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• 21. [Article] EWEB’s Vision: Payments for Ecosystem Services through a Voluntary Incentives Program; An Innovative, Incentive-Based Approach for Preserving Water Quality in the McKenzie River Watershed

  EWEB envisions the development of an investment mechanism that makes payments for ecosystem services (PES) as a way to maintain and improve water quality within the McKenzie River Watershed, Eugene’s sole ...

  Citation × Citation Citation Abstract Copy Title: EWEB’s Vision: Payments for Ecosystem Services through a Voluntary Incentives Program; An Innovative, Incentive-Based Approach for Preserving Water Quality in the McKenzie River Watershed Author: Lurie, Sue, Toth, Nancy, Morgenstern, Karl, Duncan, Sally, Fishler, Hillary, Bennett, Drew EWEB envisions the development of an investment mechanism that makes payments for ecosystem services (PES) as a way to maintain and improve water quality within the McKenzie River Watershed, Eugene’s sole source of drinking water. The public name for this concept is the Voluntary Incentives Program. Under the envisioned Voluntary Incentives Program (VIP), EWEB will provide annual dividend payments to landowners for the valuable services their properties provide including water filtration and purification. These dividends recognize the value of the natural capital provided by these lands and their downstream benefits to the residents of Eugene. A fund with sustainable financing will be established to support the dividend payments and the infrastructure necessary to operate the Voluntary Incentives Program. Financing will come from a variety of sources but could be initially endowed through existing water funds (under existing rate structure) or another utility funding mechanism. Additional possible financing sources other than EWEB include corporations, a voter approved bond measure, development impact fees, and state and federal mitigation programs. EWEB will establish a stewardship boundary identifying riparian forests and floodplains that are eligible to enroll in the VIP. Participation is open to private landowners, local governments, and non-profit organizations that own land within the designated boundary. Based on EWEB’s preliminary analysis, an estimated 6,500 acres of riparian and floodplain areas along the McKenzie and major tributaries are eligible to enroll. Land within the stewardship boundary will need to meet a threshold in order to receive payments. This threshold will be determined by adapting existing riparian forest and wetland habitat standards and definitions from NRCS, USFS, Defenders of Wildlife and other entities to establish the criteria for participation in the VIP. Appropriate infrastructure is essential to implement and run the incentives program: critical tasks include managing the fund, making payments to VIP participants, assessing the quality of land enrolled, negotiating agreements, monitoring properties and verifying compliance, and educating and communicating with the community. The VIP will rely upon a coalition of existing organizations, which will form a watershed investment district (WID), to provide this critical infrastructure, without which successful implementation of the VIP is highly unlikely. Title: EWEB’s Vision: Payments for Ecosystem Services through a Voluntary Incentives Program; An Innovative, Incentive-Based Approach for Preserving Water Quality in the McKenzie River Watershed Author: Lurie, Sue, Toth, Nancy, Morgenstern, Karl, Duncan, Sally, Fishler, Hillary, Bennett, Drew EWEB envisions the development of an investment mechanism that makes payments for ecosystem services (PES) as a way to maintain and improve water quality within the McKenzie River Watershed, Eugene’s sole source of drinking water. The public name for this concept is the Voluntary Incentives Program. Under the envisioned Voluntary Incentives Program (VIP), EWEB will provide annual dividend payments to landowners for the valuable services their properties provide including water filtration and purification. These dividends recognize the value of the natural capital provided by these lands and their downstream benefits to the residents of Eugene. A fund with sustainable financing will be established to support the dividend payments and the infrastructure necessary to operate the Voluntary Incentives Program. Financing will come from a variety of sources but could be initially endowed through existing water funds (under existing rate structure) or another utility funding mechanism. Additional possible financing sources other than EWEB include corporations, a voter approved bond measure, development impact fees, and state and federal mitigation programs. EWEB will establish a stewardship boundary identifying riparian forests and floodplains that are eligible to enroll in the VIP. Participation is open to private landowners, local governments, and non-profit organizations that own land within the designated boundary. Based on EWEB’s preliminary analysis, an estimated 6,500 acres of riparian and floodplain areas along the McKenzie and major tributaries are eligible to enroll. Land within the stewardship boundary will need to meet a threshold in order to receive payments. This threshold will be determined by adapting existing riparian forest and wetland habitat standards and definitions from NRCS, USFS, Defenders of Wildlife and other entities to establish the criteria for participation in the VIP. Appropriate infrastructure is essential to implement and run the incentives program: critical tasks include managing the fund, making payments to VIP participants, assessing the quality of land enrolled, negotiating agreements, monitoring properties and verifying compliance, and educating and communicating with the community. The VIP will rely upon a coalition of existing organizations, which will form a watershed investment district (WID), to provide this critical infrastructure, without which successful implementation of the VIP is highly unlikely. Close

• 22. [Article] Factors influencing spawning site choice by female Garibaldi, Hypsypops rubicundus (Pisces: Pomacentridae)

  The garibaldi, Hypsypops rubicundus, is a sexually monochromatic pomacentrid found in rocky subtidal areas of Southern and Baja California. During the spawning season, males attempt to attract females ...

  Citation × Citation Citation Abstract Copy Title: Factors influencing spawning site choice by female Garibaldi, Hypsypops rubicundus (Pisces: Pomacentridae) Author: Sikkel, Paul C. The garibaldi, Hypsypops rubicundus, is a sexually monochromatic pomacentrid found in rocky subtidal areas of Southern and Baja California. During the spawning season, males attempt to attract females to a nest of red algae located within individually defended territories. Females were observed to enter the nests of several males before selecting one for spawning and were significantly more likely to spawn in nests that contained eggs in the early stages of development than nests with no eggs or with late stage embryos. Among empty nests, the density of red turf algae and the number of nesting males surrounding the nest were inversely related to the amount of time elapsed until a nesting male acquired the first clutch of a brood. The percentage of long algae in the nest was positively related to this measure. These results demonstrate active female choice of spawning site by female garibaldi and suggest that certain characteristics of the male-defended nest influence this choice. Title: Factors influencing spawning site choice by female Garibaldi, Hypsypops rubicundus (Pisces: Pomacentridae) Author: Sikkel, Paul C. The garibaldi, Hypsypops rubicundus, is a sexually monochromatic pomacentrid found in rocky subtidal areas of Southern and Baja California. During the spawning season, males attempt to attract females to a nest of red algae located within individually defended territories. Females were observed to enter the nests of several males before selecting one for spawning and were significantly more likely to spawn in nests that contained eggs in the early stages of development than nests with no eggs or with late stage embryos. Among empty nests, the density of red turf algae and the number of nesting males surrounding the nest were inversely related to the amount of time elapsed until a nesting male acquired the first clutch of a brood. The percentage of long algae in the nest was positively related to this measure. These results demonstrate active female choice of spawning site by female garibaldi and suggest that certain characteristics of the male-defended nest influence this choice. Close

• 23. [Article] Effects of juvenile steelhead on juvenile chinook salmon behavior and physiology

  Experiments were designed to determine whether and how steelhead Oncorhynchus mykiss affect chinook salmon O. tshawytscha when the two species are confined together. In a behavioral experiment, we observed ...

  Citation × Citation Citation Abstract Copy Title: Effects of juvenile steelhead on juvenile chinook salmon behavior and physiology Author: Schreck, Carl B., Congleton, J. L., Kelsey, D. A., Davis, L. E. Experiments were designed to determine whether and how steelhead Oncorhynchus mykiss affect chinook salmon O. tshawytscha when the two species are confined together. In a behavioral experiment, we observed groups of juvenile chinook salmon and steelhead together and groups of chinook salmon alone to determine whether the steelhead were aggressive and their presence changed the behavior of chinook salmon. We also performed two runs of a physiological experiment to determine whether the addition of steelhead to tanks containing chinook salmon would stress the chinook salmon, as determined by a change in their plasma cortisol levels. Behavioral changes were observed in the chinook salmon when they were held with steelhead; they reduced their movements, darted less, were attacked up to 16 times as often, and were found less frequently in the shade than chinook salmon held without steelhead. Steelhead were found to establish territories and defend them with chases, charges, and nips. In their attempts to establish and defend territories, the steelhead attacked the chinook salmon as often as they attacked other steelhead, but the chinook salmon showed little aggression toward the steelhead. Cortisol concentrations were significantly higher for chinook salmon in tanks receiving steelhead than in tanks receiving additional chinook salmon or no additional fish after 2 h (in one of the two experimental runs) and after 32 h (in both experimental runs; combined data). These results suggest that confining steelhead and chinook salmon together, such as in raceways and barges in the Columbia River system and in other situations, is stressful to the chinook salmon. Title: Effects of juvenile steelhead on juvenile chinook salmon behavior and physiology Author: Schreck, Carl B., Congleton, J. L., Kelsey, D. A., Davis, L. E. Experiments were designed to determine whether and how steelhead Oncorhynchus mykiss affect chinook salmon O. tshawytscha when the two species are confined together. In a behavioral experiment, we observed groups of juvenile chinook salmon and steelhead together and groups of chinook salmon alone to determine whether the steelhead were aggressive and their presence changed the behavior of chinook salmon. We also performed two runs of a physiological experiment to determine whether the addition of steelhead to tanks containing chinook salmon would stress the chinook salmon, as determined by a change in their plasma cortisol levels. Behavioral changes were observed in the chinook salmon when they were held with steelhead; they reduced their movements, darted less, were attacked up to 16 times as often, and were found less frequently in the shade than chinook salmon held without steelhead. Steelhead were found to establish territories and defend them with chases, charges, and nips. In their attempts to establish and defend territories, the steelhead attacked the chinook salmon as often as they attacked other steelhead, but the chinook salmon showed little aggression toward the steelhead. Cortisol concentrations were significantly higher for chinook salmon in tanks receiving steelhead than in tanks receiving additional chinook salmon or no additional fish after 2 h (in one of the two experimental runs) and after 32 h (in both experimental runs; combined data). These results suggest that confining steelhead and chinook salmon together, such as in raceways and barges in the Columbia River system and in other situations, is stressful to the chinook salmon. Close

• 24. [Article] Effects of steelhead trout (Oncorhynchus mykiss) on chinook salmon (O. tshawytscha) behavior and physiology

  Three experiments were designed to determine if and how steelhead trout, Oncorhynchus mykiss, may affect chinook salmon, O. tshawytscha, when they are confined together as in a raceway or on a barge. We ...

  Citation × Citation Citation Abstract Copy Title: Effects of steelhead trout (Oncorhynchus mykiss) on chinook salmon (O. tshawytscha) behavior and physiology Author: Kelsey, Denise A. Three experiments were designed to determine if and how steelhead trout, Oncorhynchus mykiss, may affect chinook salmon, O. tshawytscha, when they are confined together as in a raceway or on a barge. We observed groups of chinook and steelhead together and groups of only chinook in a behavioral experiment to determine if steelhead are aggressive and if their presence changed the behavior of chinook. Two physiological experiments were completed to determine if the loading of steelhead on top of chinook and if the introduction of odor from rainbow trout (steelhead not available) caused a change in plasma cortisol levels in chinook. It was found that chinook showed characteristics of a schooling species, while steelhead exhibited territory holding characteristics. Behavioral changes in chinook were observed when steelhead were present. Chinook grouped with steelhead reduced their movements, darted less, were attacked up to 16 times more often, and were found less frequently in the shade than groups of only chinook. Steelhead were found to establish territories and defend them with chases, charges, and nips. In attempts to establish territories and defend them, steelhead attacked chinook as often as they attacked other steelhead even though chinook showed little aggression toward steelhead. In a physiological experiment, chinook experienced the loading of salmonids into their tank. Chinook had higher levels of plasma cortisol at 2 and 32 hours after the loading of steelhead than chinook that were loaded with chinook or controls (no loading). A second physiological experiment with odor showed that chinook that received rainbow odor and those that received chinook odor had similar levels of plasma cortisol. Cortisol levels (two hours after the odor was introduced) were higher in chinook receiving either of the scented waters than in those that did not receive any odor. In conclusion, all three experiments indicated that the presence of juvenile steelhead trout affect juvenile chinook salmon behavior and physiology. Title: Effects of steelhead trout (Oncorhynchus mykiss) on chinook salmon (O. tshawytscha) behavior and physiology Author: Kelsey, Denise A. Three experiments were designed to determine if and how steelhead trout, Oncorhynchus mykiss, may affect chinook salmon, O. tshawytscha, when they are confined together as in a raceway or on a barge. We observed groups of chinook and steelhead together and groups of only chinook in a behavioral experiment to determine if steelhead are aggressive and if their presence changed the behavior of chinook. Two physiological experiments were completed to determine if the loading of steelhead on top of chinook and if the introduction of odor from rainbow trout (steelhead not available) caused a change in plasma cortisol levels in chinook. It was found that chinook showed characteristics of a schooling species, while steelhead exhibited territory holding characteristics. Behavioral changes in chinook were observed when steelhead were present. Chinook grouped with steelhead reduced their movements, darted less, were attacked up to 16 times more often, and were found less frequently in the shade than groups of only chinook. Steelhead were found to establish territories and defend them with chases, charges, and nips. In attempts to establish territories and defend them, steelhead attacked chinook as often as they attacked other steelhead even though chinook showed little aggression toward steelhead. In a physiological experiment, chinook experienced the loading of salmonids into their tank. Chinook had higher levels of plasma cortisol at 2 and 32 hours after the loading of steelhead than chinook that were loaded with chinook or controls (no loading). A second physiological experiment with odor showed that chinook that received rainbow odor and those that received chinook odor had similar levels of plasma cortisol. Cortisol levels (two hours after the odor was introduced) were higher in chinook receiving either of the scented waters than in those that did not receive any odor. In conclusion, all three experiments indicated that the presence of juvenile steelhead trout affect juvenile chinook salmon behavior and physiology. Close

• 25. [Article] How Mega-Development is Killing the Nature and Culture of Uruguay

  43 pages. A thesis presented to the Department of Journalism and the Clark Honors College of the University of Oregon in partial fulfillment of the requirements for degree of Bachelor of Arts, Spring 2...

  Citation × Citation Citation Abstract Copy Title: How Mega-Development is Killing the Nature and Culture of Uruguay Author: Reid, Carolina Year: 2014 43 pages. A thesis presented to the Department of Journalism and the Clark Honors College of the University of Oregon in partial fulfillment of the requirements for degree of Bachelor of Arts, Spring 2014. Title: How Mega-Development is Killing the Nature and Culture of Uruguay Author: Reid, Carolina Year: 2014 43 pages. A thesis presented to the Department of Journalism and the Clark Honors College of the University of Oregon in partial fulfillment of the requirements for degree of Bachelor of Arts, Spring 2014. Close

• 26. [Article] Territorial behavior in juvenile steelhead trout (Oncorhynchus mykiss) : how redside shiner (Richardsonius balteatus) influence intraspecific interactions

  Juvenile steelhead are known to associate with shiner groups, though they also compete for food. Steelhead form dominance hierarchies within cohorts and aggressively defend feeding territories against ...

  Citation × Citation Citation Abstract Copy Title: Territorial behavior in juvenile steelhead trout (Oncorhynchus mykiss) : how redside shiner (Richardsonius balteatus) influence intraspecific interactions Author: Tinus, Craig A. Juvenile steelhead are known to associate with shiner groups, though they also compete for food. Steelhead form dominance hierarchies within cohorts and aggressively defend feeding territories against all other fish. This study focused on the differential effect of shiner competition on steelhead of different social standing. Survival of subordinate juvenile steelhead was significantly enhanced by the presence of redside shiner under laboratory conditions. A factorial experiment in 80 L tanks examined the relative effects of 0, 3, and 9 shiner at 15° and 20°C on the growth and survival of 3 juvenile steelhead per tank. No temperature effect was detected and there was no significant difference in steelhead growth though statistical power was low (n=5). The largest steelhead did not die in any treatment and no steelhead died in the presence of 9 shiner. In treatments where no shiner were present mortality in the smallest steelhead was 80% (p-value<0.01). Aggressive interactions between steelhead allowed pathogens to colonize breaks in the skin of stressed fish resulting in death. In 6800 L recirculating stream channels with natural substrate, 10 steelhead were held either alone or with 20 shiner at 15°C. No steelhead died and their growth was not significantly different between treatments, but, in the absence of shiner fin damage was 16 times greater (p-value<0.01) in the smallest three steelhead. If a shiner group was present the smallest steelhead appeared to shoal with shiner to avoid attack by dominant steelhead. Title: Territorial behavior in juvenile steelhead trout (Oncorhynchus mykiss) : how redside shiner (Richardsonius balteatus) influence intraspecific interactions Author: Tinus, Craig A. Juvenile steelhead are known to associate with shiner groups, though they also compete for food. Steelhead form dominance hierarchies within cohorts and aggressively defend feeding territories against all other fish. This study focused on the differential effect of shiner competition on steelhead of different social standing. Survival of subordinate juvenile steelhead was significantly enhanced by the presence of redside shiner under laboratory conditions. A factorial experiment in 80 L tanks examined the relative effects of 0, 3, and 9 shiner at 15° and 20°C on the growth and survival of 3 juvenile steelhead per tank. No temperature effect was detected and there was no significant difference in steelhead growth though statistical power was low (n=5). The largest steelhead did not die in any treatment and no steelhead died in the presence of 9 shiner. In treatments where no shiner were present mortality in the smallest steelhead was 80% (p-value<0.01). Aggressive interactions between steelhead allowed pathogens to colonize breaks in the skin of stressed fish resulting in death. In 6800 L recirculating stream channels with natural substrate, 10 steelhead were held either alone or with 20 shiner at 15°C. No steelhead died and their growth was not significantly different between treatments, but, in the absence of shiner fin damage was 16 times greater (p-value<0.01) in the smallest three steelhead. If a shiner group was present the smallest steelhead appeared to shoal with shiner to avoid attack by dominant steelhead. Close

• 27. [Article] Habitat utilization by spotted owls in the West-Central Cascades of Oregon

  Eight adult spotted owls (Strix occidentalis) on the west slope of the Cascade Range in Oregon were fitted with radio transmitters and observed for periods ranging from 271 to 383 days. Mean home range ...

  Citation × Citation Citation Abstract Copy Title: Habitat utilization by spotted owls in the West-Central Cascades of Oregon Author: Forsman, Eric D. Eight adult spotted owls (Strix occidentalis) on the west slope of the Cascade Range in Oregon were fitted with radio transmitters and observed for periods ranging from 271 to 383 days. Mean home range size for the owls was 1,177 ha (range = 920-1,376 ha). There was no significant difference between home range size of males and females. On the average, home ranges of individuals occupying adjacent territories overlapped by 12 percent (range = 3-25 percent). Home ranges of paired individuals overlapped by 50 to 73 percent. The size and shape of home ranges varied on a seasonal basis; generally, the largest home ranges were observed during winter. Nest sites were centrally located within the home ranges utilized by the owls. On the radiotelemetry study area, the mean nearest neighbor distance between nests of adjacent pairs was 2.88 km. On 11 other areas in western Oregon where suitable habitat was available, mean nearest neighbor distances ranged from 1.68 to 3.04 km. The overall mean nearest neighbor distance for the 12 areas was 2.42 km. Home ranges occupied by the radio tagged owls were so large that consistent defense of the entire home range was impossible. Territorial defense appeared to take the form of a decreasing zone of influence centered around the nest; areas near each nest were defended consistently, but defense became increasingly inconsistent with distance from the nest. Discrete territorial boundaries could not be determined and probably did not exist. Territorial interactions between individuals of the same sex were much more aggressive than interactions between individuals of opposite sex. Spotted owls may have responded less aggressively toward individuals of the opposite sex because the advantage of allowing potential mate replacements to remain within the territory outweighed other considerations. Except for a few instances in which 2 of the owls foraged in recently clear-cut areas or in rock talus, virtually all foraging occurred in forests over 30 years old. Old-growth forests, which covered 36 to 64 percent of the area on individual owl home ranges, were strongly preferred for foraging by all of the owls; 92 percent of all foraging occurred in such forests. Utilization of younger forests (30 to 200 years old) was generally either less than or not significantly different than expected, indicating that younger forests were less desirable for foraging than old-growth stands. Old-growth forests were also preferred for roosting (97 percent of all roosts). Large old trees apparently were preferred for roosting during inclement weather because they provided better overhead protection from rain and snow. Small trees in the understory were preferred during warm weather because they provided greater protection from high temperatures and solar radiation. Spotted owls were primarily nocturnal. On the average, they left their day roasts to begin foraging at 14 minutes after sunset and stopped foraging at 21 minutes before sunrise. Diurnal foraging was limited primarily to opportunistic attempts to capture animals that wandered into roost areas. The usual method of foraging was to move from perch to perch at night, watching and listening for potential prey. The average rate of movement while foraging in this manner was 247 m/hr, and the average distance moved per nightly foraging period was 2,782 meters (range = 320-8,895 m). Seven species of mammals comprised the nucleus of the diet, the northern flying squirrel (Glaucomys sabrinus), red tree vole (Phenacomys longicaudus), western red-backed vole (Clethrionomys occidentalis), deer mouse (Peromyscus maniculatus), western pocket gopher (Thomomys mazama), snowshoe hare (Lepus americanus), and bushy-tailed woodrat (Neotoma cinerea). The flying squirrel, the most common animal in the diet, comprised 42 percent of all prey captured. During mid-winter, 85 percent of the prey captured were arboreal mammals. During the rest of the year, terrestrial prey became more abundant in the diet, but the flying squirrel remained the most common animal in the diet. Mean prey size and dietary composition for male and female spotted owls were not significantly different; this suggested that males and females did not partition the prey resource on the basis of size or species. Estimates of biomass consumed per owl per day on the study area ranged from 77.6 to 164.8 g, but the latter estimate was probably inflated. Four hypotheses are presented to explain why spotted owls foraged primarily in older forests and avoided recently cutover areas: (1) the biomass of prey preferred by the owls may have been greater in older forests than in cutover areas, (2) switching from arboreal mammals in older forests to terrestrial mammals in cutover areas might have resulted in increased competition with other nocturnal predators, (3) prey may have been less accessible in the dense vegetation that developed on cutover areas, and (4) spotted owls may have been more susceptible to predation in cutover areas where escape cover was absent. Title: Habitat utilization by spotted owls in the West-Central Cascades of Oregon Author: Forsman, Eric D. Eight adult spotted owls (Strix occidentalis) on the west slope of the Cascade Range in Oregon were fitted with radio transmitters and observed for periods ranging from 271 to 383 days. Mean home range size for the owls was 1,177 ha (range = 920-1,376 ha). There was no significant difference between home range size of males and females. On the average, home ranges of individuals occupying adjacent territories overlapped by 12 percent (range = 3-25 percent). Home ranges of paired individuals overlapped by 50 to 73 percent. The size and shape of home ranges varied on a seasonal basis; generally, the largest home ranges were observed during winter. Nest sites were centrally located within the home ranges utilized by the owls. On the radiotelemetry study area, the mean nearest neighbor distance between nests of adjacent pairs was 2.88 km. On 11 other areas in western Oregon where suitable habitat was available, mean nearest neighbor distances ranged from 1.68 to 3.04 km. The overall mean nearest neighbor distance for the 12 areas was 2.42 km. Home ranges occupied by the radio tagged owls were so large that consistent defense of the entire home range was impossible. Territorial defense appeared to take the form of a decreasing zone of influence centered around the nest; areas near each nest were defended consistently, but defense became increasingly inconsistent with distance from the nest. Discrete territorial boundaries could not be determined and probably did not exist. Territorial interactions between individuals of the same sex were much more aggressive than interactions between individuals of opposite sex. Spotted owls may have responded less aggressively toward individuals of the opposite sex because the advantage of allowing potential mate replacements to remain within the territory outweighed other considerations. Except for a few instances in which 2 of the owls foraged in recently clear-cut areas or in rock talus, virtually all foraging occurred in forests over 30 years old. Old-growth forests, which covered 36 to 64 percent of the area on individual owl home ranges, were strongly preferred for foraging by all of the owls; 92 percent of all foraging occurred in such forests. Utilization of younger forests (30 to 200 years old) was generally either less than or not significantly different than expected, indicating that younger forests were less desirable for foraging than old-growth stands. Old-growth forests were also preferred for roosting (97 percent of all roosts). Large old trees apparently were preferred for roosting during inclement weather because they provided better overhead protection from rain and snow. Small trees in the understory were preferred during warm weather because they provided greater protection from high temperatures and solar radiation. Spotted owls were primarily nocturnal. On the average, they left their day roasts to begin foraging at 14 minutes after sunset and stopped foraging at 21 minutes before sunrise. Diurnal foraging was limited primarily to opportunistic attempts to capture animals that wandered into roost areas. The usual method of foraging was to move from perch to perch at night, watching and listening for potential prey. The average rate of movement while foraging in this manner was 247 m/hr, and the average distance moved per nightly foraging period was 2,782 meters (range = 320-8,895 m). Seven species of mammals comprised the nucleus of the diet, the northern flying squirrel (Glaucomys sabrinus), red tree vole (Phenacomys longicaudus), western red-backed vole (Clethrionomys occidentalis), deer mouse (Peromyscus maniculatus), western pocket gopher (Thomomys mazama), snowshoe hare (Lepus americanus), and bushy-tailed woodrat (Neotoma cinerea). The flying squirrel, the most common animal in the diet, comprised 42 percent of all prey captured. During mid-winter, 85 percent of the prey captured were arboreal mammals. During the rest of the year, terrestrial prey became more abundant in the diet, but the flying squirrel remained the most common animal in the diet. Mean prey size and dietary composition for male and female spotted owls were not significantly different; this suggested that males and females did not partition the prey resource on the basis of size or species. Estimates of biomass consumed per owl per day on the study area ranged from 77.6 to 164.8 g, but the latter estimate was probably inflated. Four hypotheses are presented to explain why spotted owls foraged primarily in older forests and avoided recently cutover areas: (1) the biomass of prey preferred by the owls may have been greater in older forests than in cutover areas, (2) switching from arboreal mammals in older forests to terrestrial mammals in cutover areas might have resulted in increased competition with other nocturnal predators, (3) prey may have been less accessible in the dense vegetation that developed on cutover areas, and (4) spotted owls may have been more susceptible to predation in cutover areas where escape cover was absent. Close

• 28. [Article] Impacts of earlier emerging steelhead fry of hatchery origin on the social structure, distribution, and growth of wild steelhead fry

  Newly emerged steelhead fry (Oncorhynchus mvkiss) of hatchery and wild origins were studied in laboratory stream channels and natural streams. Objectives of the study were to determine if and how earlier emerging ...

  Citation × Citation Citation Abstract Copy Title: Impacts of earlier emerging steelhead fry of hatchery origin on the social structure, distribution, and growth of wild steelhead fry Author: Noble, Sandra M. (Sandra Marie) Newly emerged steelhead fry (Oncorhynchus mvkiss) of hatchery and wild origins were studied in laboratory stream channels and natural streams. Objectives of the study were to determine if and how earlier emerging hatchery fry influence the emigration, realized densities, growth, habitat use, social structure, and activity patterns of localized populations of wild steelhead fry when the hatchery fry have a competitive advantage conferred by larger size and prior residence. During 1986 and 1987, the above variables were observed daily among hatchery and wild steelhead fry in laboratory stream channels for 8 weeks following emergence in June. The habitat use and social activities for fry of both origins were observed weekly in natural stream reaches from June through August in 1987 to corroborate lab findings. In lab channels, both hatchery and wild fry received 2 treatments: living alone (allopatry) and living together (sympatry). In the lab, fry of hatchery origin emerged 7 to 10 d prior to wild fry and remained larger in size during the 8 weeks of study both years. In natural stream reaches, fry of each origin were observed only in allopatric situations. Wild fry in the field emerged from natural redds while hatchery fry were released in stream reaches as unfed, newly emerged (swim-up) fry. Hatchery and wild fry in lab sections were found to be very similar in their emigration rates, distances to nearest neighbor, growth rates, and use of habitat. Both fry types, regardless of treatment or environment (lab or field), established similar stable social structure and used the same types of aggressive acts. Among all lab groups, once a fry became dominant, it retained that social status to the end of the study period. Significant differences (P<.05 both years) among comparison tests were: 1) in allopatric lab sections, wild fry maintained larger densities than hatchery fry, 2) in sympatry, hatchery fry had a greater tendency to establish stable focal points and social hierarchies more readily, defend larger areas, have better condition, prefer pools with overhead cover more frequently, be more aggressive, and reach stable densities more quickly than the wild fry, 3) fewer hatchery fry in sympatry maintained nomadic positions than wild fry in both treatments, 4) in sympatry, hatchery fry directed more acts of overt aggression toward wild fry than other hatchery fry, 5) wild fry in sympatry usually used defensive or less offensive acts of aggression when interacting with other fry, 6) fry of both origins in natural stream reaches maintained greater distances to their nearest neighbor than fry in allopatric lab sections, 7) dominant hatchery fry in both treatments maintained larger focal areas than subdominant fry, 8) hatchery fry maintained longer lengths than wild fry through the duration of the study, and 9) hatchery fry were more aggressive in sympatry than in allopatry. Potential differences (P<.05 in one year and P<.1 in the other year) were: 1) wild fry in sympatry had lower realized densities, maintained smaller focal areas, had greater proportions of nomadic individuals, and established stable social hierarchies slower than wild fry in allopatric lab sections, 2) wild fry in sympatry had poorer condition than all other fry groups in lab sections, 3) in sympatry, wild fry were the recipients of the majority of aggressive acts perpetrated by hatchery fry and other wild fry and usually assumed the subordinate positions within the social hierarchy, 4) all fry in the lab showed a high preference for pools with overhead cover and low preference for gravel and fines and run areas, and 5) wild fry in allopatric lab sections were more socially active than hatchery fry while the reverse was observed in the natural streams. Any influences that could be attributed to inherent differences between stock origins were probably masked by size differences between fry types. The study would have been more complete had I included sympatric lab sections where wild fry emerged first and where fry types emerged simultaneously, and sympatric reaches in natural streams. Results were further confounded by the limited number of wild adults used for broodstock in the lab segment of this study. Progeny produced from so few adults (5 adults of each sex each year) would have very limited genotypic variation compared to what occurs in natural streams. This may partially explain why some findings from lab sections and natural stream reaches differed. Likewise, genotypic expression among wild fry in lab sections may have varied greatly between years. This could explain differences found between years in behavior of wild fry in similar lab treatments. Although this study does not simulate all possible scenarios, results support suspicions that introductions of hatchery fry of larger size and earlier emergence into streams containing wild stocks could disrupt the social structure and negatively influence the realized densities, spatial distribution, growth, and behavior of wild juveniles in recipient streams. Title: Impacts of earlier emerging steelhead fry of hatchery origin on the social structure, distribution, and growth of wild steelhead fry Author: Noble, Sandra M. (Sandra Marie) Newly emerged steelhead fry (Oncorhynchus mvkiss) of hatchery and wild origins were studied in laboratory stream channels and natural streams. Objectives of the study were to determine if and how earlier emerging hatchery fry influence the emigration, realized densities, growth, habitat use, social structure, and activity patterns of localized populations of wild steelhead fry when the hatchery fry have a competitive advantage conferred by larger size and prior residence. During 1986 and 1987, the above variables were observed daily among hatchery and wild steelhead fry in laboratory stream channels for 8 weeks following emergence in June. The habitat use and social activities for fry of both origins were observed weekly in natural stream reaches from June through August in 1987 to corroborate lab findings. In lab channels, both hatchery and wild fry received 2 treatments: living alone (allopatry) and living together (sympatry). In the lab, fry of hatchery origin emerged 7 to 10 d prior to wild fry and remained larger in size during the 8 weeks of study both years. In natural stream reaches, fry of each origin were observed only in allopatric situations. Wild fry in the field emerged from natural redds while hatchery fry were released in stream reaches as unfed, newly emerged (swim-up) fry. Hatchery and wild fry in lab sections were found to be very similar in their emigration rates, distances to nearest neighbor, growth rates, and use of habitat. Both fry types, regardless of treatment or environment (lab or field), established similar stable social structure and used the same types of aggressive acts. Among all lab groups, once a fry became dominant, it retained that social status to the end of the study period. Significant differences (P<.05 both years) among comparison tests were: 1) in allopatric lab sections, wild fry maintained larger densities than hatchery fry, 2) in sympatry, hatchery fry had a greater tendency to establish stable focal points and social hierarchies more readily, defend larger areas, have better condition, prefer pools with overhead cover more frequently, be more aggressive, and reach stable densities more quickly than the wild fry, 3) fewer hatchery fry in sympatry maintained nomadic positions than wild fry in both treatments, 4) in sympatry, hatchery fry directed more acts of overt aggression toward wild fry than other hatchery fry, 5) wild fry in sympatry usually used defensive or less offensive acts of aggression when interacting with other fry, 6) fry of both origins in natural stream reaches maintained greater distances to their nearest neighbor than fry in allopatric lab sections, 7) dominant hatchery fry in both treatments maintained larger focal areas than subdominant fry, 8) hatchery fry maintained longer lengths than wild fry through the duration of the study, and 9) hatchery fry were more aggressive in sympatry than in allopatry. Potential differences (P<.05 in one year and P<.1 in the other year) were: 1) wild fry in sympatry had lower realized densities, maintained smaller focal areas, had greater proportions of nomadic individuals, and established stable social hierarchies slower than wild fry in allopatric lab sections, 2) wild fry in sympatry had poorer condition than all other fry groups in lab sections, 3) in sympatry, wild fry were the recipients of the majority of aggressive acts perpetrated by hatchery fry and other wild fry and usually assumed the subordinate positions within the social hierarchy, 4) all fry in the lab showed a high preference for pools with overhead cover and low preference for gravel and fines and run areas, and 5) wild fry in allopatric lab sections were more socially active than hatchery fry while the reverse was observed in the natural streams. Any influences that could be attributed to inherent differences between stock origins were probably masked by size differences between fry types. The study would have been more complete had I included sympatric lab sections where wild fry emerged first and where fry types emerged simultaneously, and sympatric reaches in natural streams. Results were further confounded by the limited number of wild adults used for broodstock in the lab segment of this study. Progeny produced from so few adults (5 adults of each sex each year) would have very limited genotypic variation compared to what occurs in natural streams. This may partially explain why some findings from lab sections and natural stream reaches differed. Likewise, genotypic expression among wild fry in lab sections may have varied greatly between years. This could explain differences found between years in behavior of wild fry in similar lab treatments. Although this study does not simulate all possible scenarios, results support suspicions that introductions of hatchery fry of larger size and earlier emergence into streams containing wild stocks could disrupt the social structure and negatively influence the realized densities, spatial distribution, growth, and behavior of wild juveniles in recipient streams. Close