-fire year, biomass on sites seeded to S. cereale was >1653 kg ha^-1, dominated by S. cereale. In 1994, no differences in total aboveground biomass were detected between S. cereale, F idahoensis, S. hystrix, and the Control, indicating that the erosion reducing benefits of grass-seeding did not last for more than one year. Treatments seeded with S. cereale produced 58% less native forb biomass than controls (350 versus 825 kg ha^-1). Seeding S. cereale also reduced by 69% the biomass ofStipa occidentalis, the most common native grass in the area (311 kg ha^-1 on control sites versus 96 kg ha^-1 on S. cereale sites). In 1993 and 1994, fewer species were recorded on S. cereale sites than on all other sites. Grass-seeded sites exhibited no significant differences in frequency of noxious weeds in either of the two years following wildfire and seeding, rejecting the hypothesis that grass-seeding reduces noxious weeds. However, native graminoid frequency was reduced on all grass-seeded treatments. There were no significant differences between grass-seeding treatments in height and density of natural P. ponderosa and P. tridentata seedlings or growth and survival of planted P. ponderosa and P. tridentata seedlings. While salvage-logging provides an economic benefit to local communities through the extraction of commercially valuable timber, there is an ecological cost. Reduction in ecosystem structure, production, species richness, diversity and alterations in species composition are important ecological consequences of salvage logging. In addition, this study demonstrates that salvage-logging retards the re-establishment and early growth of Pinus ponderosa and Purshia tridentata, two important wildfire restoration priorities. Grass-seeding also results in significant ecological changes. Alterations in ecosystem production, species richness, diversity and species composition are important ecological consequences of grass-seeding. Managers should consider these long-term influences on ecosystem composition and structure when faced with decisions concerning post-fire rehabilitation and management. Title: Ecological effects of post-wildfire management activities (salvage-logging and grass-seeding) on vegetation composition, diversity, biomass, and growth and survival of Pinus ponderosa and Purshia tridentata Author: Sexton, Timothy Ogden Salvage-logging and artificial seeding of grass following wildfire are common practices in coniferous forests of the western United States, yet few studies have quantified the ecological effects of these post-fire activities. The effects of post-wildfire salvage-logging and grass-seeding on vegetation composition, aboveground biomass, and growth and survival of Pinus ponderosa and Purshia tridentata were quantified on the area burned by the 1992 Lone Pine Fire, Winema National Forest in the Klamath Basin, Oregon. Prior to the fire, the area was dominated by uneven-aged stands of Pinus ponderosa with Purshia tridentata and Stipa occidentalis in the understory. The fire was a stand-replacement disturbance, where the majority of trees, the herbaceous component, and crowns of understory shrubs were killed. Salvage logging resulted in a significant decrease in understory biomass, species richness, species diversity, and growth and survival of P. ponderosa and P. tridentata. In addition plant community composition was shifted from native forb dominance to grass dominance. In 1993, the understory biomass of salvage-logged sites was 38% of the aboveground biomass produced on nonsalvaged sites (322 kg ha^-1 vs 843 kg ha^-1). In 1994, salvage-logged sites produced only 27% of the biomass produced on nonsalvaged sites (402 kg ha^-1 vs 1468 kg ha^-1). Salvage-logging reduced species richness, species diversity, and altered species composition. The first and second years following logging, species richness was reduced by 13% (20 versus 23), and 30% (15 versus 22), respectively. In 1993 and 1994, native forb frequency on nonsalvaged sites was 80% and 77% respectively, while salvage-logged sites recorded 68% and 31% respectively. Conversely, graminoid frequency was significantly higher on salvage-logged sites. In 1994, native graminoid frequency was 35% in nonsalvaged sites and > 61% on salvage-logged sites. During the first two years following salvage-logging, mean height growth of naturally-regenerated Pinus ponderosa was significantly lower on salvaged sites (9.4 versus 7.8 cm yr^-1) as was density of natural Purshia tridentata seedlings (313 versus 530 seedlings ha^-1). Survival in salvage-logged treatments was 22% lower for planted Purshia tridentata seedlings (57% versus 45%). Height growth in salvage-logged treatments was 16% lower for planted Pinus ponderosa (4.4 versus 3.7 cm). Total aboveground biomass on nonsalvaged burned sites (controls) averaged 843 kg ha^-1 in 1993, and 1473 kg ha^-1 in 1994. In 1993, the first year following the fire, sites seeded to Secale cereale produced = 1995 kg ha^-1 total aboveground biomass; most of the biomass was S. cereale. Those seeded sites produced 89% less native forb biomass than controls (82 versus 780 kg ha^-1), and = 80% less native forb biomass than areas seeded to the native grasses Sitanion hystrix and Festuca idahoensis. In 1994, the second post-fire year, biomass on sites seeded to S. cereale was >1653 kg ha^-1, dominated by S. cereale. In 1994, no differences in total aboveground biomass were detected between S. cereale, F idahoensis, S. hystrix, and the Control, indicating that the erosion reducing benefits of grass-seeding did not last for more than one year. Treatments seeded with S. cereale produced 58% less native forb biomass than controls (350 versus 825 kg ha^-1). Seeding S. cereale also reduced by 69% the biomass ofStipa occidentalis, the most common native grass in the area (311 kg ha^-1 on control sites versus 96 kg ha^-1 on S. cereale sites). In 1993 and 1994, fewer species were recorded on S. cereale sites than on all other sites. Grass-seeded sites exhibited no significant differences in frequency of noxious weeds in either of the two years following wildfire and seeding, rejecting the hypothesis that grass-seeding reduces noxious weeds. However, native graminoid frequency was reduced on all grass-seeded treatments. There were no significant differences between grass-seeding treatments in height and density of natural P. ponderosa and P. tridentata seedlings or growth and survival of planted P. ponderosa and P. tridentata seedlings. While salvage-logging provides an economic benefit to local communities through the extraction of commercially valuable timber, there is an ecological cost. Reduction in ecosystem structure, production, species richness, diversity and alterations in species composition are important ecological consequences of salvage logging. In addition, this study demonstrates that salvage-logging retards the re-establishment and early growth of Pinus ponderosa and Purshia tridentata, two important wildfire restoration priorities. Grass-seeding also results in significant ecological changes. Alterations in ecosystem production, species richness, diversity and species composition are important ecological consequences of grass-seeding. Managers should consider these long-term influences on ecosystem composition and structure when faced with decisions concerning post-fire rehabilitation and management. Close

• 4877. [Article] Status and Distribution of Native Fishes in the Goose Lake Basin Information Reports number 2008-02

  Abstract -- This study describes the current distribution of the nine native fish species in the Oregon portion of the Goose Lake basin (Lake County): Goose Lake redband trout Oncorhynchus mykiss ssp., ...

  Citation × Citation Citation Abstract Copy Title: Status and Distribution of Native Fishes in the Goose Lake Basin Information Reports number 2008-02 Abstract -- This study describes the current distribution of the nine native fish species in the Oregon portion of the Goose Lake basin (Lake County): Goose Lake redband trout Oncorhynchus mykiss ssp., Goose Lake lamprey Entosphenus sp., Goose Lake tui chub Siphateles bicolor thalassinus, Goose Lake sucker Catostomus occidentalis lacusanserinus, Modoc sucker Catostomus microps, Pit-Klamath brook lamprey Entosphenus lethophagus, speckled dace Rhinichthys osculus, Pit roach Lavinia symmetricus mitrulus, and Pit sculpin Cottus pitensis. The Goose Lake basin is an endorheic, or topographically closed basin located in south central Oregon and northeastern California. The basin is within the usually closed northeastern extremity of the adjoining Sacramento River basin, astride the Oregon-California boundary. Although most of the lake lies in California, most of its valley and nearly two-thirds of the total drainage area (~722 sq. mi.) are in Oregon. The largest streams in the basin are Drews, Cottonwood, and Thomas Creeks. Annual precipitation averages about 36 cm per year (Phillips and van Denburgh 1971). Goose Lake overflowed briefly into the North Fork Pit River in 1868 and 1881, but storage and diversion of irrigation water has substantially reduced the inflow and future overflow is unlikely (USGS 1971). The lakebed was dry in the summers of 1926, 1929- 1934, and 1992. About half the basin is forestland, 20% is hay fields and pastureland, and 16% is shrub and rangeland. Currently, almost 35% of the inflow is diverted for irrigation (OWRD 1989). The Goose Lake basin is home to four endemic fish taxa: the Goose Lake redband trout, lamprey, sucker, and tui chub. Endemic fishes of the Goose Lake basin split their life histories between Goose Lake and its tributaries, as opposed to the five native but non-endemic species that primarily occupy stream habitats. Pit roach and all endemic fishes except Goose Lake tui chub are listed as a “species of concern” by the USFWS, a designation that implies there is concern about species viability, but not enough information is known to initiate a listing review for threatened or endangered status. The Modoc sucker was listed as a federally endangered species in 1985 (USFWS 1985). No formal recovery plan was required due to an existing “Action Plan for the Recovery of the Modoc Sucker” (USFWS 1984). Most of the recovery actions outlined in the action plan were either completed or are no longer relevant (Stewart Reid, Western Fishes, personal communication). However, actions 26 and 27 pertaining to range expansion remain incomplete. Action 26 suggests reclassification to threatened upon establishment of safe populations (for 3-5 years) throughout the Rush and Turner Creek watersheds in the Pit River basin. Action 27 suggests delisting upon establishing safe populations in two other historic streams. At the time of listing, the historic range of Modoc sucker was thought to have included only two small tributaries of the Pit River in Modoc and Lassen Counties, Ash and Turner Creeks (USFWS 1985). Therefore, a major recovery goal was to expand the species’ range with additional populations (USFWS 1984). In 2001, reexamination of historical documents and museum specimens established that Modoc suckers had also historically occupied Thomas Creek in the Goose Lake basin. Field collections in 2001, with subsequent morphological and genetic analysis, confirmed that the population was still present in Thomas Creek (Stewart Reid, Western Fishes, personal communication); however, the broader range of Modoc sucker in the Goose Lake watershed was not known. In 1995, the Goose Lake Fishes Working Group drafted a conservation plan for “prelisting” recovery of all native fish in response to severe drought and habitat degradation (GLFWG 1995). The Aquatic Inventories Project of the Oregon Department of Fish and Wildlife (ODFW) conducted habitat and fish distribution surveys (1991-1995) to obtain baseline information to help inform recovery efforts (ODFW, unpublished data). Since then, field work to monitor the distribution and abundance of Goose Lake fishes has been limited and sporadic, targeting only Goose Lake redband trout and Modoc sucker (Dambacher 2001; Reid 2007). No comprehensive follow up work has been conducted to evaluate fish response to climatic conditions, habitat restoration projects, and continued irrigation activities. ODFW recently drafted a status review of native fish of Oregon (ODFW 2005). Except for redband trout, Goose Lake fishes were not included in the status review due to a lack of new information since the previous status review in 1995 (Kostow et al. 1995). Further, the review of Goose Lake redband trout was limited by a lack of long-term data series. The first objective of this study was to document the current distribution of native fishes in Oregon’s portion of the Goose Lake basin and assess changes in distribution that may have occurred since the last surveys were conducted 12 years ago. The second objective was to provide new information about the distribution of Modoc suckers within the basin. The third objective was to determine relative abundance and age-class diversity of native fishes at randomly selected sample sites. All objectives were addressed throughout the potential riverine distribution of fish in the Oregon portion of the Goose Lake basin. Information gathered in this study is critical to effective conservation and management of each species and its habitat. In addition, this report describes the distribution and relative abundance of nonnative fishes (fathead minnow (Pimephales promelas), brown bullhead (Ameiurus nebulosus), white crappie (Pomoxis annularis), yellow perch (Perca flavescens), pumpkinseed (Lepomis gibbosus), and brook trout (Salvelinus fontinalis)) in the basin. Unlike prior efforts, this study used a statisticallybased design to select sample points with the aim of achieving a representative sample across the Oregon portion of the Goose Lake watershed. Additionally, a wide array of fish sampling gear was employed to maximize our ability to capture all fish species present across the diversity of habitat types encountered. Title: Status and Distribution of Native Fishes in the Goose Lake Basin Information Reports number 2008-02 Abstract -- This study describes the current distribution of the nine native fish species in the Oregon portion of the Goose Lake basin (Lake County): Goose Lake redband trout Oncorhynchus mykiss ssp., Goose Lake lamprey Entosphenus sp., Goose Lake tui chub Siphateles bicolor thalassinus, Goose Lake sucker Catostomus occidentalis lacusanserinus, Modoc sucker Catostomus microps, Pit-Klamath brook lamprey Entosphenus lethophagus, speckled dace Rhinichthys osculus, Pit roach Lavinia symmetricus mitrulus, and Pit sculpin Cottus pitensis. The Goose Lake basin is an endorheic, or topographically closed basin located in south central Oregon and northeastern California. The basin is within the usually closed northeastern extremity of the adjoining Sacramento River basin, astride the Oregon-California boundary. Although most of the lake lies in California, most of its valley and nearly two-thirds of the total drainage area (~722 sq. mi.) are in Oregon. The largest streams in the basin are Drews, Cottonwood, and Thomas Creeks. Annual precipitation averages about 36 cm per year (Phillips and van Denburgh 1971). Goose Lake overflowed briefly into the North Fork Pit River in 1868 and 1881, but storage and diversion of irrigation water has substantially reduced the inflow and future overflow is unlikely (USGS 1971). The lakebed was dry in the summers of 1926, 1929- 1934, and 1992. About half the basin is forestland, 20% is hay fields and pastureland, and 16% is shrub and rangeland. Currently, almost 35% of the inflow is diverted for irrigation (OWRD 1989). The Goose Lake basin is home to four endemic fish taxa: the Goose Lake redband trout, lamprey, sucker, and tui chub. Endemic fishes of the Goose Lake basin split their life histories between Goose Lake and its tributaries, as opposed to the five native but non-endemic species that primarily occupy stream habitats. Pit roach and all endemic fishes except Goose Lake tui chub are listed as a “species of concern” by the USFWS, a designation that implies there is concern about species viability, but not enough information is known to initiate a listing review for threatened or endangered status. The Modoc sucker was listed as a federally endangered species in 1985 (USFWS 1985). No formal recovery plan was required due to an existing “Action Plan for the Recovery of the Modoc Sucker” (USFWS 1984). Most of the recovery actions outlined in the action plan were either completed or are no longer relevant (Stewart Reid, Western Fishes, personal communication). However, actions 26 and 27 pertaining to range expansion remain incomplete. Action 26 suggests reclassification to threatened upon establishment of safe populations (for 3-5 years) throughout the Rush and Turner Creek watersheds in the Pit River basin. Action 27 suggests delisting upon establishing safe populations in two other historic streams. At the time of listing, the historic range of Modoc sucker was thought to have included only two small tributaries of the Pit River in Modoc and Lassen Counties, Ash and Turner Creeks (USFWS 1985). Therefore, a major recovery goal was to expand the species’ range with additional populations (USFWS 1984). In 2001, reexamination of historical documents and museum specimens established that Modoc suckers had also historically occupied Thomas Creek in the Goose Lake basin. Field collections in 2001, with subsequent morphological and genetic analysis, confirmed that the population was still present in Thomas Creek (Stewart Reid, Western Fishes, personal communication); however, the broader range of Modoc sucker in the Goose Lake watershed was not known. In 1995, the Goose Lake Fishes Working Group drafted a conservation plan for “prelisting” recovery of all native fish in response to severe drought and habitat degradation (GLFWG 1995). The Aquatic Inventories Project of the Oregon Department of Fish and Wildlife (ODFW) conducted habitat and fish distribution surveys (1991-1995) to obtain baseline information to help inform recovery efforts (ODFW, unpublished data). Since then, field work to monitor the distribution and abundance of Goose Lake fishes has been limited and sporadic, targeting only Goose Lake redband trout and Modoc sucker (Dambacher 2001; Reid 2007). No comprehensive follow up work has been conducted to evaluate fish response to climatic conditions, habitat restoration projects, and continued irrigation activities. ODFW recently drafted a status review of native fish of Oregon (ODFW 2005). Except for redband trout, Goose Lake fishes were not included in the status review due to a lack of new information since the previous status review in 1995 (Kostow et al. 1995). Further, the review of Goose Lake redband trout was limited by a lack of long-term data series. The first objective of this study was to document the current distribution of native fishes in Oregon’s portion of the Goose Lake basin and assess changes in distribution that may have occurred since the last surveys were conducted 12 years ago. The second objective was to provide new information about the distribution of Modoc suckers within the basin. The third objective was to determine relative abundance and age-class diversity of native fishes at randomly selected sample sites. All objectives were addressed throughout the potential riverine distribution of fish in the Oregon portion of the Goose Lake basin. Information gathered in this study is critical to effective conservation and management of each species and its habitat. In addition, this report describes the distribution and relative abundance of nonnative fishes (fathead minnow (Pimephales promelas), brown bullhead (Ameiurus nebulosus), white crappie (Pomoxis annularis), yellow perch (Perca flavescens), pumpkinseed (Lepomis gibbosus), and brook trout (Salvelinus fontinalis)) in the basin. Unlike prior efforts, this study used a statisticallybased design to select sample points with the aim of achieving a representative sample across the Oregon portion of the Goose Lake watershed. Additionally, a wide array of fish sampling gear was employed to maximize our ability to capture all fish species present across the diversity of habitat types encountered. Close

• 4878. [Article] Status, Distribution, and Life History Investigations of Warner Suckers, 2006-2010 Information Reports number 2011-02

  Abstract -- The Warner sucker Catostomus warnerensis is endemic to the Warner Valley, a subbasin of the Great Basin in southeastern Oregon and northwestern Nevada. This species was historically abundant ...

  Citation × Citation Citation Abstract Copy Title: Status, Distribution, and Life History Investigations of Warner Suckers, 2006-2010 Information Reports number 2011-02 Abstract -- The Warner sucker Catostomus warnerensis is endemic to the Warner Valley, a subbasin of the Great Basin in southeastern Oregon and northwestern Nevada. This species was historically abundant (Snyder 1908) and its historical range includes three permanent lakes (Hart, Crump, and Pelican), several ephemeral lakes, a network of sloughs and diversion canals, and three major tributary drainages (Honey, Deep, and Twentymile creeks). Warner sucker abundance and distribution has declined over the past century and it was federally listed as threatened in 1985 due to habitat fragmentation and threats posed by the proliferation of piscivorous non-native game fishes (U.S. Fish and Wildlife Service 1985). The Warner Valley is a northeast-southwest trending endorheic basin that extends approximately 90 km (Figure 1). The elevation of the valley floor is approximately 1,370 m and the basin is bound by fault block escarpments, the Warner Rim on the west and Hart Mountain and Poker Jim Ridge on the east. The Warner basin was formed during the middle Tertiary and late Quaternary geologic periods as a result of volcanic and tectonic activity (Baldwin 1974). Abundant precipitation during the Pleistocene Epoch resulted in the formation of Pluvial Lake Warner (Hubbs and Miller 1948). At its maximum extent approximately 11,000 years ago, the lake reached approximately 100 m in depth and 1,300 km2 in area (Snyder et al. 1964; Weide 1975). The Warner sucker inhabits the lakes and low gradient stream reaches of the Warner Valley. The metapopulation of Warner suckers is comprised of two life history forms: lake and stream morphs. The lake suckers display a lacustrine-adfluvial pattern in which they spend most of the year in the lake and spawn in the streams. However, when upstream migration is hindered by low stream flows during drought years or by irrigation diversion dams, lake suckers may spawn in nearshore areas of the lakes (White et al. 1990). Large lake-dwelling populations of introduced fishes in the lakes likely reduce sucker recruitment by predation on young suckers (U.S. Fish and Wildlife Service 1998). Periodic lake desiccation also threatens the lake suckers. The stream suckers display a fluvial life-history pattern and spawn in the three major tributary drainages (Honey, Deep, and Twentymile Creeks). Threats specific to the stream form include water withdrawals for irrigation and impacts from grazing. Stream suckers recolonized the lakes after past drying events (mid-1930’s and early-1990’s). The Recovery Plan for the Threatened and Rare Native Fishes of the Warner Basin and Alkali Subbasin (U.S. Fish and Wildlife Service 1998) sets three recovery criteria for delisting the species. These criteria require that: (1) a self-sustaining metapopulation is distributed throughout the drainages of Twentymile Creek, Honey Creek, and below the falls on Deep Creek, and in Pelican, Crump, and Hart Lakes; (2) passage is restored within and among these drainages so that individual populations of Warner suckers can function as a metapopulation; and (3) no threats exist that would likely threaten the survival of the species over a significant portion of its range. The Oregon Department of Fish and Wildlife’s (ODFW’s) Native Fish Investigations Project conducted investigations from 2006 through 2010 to describe the conservation (recovery) status of Warner suckers. The objectives of our investigations were to: 1) describe the current distribution of suckers in the Warner subbasin, 2) estimate their abundance in the lakes and streams, 3) collect life history information, and 4) describe the primary factors that currently limit the sucker’s ability to maintain a functioning metapopulation, including connectivity/fragmentation of habitats and factors affecting successful recruitment in the lake and stream environments. Previous similar studies were conducted in 1990, 1991, 1994, 1995, 1996, 1997, and 2001 (White et al. 1990; White et al. 1991; Allen et al. 1994; Allen et al. 1995; Allen et al. 1996; Bosse et al. 1997; Hartzell et al. 2001). We addressed these objectives by implementing the following tasks: 1) conducting surveys in Hart and Crump Lakes to describe the distribution and quantify the abundance of Warner suckers, search for evidence of recent recruitment, estimate sucker abundance relative to nonnative fish abundance, and describe certain life history characteristics, 2) tagging suckers with Passive Integrated Transponder (PIT) tags in the lakes and tributaries to estimate growth rates and describe seasonal movements, 3) radio tracking suckers in the lakes and tributaries to describe seasonal movements, 4) fishing screw traps in Warner basin tributaries to monitor downstream movements, 5) operating a trap at a fish ladder on a Warner tributary to assess upstream passage success, 6) conducting surveys in Warner basin tributaries to describe the current distribution of stream resident populations of Warner suckers and to quantify their abundance, 7) describing associations between the distribution of suckers and habitat variables in Twentymile Creek, 8) trapping larval suckers in the tributaries to describe the relative abundance and timing of larval movements, 9) describing life history parameters including growth rates, length frequency distributions, length at maturity, and weight-length relationships, 10) evaluating a nonlethal ageing technique, 11) describing the distribution and abundance of the Warner suckers at Summer Lake Wildlife Management area, where a self-sustaining population became established after fish salvage from Hart Lake during the 1992 drought, and 12) collecting tissue samples for future genetic analyses. This report compiles the results of this work, synthesizes and interprets findings relative to the conservation status of the species, and recommends future studies. Title: Status, Distribution, and Life History Investigations of Warner Suckers, 2006-2010 Information Reports number 2011-02 Abstract -- The Warner sucker Catostomus warnerensis is endemic to the Warner Valley, a subbasin of the Great Basin in southeastern Oregon and northwestern Nevada. This species was historically abundant (Snyder 1908) and its historical range includes three permanent lakes (Hart, Crump, and Pelican), several ephemeral lakes, a network of sloughs and diversion canals, and three major tributary drainages (Honey, Deep, and Twentymile creeks). Warner sucker abundance and distribution has declined over the past century and it was federally listed as threatened in 1985 due to habitat fragmentation and threats posed by the proliferation of piscivorous non-native game fishes (U.S. Fish and Wildlife Service 1985). The Warner Valley is a northeast-southwest trending endorheic basin that extends approximately 90 km (Figure 1). The elevation of the valley floor is approximately 1,370 m and the basin is bound by fault block escarpments, the Warner Rim on the west and Hart Mountain and Poker Jim Ridge on the east. The Warner basin was formed during the middle Tertiary and late Quaternary geologic periods as a result of volcanic and tectonic activity (Baldwin 1974). Abundant precipitation during the Pleistocene Epoch resulted in the formation of Pluvial Lake Warner (Hubbs and Miller 1948). At its maximum extent approximately 11,000 years ago, the lake reached approximately 100 m in depth and 1,300 km2 in area (Snyder et al. 1964; Weide 1975). The Warner sucker inhabits the lakes and low gradient stream reaches of the Warner Valley. The metapopulation of Warner suckers is comprised of two life history forms: lake and stream morphs. The lake suckers display a lacustrine-adfluvial pattern in which they spend most of the year in the lake and spawn in the streams. However, when upstream migration is hindered by low stream flows during drought years or by irrigation diversion dams, lake suckers may spawn in nearshore areas of the lakes (White et al. 1990). Large lake-dwelling populations of introduced fishes in the lakes likely reduce sucker recruitment by predation on young suckers (U.S. Fish and Wildlife Service 1998). Periodic lake desiccation also threatens the lake suckers. The stream suckers display a fluvial life-history pattern and spawn in the three major tributary drainages (Honey, Deep, and Twentymile Creeks). Threats specific to the stream form include water withdrawals for irrigation and impacts from grazing. Stream suckers recolonized the lakes after past drying events (mid-1930’s and early-1990’s). The Recovery Plan for the Threatened and Rare Native Fishes of the Warner Basin and Alkali Subbasin (U.S. Fish and Wildlife Service 1998) sets three recovery criteria for delisting the species. These criteria require that: (1) a self-sustaining metapopulation is distributed throughout the drainages of Twentymile Creek, Honey Creek, and below the falls on Deep Creek, and in Pelican, Crump, and Hart Lakes; (2) passage is restored within and among these drainages so that individual populations of Warner suckers can function as a metapopulation; and (3) no threats exist that would likely threaten the survival of the species over a significant portion of its range. The Oregon Department of Fish and Wildlife’s (ODFW’s) Native Fish Investigations Project conducted investigations from 2006 through 2010 to describe the conservation (recovery) status of Warner suckers. The objectives of our investigations were to: 1) describe the current distribution of suckers in the Warner subbasin, 2) estimate their abundance in the lakes and streams, 3) collect life history information, and 4) describe the primary factors that currently limit the sucker’s ability to maintain a functioning metapopulation, including connectivity/fragmentation of habitats and factors affecting successful recruitment in the lake and stream environments. Previous similar studies were conducted in 1990, 1991, 1994, 1995, 1996, 1997, and 2001 (White et al. 1990; White et al. 1991; Allen et al. 1994; Allen et al. 1995; Allen et al. 1996; Bosse et al. 1997; Hartzell et al. 2001). We addressed these objectives by implementing the following tasks: 1) conducting surveys in Hart and Crump Lakes to describe the distribution and quantify the abundance of Warner suckers, search for evidence of recent recruitment, estimate sucker abundance relative to nonnative fish abundance, and describe certain life history characteristics, 2) tagging suckers with Passive Integrated Transponder (PIT) tags in the lakes and tributaries to estimate growth rates and describe seasonal movements, 3) radio tracking suckers in the lakes and tributaries to describe seasonal movements, 4) fishing screw traps in Warner basin tributaries to monitor downstream movements, 5) operating a trap at a fish ladder on a Warner tributary to assess upstream passage success, 6) conducting surveys in Warner basin tributaries to describe the current distribution of stream resident populations of Warner suckers and to quantify their abundance, 7) describing associations between the distribution of suckers and habitat variables in Twentymile Creek, 8) trapping larval suckers in the tributaries to describe the relative abundance and timing of larval movements, 9) describing life history parameters including growth rates, length frequency distributions, length at maturity, and weight-length relationships, 10) evaluating a nonlethal ageing technique, 11) describing the distribution and abundance of the Warner suckers at Summer Lake Wildlife Management area, where a self-sustaining population became established after fish salvage from Hart Lake during the 1992 drought, and 12) collecting tissue samples for future genetic analyses. This report compiles the results of this work, synthesizes and interprets findings relative to the conservation status of the species, and recommends future studies. Close

• 4879. [Article] Forest macro-arthropods as potential indicators of ecosystem conditions in Western Idaho : an analysis of community composition, biological diversity, and community structure

  Adaptive ecosystem management is a new paradigm for managing federal forests which requires regular monitoring of ecosystem function and diversity to measure the effects of management. Managers need new ...

  Citation × Citation Citation Abstract Copy Title: Forest macro-arthropods as potential indicators of ecosystem conditions in Western Idaho : an analysis of community composition, biological diversity, and community structure Author: Ruby, Margaret E. Adaptive ecosystem management is a new paradigm for managing federal forests which requires regular monitoring of ecosystem function and diversity to measure the effects of management. Managers need new strategies and tools to help them assess their progress in maintaining healthy, productive and biologically diverse forests. Biomonitoring of select forest macro-arthropod species can provide useful information on the effects of management on forest biodiversity and ecosystem function. The purpose of this study was threefold: (1) to inventory the macro-arthropod community and important environmental variables in the Bear Creek and Indian Creek study area within the Payette National Forest (PNF) in Western Idaho; (2) to compare measures of community composition, diversity, and structure in forest macro-arthropod communities between patches of different sizes and treatment; and (3) to assist PNF managers in their ecosystem management efforts by providing principles to guide the use of macro-arthropods as indicators of changing forest conditions. Transects with pitfall traps were used to collect macro-arthropods at 22 sites in the Bear Creek and Indian Creek study area during the summer of 1994. Five forest patch types in Abies grandis habitat types were sampled. Intact forest patches of 100 or more hectares, and large patches of 50-100 hectares, ranged in age between 50 and 250 years old with multistoried structure. Small patches up to 10 hectares were remnants or fragments of formerly intact forest isolated by logging. A plantation patch was 15 years old with patchy understory and forb cover. Clearcut patches had little or no overstory, and variable understory, and forb layers. At each transect, soil samples were collected and six environmental descriptor variables were analyzed according to patch treatment and patch size. These site descriptors were: basal area (ft²/acre); percent canopy cover for the overstory, understory; and forb layers; litter depth (cm), and percent soil moisture content. Differences detected using an ANOVA and T-tests are discussed in the Results section. Arthropod community composition, diversity, and structure were described according to relative abundance, and four measures of diversity. They were also described by membership in seventeen orders and/or super-families; ten functional groups; two disperser classes (long or short distance); and three species indicator classes. A total of 5455 macro-arthropod individuals, representing 17 orders and/or super-families and 219 species were collected in the Bear Creek and Indian Creek study area. While macro-arthropod fauna relative abundance did not vary significantly by treatment (ANOVA p<0.3), it did vary significantly by patch size (ANOVA p<0.03). Fauna relative abundance was 35% greater in clearcut patches than in large patches (T-test p<0.09). Fauna relative abundance in small patches was twice that of intact (T-test p<0.03) and large (T-test p<0.02) patches. Taxonomic diversity (number of genera/taxa) of beetle, ant, and bug taxa differed significantly according to treatment type(each ANOVA p<0.05). For the top four taxa (beetles, ants, spiders, and bugs), taxonomic diversity was highest in the plantation and clearcut patches. Ants and bugs had their highest taxonomic diversity in the plantation patch (separate T-tests p<0.05) while the taxonomic diversity of beetles was highest in clear-cut patches (T-test p<0.05). Beetle and ant taxonomic diversity varied significantly by patch size (each ANOVA p<0.05). For beetles and bugs, small patches were twice as diverse as intact patches (separate T-tests p<0.04) and 1.5 times that of large patches. Ant diversity was similarly distributed amongst the patch sizes, with significant differences between small and intact and between small and large patches (separate T-tests p<0.05). Of the four species diversity measures employed, only two, [alpha] and JK1 (both measures of richness), were found to vary significantly by patch treatment and size. Evenness (E) and the Shannon Diversity Index (H') failed to detect differences in the majority of tests. Fauna [alpha] and JK1 differed significantly by treatment type (each ANOVA p<0.05). Richness in clearcut patches was nearly twice in intact and large patches, followed by plantation and large patches. Fauna [alpha] and JKl also differed significantly by patch size (each ANOVA p <0.001), with small patch fauna twice as rich as that in large and intact patches (separate T-tests p <0.01). Of the top four functional groups, predators were the most abundant and had the highest taxonomic diversity (number of genera/functional group), followed by herbivores, fungivores and parasites. Predators and herbivores showed increasing taxonomic diversity with decreasing patch size, from intact to large to small (ANOVA p< 0.05). Similarly, predators and herbivores exhibited increasing taxonomic diversity with increasing levels of management: from intact and large to plantation and clear-cuts (ANOVA p< 0.05). Predators and herbivores were most numerous in the managed and small patches. Fungivore taxonomic diversity was also highest in the small and managed patches, though neither patch size nor treatment differences were significant (ANOVA p<0.85). Parasite taxonomic diversity differed by patch size with highest generic diversity in the small patches (ANOVA p<0.l) and by treatment type with generic diversity highest in plantations and clearcuts followed in order by large and intact patches (ANOVA p<0.l). Twice as many genera were long distance dispersers as were short distance dispersers. Relative abundance of long distance dispersers varied significantly by patch treatment and patch size (each ANOVA p<0.0l). Long distance dispersers were most numerous in clear-cut patches, followed in order by plantation, small, large, and intact patches. Relative abundance of short distance dispersers was not significantly different between treatment types (ANOVA p<0.20) but was significantly different between patch sizes (ANOVA p<0.0l). Short distance dispersers were most numerous in small patches followed by plantation, large, and intact and least numerous in clearcut patches. An indicator species analysis of 121 Bear Creek and Indian Creek genera (Dufrene and Legendre 1997), revealed sub-groups of species with 75 to 100 percent "perfect indication" or affiliation for specific patch types. When intact and large patches were pooled and analyzed against all treated patches (plantation and clearcut patches), a list of 36 genera with 75 to 100 percent "perfect affiliation" for intact or large patches was produced (MRPP p<0.05). Small patches had 42 indicators with 75 to 100 percent "perfect indication" when compared with the pooled intact and large patches (MRPP p<0.l). Conclusions Macro-arthropod community composition, diversity and structure did vary, usually significantly, by patch treatment and size. Useful measures of generic diversity include richness estimators [alpha], [beta], and JK1. Examination of taxonomic diversity was also useful, especially for the more mobile arthropods. Pitfall traps provided copious data on the structure of the community in regards to predators and herbivores. Pitfalls, however, did not provide much information about the status of fungivores and parasites in the various different patches. Another trapping method such as the berlaise funnel, would likely provide more information about those functional groups which are likely operating at a finer scale of resolution than that tested by the pitfall trap. Employing both methods would provide a much better assessment of the community of arthropods living on the forest floor. The indicator species analysis program also provided very useful lists of species which are affiliated with particular patch conditions. Taken together, these measures could be adopted for use by forest managers to allow them to assess and monitor the effects of a management regime on the structure and composition of macro-arthropod communities as part of a comprehensive adaptive management plan. Title: Forest macro-arthropods as potential indicators of ecosystem conditions in Western Idaho : an analysis of community composition, biological diversity, and community structure Author: Ruby, Margaret E. Adaptive ecosystem management is a new paradigm for managing federal forests which requires regular monitoring of ecosystem function and diversity to measure the effects of management. Managers need new strategies and tools to help them assess their progress in maintaining healthy, productive and biologically diverse forests. Biomonitoring of select forest macro-arthropod species can provide useful information on the effects of management on forest biodiversity and ecosystem function. The purpose of this study was threefold: (1) to inventory the macro-arthropod community and important environmental variables in the Bear Creek and Indian Creek study area within the Payette National Forest (PNF) in Western Idaho; (2) to compare measures of community composition, diversity, and structure in forest macro-arthropod communities between patches of different sizes and treatment; and (3) to assist PNF managers in their ecosystem management efforts by providing principles to guide the use of macro-arthropods as indicators of changing forest conditions. Transects with pitfall traps were used to collect macro-arthropods at 22 sites in the Bear Creek and Indian Creek study area during the summer of 1994. Five forest patch types in Abies grandis habitat types were sampled. Intact forest patches of 100 or more hectares, and large patches of 50-100 hectares, ranged in age between 50 and 250 years old with multistoried structure. Small patches up to 10 hectares were remnants or fragments of formerly intact forest isolated by logging. A plantation patch was 15 years old with patchy understory and forb cover. Clearcut patches had little or no overstory, and variable understory, and forb layers. At each transect, soil samples were collected and six environmental descriptor variables were analyzed according to patch treatment and patch size. These site descriptors were: basal area (ft²/acre); percent canopy cover for the overstory, understory; and forb layers; litter depth (cm), and percent soil moisture content. Differences detected using an ANOVA and T-tests are discussed in the Results section. Arthropod community composition, diversity, and structure were described according to relative abundance, and four measures of diversity. They were also described by membership in seventeen orders and/or super-families; ten functional groups; two disperser classes (long or short distance); and three species indicator classes. A total of 5455 macro-arthropod individuals, representing 17 orders and/or super-families and 219 species were collected in the Bear Creek and Indian Creek study area. While macro-arthropod fauna relative abundance did not vary significantly by treatment (ANOVA p<0.3), it did vary significantly by patch size (ANOVA p<0.03). Fauna relative abundance was 35% greater in clearcut patches than in large patches (T-test p<0.09). Fauna relative abundance in small patches was twice that of intact (T-test p<0.03) and large (T-test p<0.02) patches. Taxonomic diversity (number of genera/taxa) of beetle, ant, and bug taxa differed significantly according to treatment type(each ANOVA p<0.05). For the top four taxa (beetles, ants, spiders, and bugs), taxonomic diversity was highest in the plantation and clearcut patches. Ants and bugs had their highest taxonomic diversity in the plantation patch (separate T-tests p<0.05) while the taxonomic diversity of beetles was highest in clear-cut patches (T-test p<0.05). Beetle and ant taxonomic diversity varied significantly by patch size (each ANOVA p<0.05). For beetles and bugs, small patches were twice as diverse as intact patches (separate T-tests p<0.04) and 1.5 times that of large patches. Ant diversity was similarly distributed amongst the patch sizes, with significant differences between small and intact and between small and large patches (separate T-tests p<0.05). Of the four species diversity measures employed, only two, [alpha] and JK1 (both measures of richness), were found to vary significantly by patch treatment and size. Evenness (E) and the Shannon Diversity Index (H') failed to detect differences in the majority of tests. Fauna [alpha] and JK1 differed significantly by treatment type (each ANOVA p<0.05). Richness in clearcut patches was nearly twice in intact and large patches, followed by plantation and large patches. Fauna [alpha] and JKl also differed significantly by patch size (each ANOVA p <0.001), with small patch fauna twice as rich as that in large and intact patches (separate T-tests p <0.01). Of the top four functional groups, predators were the most abundant and had the highest taxonomic diversity (number of genera/functional group), followed by herbivores, fungivores and parasites. Predators and herbivores showed increasing taxonomic diversity with decreasing patch size, from intact to large to small (ANOVA p< 0.05). Similarly, predators and herbivores exhibited increasing taxonomic diversity with increasing levels of management: from intact and large to plantation and clear-cuts (ANOVA p< 0.05). Predators and herbivores were most numerous in the managed and small patches. Fungivore taxonomic diversity was also highest in the small and managed patches, though neither patch size nor treatment differences were significant (ANOVA p<0.85). Parasite taxonomic diversity differed by patch size with highest generic diversity in the small patches (ANOVA p<0.l) and by treatment type with generic diversity highest in plantations and clearcuts followed in order by large and intact patches (ANOVA p<0.l). Twice as many genera were long distance dispersers as were short distance dispersers. Relative abundance of long distance dispersers varied significantly by patch treatment and patch size (each ANOVA p<0.0l). Long distance dispersers were most numerous in clear-cut patches, followed in order by plantation, small, large, and intact patches. Relative abundance of short distance dispersers was not significantly different between treatment types (ANOVA p<0.20) but was significantly different between patch sizes (ANOVA p<0.0l). Short distance dispersers were most numerous in small patches followed by plantation, large, and intact and least numerous in clearcut patches. An indicator species analysis of 121 Bear Creek and Indian Creek genera (Dufrene and Legendre 1997), revealed sub-groups of species with 75 to 100 percent "perfect indication" or affiliation for specific patch types. When intact and large patches were pooled and analyzed against all treated patches (plantation and clearcut patches), a list of 36 genera with 75 to 100 percent "perfect affiliation" for intact or large patches was produced (MRPP p<0.05). Small patches had 42 indicators with 75 to 100 percent "perfect indication" when compared with the pooled intact and large patches (MRPP p<0.l). Conclusions Macro-arthropod community composition, diversity and structure did vary, usually significantly, by patch treatment and size. Useful measures of generic diversity include richness estimators [alpha], [beta], and JK1. Examination of taxonomic diversity was also useful, especially for the more mobile arthropods. Pitfall traps provided copious data on the structure of the community in regards to predators and herbivores. Pitfalls, however, did not provide much information about the status of fungivores and parasites in the various different patches. Another trapping method such as the berlaise funnel, would likely provide more information about those functional groups which are likely operating at a finer scale of resolution than that tested by the pitfall trap. Employing both methods would provide a much better assessment of the community of arthropods living on the forest floor. The indicator species analysis program also provided very useful lists of species which are affiliated with particular patch conditions. Taken together, these measures could be adopted for use by forest managers to allow them to assess and monitor the effects of a management regime on the structure and composition of macro-arthropod communities as part of a comprehensive adaptive management plan. Close

• 4880. [Article] The geology of the Elk Mountain-Porter Ridge area, Clatsop County, northwest Oregon

  Eight Tertiary sedimentary and volcanic units crop out in the thesis area. From oldest to youngest they are the: Sager Creek formation (informal); Pittsburg Bluff Formation; Northrup Creek formation (informal); ...

  Citation × Citation Citation Abstract Copy Title: The geology of the Elk Mountain-Porter Ridge area, Clatsop County, northwest Oregon Author: Goalen, Jeffrey Scott Eight Tertiary sedimentary and volcanic units crop out in the thesis area. From oldest to youngest they are the: Sager Creek formation (informal); Pittsburg Bluff Formation; Northrup Creek formation (informal); Smuggler Cove formation (informal); Wickiup Mountain and Cannon Beach members (both informal) of the Astoria Formation; the Grande Ronde Basalt, and Frenchman Springs Member of the Wanapum Basalt, both of the Columbia River Basalt Group. Also, areally limited, unnamed sedimentary strata interbedded between flows of the Columbia River Basalt Group crop out in the study area. Quaternary deposits consist of alluvium, colluvium, and landslide debris. Rhythmically-bedded, foram-bearing, carbonaceous to micaceous mudstone and graded, fine-grained, feldspathic turbidite sandstone are the dominant lithologies of the upper Eocene (Refugian) Sager Creek formation. Plane-laminae and climbing ripple-laminae typical of Bouma b,c, and d intervals are common in the thin turbidite sandstone beds. Contemporaneous, rare, thick, sandstones may represent submarine feeder channels that supplied the more widespread, thinly-bedded overbank turbidite sandstones. Foraminiferal paleobathymetry indicates that deposition was in bathyal water depths. Sager Creek deposition was followed by a regression or offlap of the late Eocene to early Oligocene (Refugian) sea as indicated by the molluscan fossils and thick, bioturbated sandstone of the predominantly shallow-marine Pittsburg Bluff Formation. The lower part of the formation consists of outer shelf, glauconitic, fossiliferous sandstone and subordinate mudstone. Higher in the section are middle-shelf, fine-grained, bioturbated, tuffaceous, arkosic sandstones. These sandstones contain minor glauconite, wave- and storm-generated molluscan shell hash beds, carbonized wood fragments, and calcareous concretions. Deposition occurred in 20 - 50 m open-marine shelf water depths; however, the depositional environment may have shallowed to a bay-like setting (Moore, 1982, written communication). The upper part of the unit consists of thin- to medium-bedded, carbonaceous siltstone and mudstone with minor ashfall tuffs. Therefore, deposition of the Pittsburg Bluff Formation occurred as a shallowing-upward, then deepening depositional episode, punctuated by contemporaneous, intermittent eruption of nearby calc-alkaline western Cascade volcanoes. The deep-marine Oligocene to lower Miocene (Zemorrian to Saucesian) Northrup Creek formation is predominantly composed of thinly-laminated mudstone interbedded with thin, very fine-grained, graded, micaceous arkosic sandstone. Bouma c-d-e and a-b-e sequences are common in the turbidite sandstone; Bouma a-e intervals occur locally in sandstone/mudstone couplets. Paleocurrent indicators suggest that the predominant transport direction of these turbidite deposits was northeast to southwest. Abraded, carbonaceous plant debris and mica are abundant in the sandstone laminations. The upper part of the formation consists of thick, mollusk-bearing, moderately- to well-sorted, arkosic sandstone and minor polymict grit beds. A shallowing-upward, high-energy, shallow-marine shelf environment of deposition is indicated for the upper part of the formation. Contemporaneous with shelf and slope deposition of the Sager Creek, Pittsburg Bluff and Northrup Creek formations, the deep-water late Eocene to early Miocene Smuggler Cove formation was deposited as a distal or lateral correlative in a low-energy, outer shelf to slope setting. This foram-bearing unit consists of thick, bioturbated, bathyal, tuffaceous mudstone and siltstone with minor thin- to thick-bedded ashfall and current-reworked tuff. A marine onlap is indicated by the conformable relationship between the upper sandstone unit of the Northrup Creek formation and the overlying Smuggler Cove formation in the eastern part of the thesis area. The overlying lower to middle Miocene Astoria Formation contains two members in the thesis area: a high-energy, shallow shelf, fine- to medium-grained, fossiliferous, micaceous arkosic sandstone (Wickiup Mountain member), and an overlying, thinly-laminated, deepmarine mudstone (Cannon Beach member). Diatom floras indicate that a thermal "oceanographic irregularity" (water warmer than normal) may have occurred during deposition of the Cannon Beach member. At least six, and possibly eight, flows of the middle Miocene Grande Ronde Basalt (Columbia River Basalt Group) are present in the thesis area. Individual flows have been ascribed to the (N1?), R2, and N2 magnetozones. Geochemically, the flows consist of low MgO high Ti02, low MgO low Ti02, and high MgO subtypes. These subaerial to submarine flows are correlated to Mangan and others (1986) chemical subtypes 2D, 5C, 5A, and 4A of the Columbia Plateau-derived Grande Ronde Basalt of eastern Washington and eastern Oregon. The correlations are based on similarity of age, major element chemistry, stratigraphic position, and magnetic polarity. Field evidence suggests that thick, submarine pillow and breccia complexes generated sufficient pressure to autoinvasively inject into the Eocene to middle Miocene sedimentary strata of the Plympton/Porter ridge and Elk Mountain areas. This process apparently formed many randomly-oriented dike- and sill-like intrusions in the western part of the thesis area. However, in the eastern and central parts of the thesis area, three sub-parallel dikes (the Northrup, Beneke, and Fishhawk Falls dikes) extend along linear trends for tens of kilometers. This suggests that their emplacement was, in part, influenced by earlier or contemporaneous regional tectonism. At least two, and as many as five, flows of the Frenchman Springs Member of the Wanapum Basalt occur in the thesis area: one to two abundantly plagioclase-phyric Basalt of Ginkgo flow(s), and one to three Basalt of Sand Hollow flows (terminology after Beeson and Tolan, 1985). These flows consist of subaqueous pillow palagonite breccia and vesicular, columnar-jointed, subaerial basalt. No Frenchman Springs flows are invasive in the thesis area. Local middle Miocene sedimentary interbeds between flows of Grande Ronde and Frenchman Springs Basalt are lithologically and sedimentologically diverse. Common lithologies are fine- to mediumgrained, arkosic sandstone, coarse-grained basaltic sandstone, and structureless mudstone. Depositional environments represented by these strata are fluvial, marginal-marine, and shallow-marine. Thicknesses of individual interbeds range from 0.2 to 50 m. The thesis area is located on the northwest flank of the Oregon Coast Range anticline, adjacent to the Nehalem Arch. Large-scale northeast-trending oblique to strike-slip left-lateral faults coupled with northwest-trending oblique to strike-slip right-lateral faults dominate the structure of the area. These faults may have formed as conjugate shears (Riedel shear) caused by north-south compressive stress related to the oblique subduction of the Juan de Fuca Plate beneath the North American Plate. Five episodes of deformation are suggested by faults, dike orientations, unconformities, and other geological relationships within the thesis area: a late Eocene north-south compressional episode, an early Oligocene to early Miocene uplift, a middle Miocene northwest-southeast extensional episode, a post-middle Miocene to Pliocene (?) north-south compressional episode, and a north-south extensional event that occurred between the post-middle Miocene and Recent. Although crushed rock (for road and revetment construction) is currently the primary mineral resource within the thesis area, several potential fault traps on this northwest flank of the Nehalem Arch may contain significant reserves of natural gas. This conclusion is based upon field, laboratory, and subsurface (well) data. The Clark and Wilson sandstone of the middle to late Eocene Cowlitz Formation, the producing unit at the nearby Mist gas field, represents the most attractive target horizon. Additionally, the porous and permeable upper sandstone unit of the Northrup Creek formation could contain shallow hydrocarbon reserves beneath the northern part of the area. Title: The geology of the Elk Mountain-Porter Ridge area, Clatsop County, northwest Oregon Author: Goalen, Jeffrey Scott Eight Tertiary sedimentary and volcanic units crop out in the thesis area. From oldest to youngest they are the: Sager Creek formation (informal); Pittsburg Bluff Formation; Northrup Creek formation (informal); Smuggler Cove formation (informal); Wickiup Mountain and Cannon Beach members (both informal) of the Astoria Formation; the Grande Ronde Basalt, and Frenchman Springs Member of the Wanapum Basalt, both of the Columbia River Basalt Group. Also, areally limited, unnamed sedimentary strata interbedded between flows of the Columbia River Basalt Group crop out in the study area. Quaternary deposits consist of alluvium, colluvium, and landslide debris. Rhythmically-bedded, foram-bearing, carbonaceous to micaceous mudstone and graded, fine-grained, feldspathic turbidite sandstone are the dominant lithologies of the upper Eocene (Refugian) Sager Creek formation. Plane-laminae and climbing ripple-laminae typical of Bouma b,c, and d intervals are common in the thin turbidite sandstone beds. Contemporaneous, rare, thick, sandstones may represent submarine feeder channels that supplied the more widespread, thinly-bedded overbank turbidite sandstones. Foraminiferal paleobathymetry indicates that deposition was in bathyal water depths. Sager Creek deposition was followed by a regression or offlap of the late Eocene to early Oligocene (Refugian) sea as indicated by the molluscan fossils and thick, bioturbated sandstone of the predominantly shallow-marine Pittsburg Bluff Formation. The lower part of the formation consists of outer shelf, glauconitic, fossiliferous sandstone and subordinate mudstone. Higher in the section are middle-shelf, fine-grained, bioturbated, tuffaceous, arkosic sandstones. These sandstones contain minor glauconite, wave- and storm-generated molluscan shell hash beds, carbonized wood fragments, and calcareous concretions. Deposition occurred in 20 - 50 m open-marine shelf water depths; however, the depositional environment may have shallowed to a bay-like setting (Moore, 1982, written communication). The upper part of the unit consists of thin- to medium-bedded, carbonaceous siltstone and mudstone with minor ashfall tuffs. Therefore, deposition of the Pittsburg Bluff Formation occurred as a shallowing-upward, then deepening depositional episode, punctuated by contemporaneous, intermittent eruption of nearby calc-alkaline western Cascade volcanoes. The deep-marine Oligocene to lower Miocene (Zemorrian to Saucesian) Northrup Creek formation is predominantly composed of thinly-laminated mudstone interbedded with thin, very fine-grained, graded, micaceous arkosic sandstone. Bouma c-d-e and a-b-e sequences are common in the turbidite sandstone; Bouma a-e intervals occur locally in sandstone/mudstone couplets. Paleocurrent indicators suggest that the predominant transport direction of these turbidite deposits was northeast to southwest. Abraded, carbonaceous plant debris and mica are abundant in the sandstone laminations. The upper part of the formation consists of thick, mollusk-bearing, moderately- to well-sorted, arkosic sandstone and minor polymict grit beds. A shallowing-upward, high-energy, shallow-marine shelf environment of deposition is indicated for the upper part of the formation. Contemporaneous with shelf and slope deposition of the Sager Creek, Pittsburg Bluff and Northrup Creek formations, the deep-water late Eocene to early Miocene Smuggler Cove formation was deposited as a distal or lateral correlative in a low-energy, outer shelf to slope setting. This foram-bearing unit consists of thick, bioturbated, bathyal, tuffaceous mudstone and siltstone with minor thin- to thick-bedded ashfall and current-reworked tuff. A marine onlap is indicated by the conformable relationship between the upper sandstone unit of the Northrup Creek formation and the overlying Smuggler Cove formation in the eastern part of the thesis area. The overlying lower to middle Miocene Astoria Formation contains two members in the thesis area: a high-energy, shallow shelf, fine- to medium-grained, fossiliferous, micaceous arkosic sandstone (Wickiup Mountain member), and an overlying, thinly-laminated, deepmarine mudstone (Cannon Beach member). Diatom floras indicate that a thermal "oceanographic irregularity" (water warmer than normal) may have occurred during deposition of the Cannon Beach member. At least six, and possibly eight, flows of the middle Miocene Grande Ronde Basalt (Columbia River Basalt Group) are present in the thesis area. Individual flows have been ascribed to the (N1?), R2, and N2 magnetozones. Geochemically, the flows consist of low MgO high Ti02, low MgO low Ti02, and high MgO subtypes. These subaerial to submarine flows are correlated to Mangan and others (1986) chemical subtypes 2D, 5C, 5A, and 4A of the Columbia Plateau-derived Grande Ronde Basalt of eastern Washington and eastern Oregon. The correlations are based on similarity of age, major element chemistry, stratigraphic position, and magnetic polarity. Field evidence suggests that thick, submarine pillow and breccia complexes generated sufficient pressure to autoinvasively inject into the Eocene to middle Miocene sedimentary strata of the Plympton/Porter ridge and Elk Mountain areas. This process apparently formed many randomly-oriented dike- and sill-like intrusions in the western part of the thesis area. However, in the eastern and central parts of the thesis area, three sub-parallel dikes (the Northrup, Beneke, and Fishhawk Falls dikes) extend along linear trends for tens of kilometers. This suggests that their emplacement was, in part, influenced by earlier or contemporaneous regional tectonism. At least two, and as many as five, flows of the Frenchman Springs Member of the Wanapum Basalt occur in the thesis area: one to two abundantly plagioclase-phyric Basalt of Ginkgo flow(s), and one to three Basalt of Sand Hollow flows (terminology after Beeson and Tolan, 1985). These flows consist of subaqueous pillow palagonite breccia and vesicular, columnar-jointed, subaerial basalt. No Frenchman Springs flows are invasive in the thesis area. Local middle Miocene sedimentary interbeds between flows of Grande Ronde and Frenchman Springs Basalt are lithologically and sedimentologically diverse. Common lithologies are fine- to mediumgrained, arkosic sandstone, coarse-grained basaltic sandstone, and structureless mudstone. Depositional environments represented by these strata are fluvial, marginal-marine, and shallow-marine. Thicknesses of individual interbeds range from 0.2 to 50 m. The thesis area is located on the northwest flank of the Oregon Coast Range anticline, adjacent to the Nehalem Arch. Large-scale northeast-trending oblique to strike-slip left-lateral faults coupled with northwest-trending oblique to strike-slip right-lateral faults dominate the structure of the area. These faults may have formed as conjugate shears (Riedel shear) caused by north-south compressive stress related to the oblique subduction of the Juan de Fuca Plate beneath the North American Plate. Five episodes of deformation are suggested by faults, dike orientations, unconformities, and other geological relationships within the thesis area: a late Eocene north-south compressional episode, an early Oligocene to early Miocene uplift, a middle Miocene northwest-southeast extensional episode, a post-middle Miocene to Pliocene (?) north-south compressional episode, and a north-south extensional event that occurred between the post-middle Miocene and Recent. Although crushed rock (for road and revetment construction) is currently the primary mineral resource within the thesis area, several potential fault traps on this northwest flank of the Nehalem Arch may contain significant reserves of natural gas. This conclusion is based upon field, laboratory, and subsurface (well) data. The Clark and Wilson sandstone of the middle to late Eocene Cowlitz Formation, the producing unit at the nearby Mist gas field, represents the most attractive target horizon. Additionally, the porous and permeable upper sandstone unit of the Northrup Creek formation could contain shallow hydrocarbon reserves beneath the northern part of the area. Close