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• 2821. [Article] 2008 Amphibian Distribution Surveys in Wadeable Streams and Ponds in Western and Southeast Oregon, Information Reports Number 2010-05

  Abstract -- The ODFW Oregon Conservation Strategy identified monitoring needs for 17 amphibian species native to the state of Oregon that are designated as “Strategy species”, or Species of Greatest Conservation ...

  Citation × Citation Citation Abstract Copy Title: 2008 Amphibian Distribution Surveys in Wadeable Streams and Ponds in Western and Southeast Oregon, Information Reports Number 2010-05 Abstract -- The ODFW Oregon Conservation Strategy identified monitoring needs for 17 amphibian species native to the state of Oregon that are designated as “Strategy species”, or Species of Greatest Conservation Need (per USFWS requirements for State Wildlife Action Plans). The distribution of many species of amphibians in western Oregon is sparsely documented (Oregon Conservation Strategy, page 27). Although a broad-scale survey for amphibian presence would provide important baseline information about amphibian species composition and distribution, most studies have focused on limited areas. The majority of Oregon’s amphibians rely on aquatic habitats at some point of their life, either for breeding and juvenile development or to inhabit as adults. Most aquatic amphibians breed from late winter to early summer, and adults frequently remain in or near their breeding sites into the summer. Most tadpoles and juvenile amphibians are also active in and occupy aquatic habitats during the summer. Ongoing aquatic habitat and fish surveys are opportunities to observe species and life stages (breeding adults, tadpoles and juveniles) that occupy aquatic or riparian habitats during the summer. One cost-effective approach is to combine amphibian surveys with existing aquatic habitat and fish surveys such as those conducted as part of the Oregon Plan for Salmon and Watersheds (OCSRI 1997). The Oregon Plan has been in place since 1997 and the monitoring component provides a survey framework for streams in the lower Columbia River and Oregon coast drainages. The sampling framework is also compatible with implementation of the aquatic components of the Conservation Strategy, as demonstrated by this study. This study describes the presence of amphibians in and along wadeable streams in coast and lower Columbia River drainages of Oregon, ponds and sloughs in the Willamette Valley, and selected streams in the Great Basin of southeast and central Oregon. As a component of monitoring under the Oregon Plan, the Aquatic Inventories Project (AIP) conducts aquatic habitat surveys at randomly selected and spatially balanced sites across all 1st through 4th order streams in coastal and lower Columbia River drainages. The purpose of the habitat surveys is to describe stream morphology, instream physical habitat, and riparian vegetation. Because the surveyors were already observing features within and alongside the stream channel, they were able to record observations of amphibians. The amphibian component was consistent with the survey protocol used by the US Geological Survey’s Amphibian Research and Monitoring Initiative (http://armi.usgs.gov/). The advantage of coupling an amphibian component with the OR Plan aquatic surveys was that it not only was an efficient use of resources, but more importantly, provided information using a statistically rigorous survey design across a broad geographic area. The Native Fish Investigations Project began a six year study in 2007 to document the distribution and abundance of redband trout in the Great Basin region of Eastern Oregon. The site selection procedure is comparable to the statistical standards as the Oregon Plan survey design. Amphibian data are also collected during three other survey projects, and although the site selection procedure does not conform to the same statistical standards as the Oregon Plan survey design, the projects offer a number of opportunities to collect amphibian occurrence information over a wide variety of habitats. The amphibian observations from these three projects are also included in this report. The three projects are as follows: • AIP conducts aquatic habitat surveys on selected streams throughout the state. • AIP conducts aquatic habitat surveys at stream habitat restoration projects in Western Oregon. • Native Fish Project conducts surveys of pond and slough sites for Oregon chub in the Willamette Valley. Due to the success of the 2006 and 2007 field studies, we continued our research during the summer of 2008 to improve our knowledge of distribution and community structure of amphibians. The summer 2008 surveys took place in 9 of Oregon’s 10 ecoregions (Figure 1) (Thorson et al. 2003). Ecoregions provide a framework for discussing amphibian distribution across the state because they are relatively large areas defined by distinctive geographic and ecological (flora and fauna) characteristics. The goals of our 2008 work were to: • Increase the consistency, efficiency and ability of habitat crews in identifying amphibians through improved training. • Increase knowledge of distribution, community structure, and habitat associations of amphibians in streams in: o Western Oregon coastal and lower Columbia drainages. o Ponds, sloughs and other off-channel aquatic habitats in the Willamette Valley. o Great Basin of eastern Oregon and selected streams in central Oregon. • Combine the 2008 observations with the 2006-07 results. Title: 2008 Amphibian Distribution Surveys in Wadeable Streams and Ponds in Western and Southeast Oregon, Information Reports Number 2010-05 Abstract -- The ODFW Oregon Conservation Strategy identified monitoring needs for 17 amphibian species native to the state of Oregon that are designated as “Strategy species”, or Species of Greatest Conservation Need (per USFWS requirements for State Wildlife Action Plans). The distribution of many species of amphibians in western Oregon is sparsely documented (Oregon Conservation Strategy, page 27). Although a broad-scale survey for amphibian presence would provide important baseline information about amphibian species composition and distribution, most studies have focused on limited areas. The majority of Oregon’s amphibians rely on aquatic habitats at some point of their life, either for breeding and juvenile development or to inhabit as adults. Most aquatic amphibians breed from late winter to early summer, and adults frequently remain in or near their breeding sites into the summer. Most tadpoles and juvenile amphibians are also active in and occupy aquatic habitats during the summer. Ongoing aquatic habitat and fish surveys are opportunities to observe species and life stages (breeding adults, tadpoles and juveniles) that occupy aquatic or riparian habitats during the summer. One cost-effective approach is to combine amphibian surveys with existing aquatic habitat and fish surveys such as those conducted as part of the Oregon Plan for Salmon and Watersheds (OCSRI 1997). The Oregon Plan has been in place since 1997 and the monitoring component provides a survey framework for streams in the lower Columbia River and Oregon coast drainages. The sampling framework is also compatible with implementation of the aquatic components of the Conservation Strategy, as demonstrated by this study. This study describes the presence of amphibians in and along wadeable streams in coast and lower Columbia River drainages of Oregon, ponds and sloughs in the Willamette Valley, and selected streams in the Great Basin of southeast and central Oregon. As a component of monitoring under the Oregon Plan, the Aquatic Inventories Project (AIP) conducts aquatic habitat surveys at randomly selected and spatially balanced sites across all 1st through 4th order streams in coastal and lower Columbia River drainages. The purpose of the habitat surveys is to describe stream morphology, instream physical habitat, and riparian vegetation. Because the surveyors were already observing features within and alongside the stream channel, they were able to record observations of amphibians. The amphibian component was consistent with the survey protocol used by the US Geological Survey’s Amphibian Research and Monitoring Initiative (http://armi.usgs.gov/). The advantage of coupling an amphibian component with the OR Plan aquatic surveys was that it not only was an efficient use of resources, but more importantly, provided information using a statistically rigorous survey design across a broad geographic area. The Native Fish Investigations Project began a six year study in 2007 to document the distribution and abundance of redband trout in the Great Basin region of Eastern Oregon. The site selection procedure is comparable to the statistical standards as the Oregon Plan survey design. Amphibian data are also collected during three other survey projects, and although the site selection procedure does not conform to the same statistical standards as the Oregon Plan survey design, the projects offer a number of opportunities to collect amphibian occurrence information over a wide variety of habitats. The amphibian observations from these three projects are also included in this report. The three projects are as follows: • AIP conducts aquatic habitat surveys on selected streams throughout the state. • AIP conducts aquatic habitat surveys at stream habitat restoration projects in Western Oregon. • Native Fish Project conducts surveys of pond and slough sites for Oregon chub in the Willamette Valley. Due to the success of the 2006 and 2007 field studies, we continued our research during the summer of 2008 to improve our knowledge of distribution and community structure of amphibians. The summer 2008 surveys took place in 9 of Oregon’s 10 ecoregions (Figure 1) (Thorson et al. 2003). Ecoregions provide a framework for discussing amphibian distribution across the state because they are relatively large areas defined by distinctive geographic and ecological (flora and fauna) characteristics. The goals of our 2008 work were to: • Increase the consistency, efficiency and ability of habitat crews in identifying amphibians through improved training. • Increase knowledge of distribution, community structure, and habitat associations of amphibians in streams in: o Western Oregon coastal and lower Columbia drainages. o Ponds, sloughs and other off-channel aquatic habitats in the Willamette Valley. o Great Basin of eastern Oregon and selected streams in central Oregon. • Combine the 2008 observations with the 2006-07 results. Close

• 2822. [Article] Amphibian Distribution in Wadeable Streams and Ponds in Western and Southeast Oregon, 2009-2010 Progress Reports 2011

  Abstract -- The ODFW Oregon Conservation Strategy identified monitoring needs for 17 amphibian species native to the state of Oregon that are designated as “Strategy Species”, or “Species of Greatest Conservation ...

  Citation × Citation Citation Abstract Copy Title: Amphibian Distribution in Wadeable Streams and Ponds in Western and Southeast Oregon, 2009-2010 Progress Reports 2011 Abstract -- The ODFW Oregon Conservation Strategy identified monitoring needs for 17 amphibian species native to the state of Oregon that are designated as “Strategy Species”, or “Species of Greatest Conservation Need” (per USFWS requirements for State Wildlife Action Plans). The distribution of many species of amphibians in western Oregon is sparsely documented (Oregon Conservation Strategy, page 27). Although a broad-scale survey for amphibian presence would provide important baseline information about amphibian species composition and distribution, most studies have focused on limited areas. The majority of Oregon’s amphibians rely on aquatic habitats at some point of their life, either for breeding and juvenile development or to inhabit as adults. Most aquatic amphibians breed from late winter to early summer, and adults frequently remain in or near their breeding sites into the summer. Most tadpoles and juvenile amphibians are also active in and occupy aquatic habitats during the summer. Ongoing aquatic habitat and fish surveys are opportunities to observe species and life stages (breeding adults, tadpoles and juveniles) that occupy aquatic or riparian habitats during the summer. One cost-effective approach is to combine amphibian surveys with existing aquatic habitat and fish surveys such as those conducted as part of the Oregon Plan for Salmon and Watersheds (OCSRI 1997). The Oregon Plan has been in place since 1997 and the monitoring component provides a survey framework for streams in the lower Columbia River and Oregon coast drainages. The sampling framework is also compatible with implementation of the aquatic components of the Conservation Strategy, as demonstrated by this study. This study describes the presence of amphibians in and along wadeable streams in coast and lower Columbia River drainages of Oregon, ponds and sloughs in the Willamette Valley, and selected streams in the Great Basin of southeast and central Oregon. As a component of monitoring under the Oregon Plan, the Aquatic Inventories Project (AIP) conducts aquatic habitat surveys at randomly selected and spatially balanced sites across all 1st through 4th order streams in coastal and lower Columbia River drainages. The purpose of the habitat surveys is to describe stream morphology, instream physical habitat, and riparian vegetation. Because the surveyors were already observing features within and alongside the stream channel, they were able to record observations of amphibians. The advantage of coupling an amphibian component with the OR Plan aquatic surveys was that it not only was an efficient use of resources, but more importantly, provided information using a statistically rigorous survey design across a broad geographic area. The Native Fish Investigations Project began a six year study in 2007 to document the distribution and abundance of redband trout in the Great Basin region of Eastern Oregon. The site selection procedure is comparable to the statistical standards as the Oregon Plan survey design. Amphibian data are also collected during three other survey projects, and although the site selection procedure does not conform to the same statistical standards as the Oregon Plan survey design, the projects offer a number of opportunities to collect amphibian occurrence information over a wide variety of habitats. The amphibian observations from these three projects are also included in this report. The three projects are as follows: AIP conducts aquatic habitat surveys on selected streams throughout the state. AIP conducts aquatic habitat surveys at stream habitat restoration projects in Western Oregon. Native Fish Project conducts surveys of pond and slough sites for Oregon chub in the Willamette Valley. Due to the success of the 2007 and 2008 field studies, we continued our research during the summer of 2009 and 2010 to improve our knowledge of distribution and community structure of amphibians. The summer 2009 and 2010 surveys took place in 9 of Oregon’s 10 ecoregions (Figure 1) (Thorson et al. 2003). Ecoregions provide a framework for discussing amphibian distribution across the state because they are relatively large areas defined by distinctive geographic and ecological (flora and fauna) characteristics. The goals of our 2009-2010 work were to: Increase the consistency, efficiency and ability of habitat crews in identifying amphibians through improved training. Increase knowledge of distribution, community structure, and habitat associations of amphibians in streams in: Western Oregon coastal and lower Columbia drainages. Ponds, sloughs and other off-channel aquatic habitats in the Willamette Valley. Great Basin of eastern Oregon and selected streams in central Oregon. Combine the 2009-2010 observations with the 2007-2008 results. Title: Amphibian Distribution in Wadeable Streams and Ponds in Western and Southeast Oregon, 2009-2010 Progress Reports 2011 Abstract -- The ODFW Oregon Conservation Strategy identified monitoring needs for 17 amphibian species native to the state of Oregon that are designated as “Strategy Species”, or “Species of Greatest Conservation Need” (per USFWS requirements for State Wildlife Action Plans). The distribution of many species of amphibians in western Oregon is sparsely documented (Oregon Conservation Strategy, page 27). Although a broad-scale survey for amphibian presence would provide important baseline information about amphibian species composition and distribution, most studies have focused on limited areas. The majority of Oregon’s amphibians rely on aquatic habitats at some point of their life, either for breeding and juvenile development or to inhabit as adults. Most aquatic amphibians breed from late winter to early summer, and adults frequently remain in or near their breeding sites into the summer. Most tadpoles and juvenile amphibians are also active in and occupy aquatic habitats during the summer. Ongoing aquatic habitat and fish surveys are opportunities to observe species and life stages (breeding adults, tadpoles and juveniles) that occupy aquatic or riparian habitats during the summer. One cost-effective approach is to combine amphibian surveys with existing aquatic habitat and fish surveys such as those conducted as part of the Oregon Plan for Salmon and Watersheds (OCSRI 1997). The Oregon Plan has been in place since 1997 and the monitoring component provides a survey framework for streams in the lower Columbia River and Oregon coast drainages. The sampling framework is also compatible with implementation of the aquatic components of the Conservation Strategy, as demonstrated by this study. This study describes the presence of amphibians in and along wadeable streams in coast and lower Columbia River drainages of Oregon, ponds and sloughs in the Willamette Valley, and selected streams in the Great Basin of southeast and central Oregon. As a component of monitoring under the Oregon Plan, the Aquatic Inventories Project (AIP) conducts aquatic habitat surveys at randomly selected and spatially balanced sites across all 1st through 4th order streams in coastal and lower Columbia River drainages. The purpose of the habitat surveys is to describe stream morphology, instream physical habitat, and riparian vegetation. Because the surveyors were already observing features within and alongside the stream channel, they were able to record observations of amphibians. The advantage of coupling an amphibian component with the OR Plan aquatic surveys was that it not only was an efficient use of resources, but more importantly, provided information using a statistically rigorous survey design across a broad geographic area. The Native Fish Investigations Project began a six year study in 2007 to document the distribution and abundance of redband trout in the Great Basin region of Eastern Oregon. The site selection procedure is comparable to the statistical standards as the Oregon Plan survey design. Amphibian data are also collected during three other survey projects, and although the site selection procedure does not conform to the same statistical standards as the Oregon Plan survey design, the projects offer a number of opportunities to collect amphibian occurrence information over a wide variety of habitats. The amphibian observations from these three projects are also included in this report. The three projects are as follows: AIP conducts aquatic habitat surveys on selected streams throughout the state. AIP conducts aquatic habitat surveys at stream habitat restoration projects in Western Oregon. Native Fish Project conducts surveys of pond and slough sites for Oregon chub in the Willamette Valley. Due to the success of the 2007 and 2008 field studies, we continued our research during the summer of 2009 and 2010 to improve our knowledge of distribution and community structure of amphibians. The summer 2009 and 2010 surveys took place in 9 of Oregon’s 10 ecoregions (Figure 1) (Thorson et al. 2003). Ecoregions provide a framework for discussing amphibian distribution across the state because they are relatively large areas defined by distinctive geographic and ecological (flora and fauna) characteristics. The goals of our 2009-2010 work were to: Increase the consistency, efficiency and ability of habitat crews in identifying amphibians through improved training. Increase knowledge of distribution, community structure, and habitat associations of amphibians in streams in: Western Oregon coastal and lower Columbia drainages. Ponds, sloughs and other off-channel aquatic habitats in the Willamette Valley. Great Basin of eastern Oregon and selected streams in central Oregon. Combine the 2009-2010 observations with the 2007-2008 results. Close

• 2823. [Article] Distribution and movements of Chinook salmon, Oncorhynchus tshawytscha, returning to the Yukon River basin

  Chinook salmon, Oncorhynchus tshawytscha, returning to the Yukon River basin and other large river systems in western Alaska have declined dramatically since the late 1990s. This continuing trend has ...

  Citation × Citation Citation Abstract Copy Title: Distribution and movements of Chinook salmon, Oncorhynchus tshawytscha, returning to the Yukon River basin Author: Eiler, John H. Chinook salmon, Oncorhynchus tshawytscha, returning to the Yukon River basin and other large river systems in western Alaska have declined dramatically since the late 1990s. This continuing trend has raised concerns over the future status of the returns, and severely impacted commercial and subsistence fisheries within the drainage. Management is further complicated by the mixed-stock composition of the run, the presence of other temporally similar salmon species, and the need to equitably allocate harvests between the numerous fisheries and user groups scattered throughout the basin. Detailed information is needed on Chinook salmon run characteristics to better understand and manage the returns, and facilitate conservation efforts. However, this goal is exacerbated by the massive size and remote nature of the basin, the large number of highly mobile fish, and the compressed timing of the run. To address these challenges, radio telemetry was used to determine the stock composition and spawning distribution of the returns, and the migratory characteristics of the fish. The migratory patterns exhibited by returning salmon provide a number of insights into the status of the run. Since the Yukon River is essentially free-flowing (i.e., not regulated), this study also presented an opportunity to document the distribution and upriver movements of large returns of wild Chinook salmon under natural conditions. During 2002-2004, returning adult Chinook salmon were captured in the lower Yukon River (approximately 300 km upriver from the river mouth), tagged with radio transmitters, and tracked upriver using remote tracking stations located on important migratory routes and major spawning tributaries. Aerial tracking surveys were used to locate fish in spawning areas and between stations. The fish responded well to the capture and handling procedures, with most (2,790, 98%) resuming upriver movements. Although the fish initially displayed a negative tagging response, with slower migration rates observed immediately after release, the duration of this response was relatively short (several days) and less severe as the fish moved upriver. Independent measures indicated that the swimming speeds and timing of the fish upriver from the tagging area were comparable to untagged fish, suggesting that the tagging methods used were relatively benign. Fish returned to spawning areas throughout the basin, ranging from several hundred to over 3,000 km from the tagging area. Distribution patterns were similar across years, suggesting that the principal components of the run were identified. Most spawning fish were clustered in a number of key tributaries, with smaller numbers of fish located in other spatially isolated areas. The fish typically returned to clear water tributaries that were relatively entrenched, had moderate gradients, and were associated with upland areas. Fish were largely absent in lowland reaches characterized by meandering, low gradient, highly alluvial channels often associated with main river floodplains. There was suggestive evidence of mainstem spawning in reaches of the Upper Yukon. The status of fish remaining in other mainstem areas was less certain, and may represent local spawning activity or fish that died while in-transit to upriver areas. Although Chinook salmon spawned throughout the basin, the run was dominated by two regional components (Tanana and Upper Yukon), which annually comprised over 70% of the return. Substantially fewer fish returned to other areas ranging from 2-9% of the return, although the collective contribution of these stocks was appreciable. Most regional returns consisted of several principal stocks and a number of small, spatially isolated populations. Regional and stock composition estimates were similar across years even though differences in run abundance were reported, suggesting that these abundance differences were not related to regional or stock-specific differences. Run timing was relatively compressed compared to rivers in the southern portion of the range, with most stocks passing through the lower river over a 6-week period, ranging from 16 to 38 d. Run timing was generally earlier for stocks traveling farther upriver, although exceptions were noted. Lower basin stocks were primarily later run fish. Pronounced differences were observed in the migration rates (km/d) exhibited by regional stocks. Substantially slower swimming speeds were observed for fish returning to terminal tributaries in the lower basin ranging from 28-40 km/d compared to 52-62 km/d for upper basin stocks. The migratory patterns (migration rates in sequential reaches) of the fish also showed distinct regional differences. Average migration rates through the lower river were remarkably similar for the different stocks, ranging from 57-62 km/d, with most stocks exhibiting a general decline as the fish moved farther upriver. Tanana River stocks displayed a pronounced reduction in swimming speed after leaving the Yukon River main stem, with migration rates declining to 24 km/d on average as the fish approached their terminal tributaries. Conversely, upper basin stocks exhibited a relatively gradual (but variable) overall decline in migration rate even though these fish were traveling substantially greater distances upriver. Average migration rates for upper basin stocks ranged from 43-61 km/d as the fish approached their terminal tributaries. There was substantial variation in the migratory patterns exhibited by individual fish, although these patterns tended to be similar to the patterns exhibited by the regional stocks, particularly as the fish moved farther upriver from the tagging area. The dominant source of variation among fish reflected the average migration rate, with individual fish traveling slower in the lower basin exhibiting consistently slower migration rates as they moved upriver compared to their faster moving counterparts. This migratory pattern was consistent across stocks, and on average explained 74% of the within-stock variation in migration rate represented by the multivariate data. The second source of variation in migration rate reflected a shift in the relative swimming speeds of the individual fish as they progressed upriver. Although movement rates declined for nearly all of the fish during the migration, differences were observed in the pattern of the decline. Fish with faster migration rates in the lower river exhibited a pronounced decline in swimming speed as they moved upriver, whereas fish moving slower in the lower river displayed a more gradual decline in migration rate. On average, this migratory pattern explained 22% of the within-stock variation in migration rate represented by the multivariate data. Most fish (98%) exhibited continuous upriver movements and strong fidelity to the rivers they entered. However a small number of fish (n = 66) deviated from this pattern. Some of these individuals initially passed their final destination and continued upriver for varying distances before reversing direction, swimming back downstream, and entering their terminal tributary. Although most of these excursions were relatively short (< 30 km), there were several instances where fish traveled hundreds of kilometers out of their way. Thirty-four fish tracked to terminal tributaries subsequently left these rivers, and traveled to other terminal tributaries within the basin (n = 31) or were harvested in upriver fisheries (n = 3). Although most of these incidents involved nearby tributaries, major diversions were also observed, with several fish traveling over 300 km to natal rivers after leaving the initial tributary. Chinook salmon returns to the Yukon River typically consisted of a series of distinct and sizable increases in the number fish entering the river over the course of the run, commonly referred to as pulses. A large number of fish (n = 251) were radio tagged over a 4-day period during a pulse in 2003 to provide information on the progression of the pulse as it moved upriver. The time taken by the pulse to move past subsequent upriver locations increased as the fish moved farther upriver from the tagging area, with the fish passing sites located 580 and 800 km upriver over a span of 14 and 21 d, respectively. Although not surprising considering the extensive variation in migration rates observed among individual fish, this finding does suggest that these pulses do not represent cohesive aggregates of fish moving upriver. Unlike the well established methods used to estimate other life history characteristics, the development of quantitative methods for analyzing and modeling fish movements has lagged noticeably behind, due in part to the complexity associated with movement data and (prior to the advent of telemetry) the difficulty of collecting this type of information on free-ranging individuals. Two fundamentally different analytical approaches, hierarchical linear regression models and multivariate ordination, were used during this study to evaluate factors thought to influence the upriver movements of the fish. In spite of the inherent differences, both methods provided strikingly similar results, indicating that the study findings were not dependent on the approach used, and suggesting that the results were plausible based on the information available and the weight of evidence. Both analytical methods had advantages, and provided complementary information. With hierarchical linear models, it was possible to simultaneously evaluate a wide range of explanatory variables (in our case, both biological and environmental), which provided standardized comparisons and simplified the interpretation of the results. Since both fixed and random effects were incorporated in the models, it was possible to account for sources of variation when insufficient information was available to identify the underlining factors – an important consideration since few field studies provide comprehensive data. With multivariate ordination, separate analyzes were needed to examine the relationships between the migration rates and the biotic and physical variables. In addition to being cumbersome, this limitation made it more difficult to compare the relative influence of the different factors and interactions between factors. However, ordination was very useful as an exploratory tool. Although compartmentalized by stock, across fish comparisons were simple and relatively straightforward. Because the explanatory variables were evaluated separately in relation to the ordination score assigned to the fish, it was possible to examine and compare highly correlated variables. Ordination was also able to identify overall patterns within the data and assess the relative importance. While this can be accomplished within the framework of linear regression using mixture models to determine whether multiple distributions exist within the data, the process is much simpler with ordination. The migratory patterns of the fish were influenced by a wide range of factors, with evidentiary support for complex, multi-faceted relationships. Physical features of the basin demonstrated stronger explanatory power, accounting for over 70% of the observed variation in migration rate compared to 18% for the biological characteristics of the fish. Parameter estimates associated with the steepness of the migratory route and remaining distance the fish had to travel to reach their natal rivers were most strongly correlated with migration rate, with consistent relationships observed across stocks. Migration rates were also noticeably slower in extensively braided reaches of the basin. The weaker relationships between migration rate and biotic factors may reflect stabilizing selection on long-distance migrants. Smaller fish exhibited minimally faster swimming speeds on average than larger individuals. This relationship was stronger in highly braided reaches. Run timing was positively related to migration rate for most stocks. Surprisingly, upper basin stocks traveling farther upriver displayed progressively negative relationships, suggesting that late-run fish were moving slower. Ancillary information suggests that this decline may relate to deteriorating fish condition later in the season. Title: Distribution and movements of Chinook salmon, Oncorhynchus tshawytscha, returning to the Yukon River basin Author: Eiler, John H. Chinook salmon, Oncorhynchus tshawytscha, returning to the Yukon River basin and other large river systems in western Alaska have declined dramatically since the late 1990s. This continuing trend has raised concerns over the future status of the returns, and severely impacted commercial and subsistence fisheries within the drainage. Management is further complicated by the mixed-stock composition of the run, the presence of other temporally similar salmon species, and the need to equitably allocate harvests between the numerous fisheries and user groups scattered throughout the basin. Detailed information is needed on Chinook salmon run characteristics to better understand and manage the returns, and facilitate conservation efforts. However, this goal is exacerbated by the massive size and remote nature of the basin, the large number of highly mobile fish, and the compressed timing of the run. To address these challenges, radio telemetry was used to determine the stock composition and spawning distribution of the returns, and the migratory characteristics of the fish. The migratory patterns exhibited by returning salmon provide a number of insights into the status of the run. Since the Yukon River is essentially free-flowing (i.e., not regulated), this study also presented an opportunity to document the distribution and upriver movements of large returns of wild Chinook salmon under natural conditions. During 2002-2004, returning adult Chinook salmon were captured in the lower Yukon River (approximately 300 km upriver from the river mouth), tagged with radio transmitters, and tracked upriver using remote tracking stations located on important migratory routes and major spawning tributaries. Aerial tracking surveys were used to locate fish in spawning areas and between stations. The fish responded well to the capture and handling procedures, with most (2,790, 98%) resuming upriver movements. Although the fish initially displayed a negative tagging response, with slower migration rates observed immediately after release, the duration of this response was relatively short (several days) and less severe as the fish moved upriver. Independent measures indicated that the swimming speeds and timing of the fish upriver from the tagging area were comparable to untagged fish, suggesting that the tagging methods used were relatively benign. Fish returned to spawning areas throughout the basin, ranging from several hundred to over 3,000 km from the tagging area. Distribution patterns were similar across years, suggesting that the principal components of the run were identified. Most spawning fish were clustered in a number of key tributaries, with smaller numbers of fish located in other spatially isolated areas. The fish typically returned to clear water tributaries that were relatively entrenched, had moderate gradients, and were associated with upland areas. Fish were largely absent in lowland reaches characterized by meandering, low gradient, highly alluvial channels often associated with main river floodplains. There was suggestive evidence of mainstem spawning in reaches of the Upper Yukon. The status of fish remaining in other mainstem areas was less certain, and may represent local spawning activity or fish that died while in-transit to upriver areas. Although Chinook salmon spawned throughout the basin, the run was dominated by two regional components (Tanana and Upper Yukon), which annually comprised over 70% of the return. Substantially fewer fish returned to other areas ranging from 2-9% of the return, although the collective contribution of these stocks was appreciable. Most regional returns consisted of several principal stocks and a number of small, spatially isolated populations. Regional and stock composition estimates were similar across years even though differences in run abundance were reported, suggesting that these abundance differences were not related to regional or stock-specific differences. Run timing was relatively compressed compared to rivers in the southern portion of the range, with most stocks passing through the lower river over a 6-week period, ranging from 16 to 38 d. Run timing was generally earlier for stocks traveling farther upriver, although exceptions were noted. Lower basin stocks were primarily later run fish. Pronounced differences were observed in the migration rates (km/d) exhibited by regional stocks. Substantially slower swimming speeds were observed for fish returning to terminal tributaries in the lower basin ranging from 28-40 km/d compared to 52-62 km/d for upper basin stocks. The migratory patterns (migration rates in sequential reaches) of the fish also showed distinct regional differences. Average migration rates through the lower river were remarkably similar for the different stocks, ranging from 57-62 km/d, with most stocks exhibiting a general decline as the fish moved farther upriver. Tanana River stocks displayed a pronounced reduction in swimming speed after leaving the Yukon River main stem, with migration rates declining to 24 km/d on average as the fish approached their terminal tributaries. Conversely, upper basin stocks exhibited a relatively gradual (but variable) overall decline in migration rate even though these fish were traveling substantially greater distances upriver. Average migration rates for upper basin stocks ranged from 43-61 km/d as the fish approached their terminal tributaries. There was substantial variation in the migratory patterns exhibited by individual fish, although these patterns tended to be similar to the patterns exhibited by the regional stocks, particularly as the fish moved farther upriver from the tagging area. The dominant source of variation among fish reflected the average migration rate, with individual fish traveling slower in the lower basin exhibiting consistently slower migration rates as they moved upriver compared to their faster moving counterparts. This migratory pattern was consistent across stocks, and on average explained 74% of the within-stock variation in migration rate represented by the multivariate data. The second source of variation in migration rate reflected a shift in the relative swimming speeds of the individual fish as they progressed upriver. Although movement rates declined for nearly all of the fish during the migration, differences were observed in the pattern of the decline. Fish with faster migration rates in the lower river exhibited a pronounced decline in swimming speed as they moved upriver, whereas fish moving slower in the lower river displayed a more gradual decline in migration rate. On average, this migratory pattern explained 22% of the within-stock variation in migration rate represented by the multivariate data. Most fish (98%) exhibited continuous upriver movements and strong fidelity to the rivers they entered. However a small number of fish (n = 66) deviated from this pattern. Some of these individuals initially passed their final destination and continued upriver for varying distances before reversing direction, swimming back downstream, and entering their terminal tributary. Although most of these excursions were relatively short (< 30 km), there were several instances where fish traveled hundreds of kilometers out of their way. Thirty-four fish tracked to terminal tributaries subsequently left these rivers, and traveled to other terminal tributaries within the basin (n = 31) or were harvested in upriver fisheries (n = 3). Although most of these incidents involved nearby tributaries, major diversions were also observed, with several fish traveling over 300 km to natal rivers after leaving the initial tributary. Chinook salmon returns to the Yukon River typically consisted of a series of distinct and sizable increases in the number fish entering the river over the course of the run, commonly referred to as pulses. A large number of fish (n = 251) were radio tagged over a 4-day period during a pulse in 2003 to provide information on the progression of the pulse as it moved upriver. The time taken by the pulse to move past subsequent upriver locations increased as the fish moved farther upriver from the tagging area, with the fish passing sites located 580 and 800 km upriver over a span of 14 and 21 d, respectively. Although not surprising considering the extensive variation in migration rates observed among individual fish, this finding does suggest that these pulses do not represent cohesive aggregates of fish moving upriver. Unlike the well established methods used to estimate other life history characteristics, the development of quantitative methods for analyzing and modeling fish movements has lagged noticeably behind, due in part to the complexity associated with movement data and (prior to the advent of telemetry) the difficulty of collecting this type of information on free-ranging individuals. Two fundamentally different analytical approaches, hierarchical linear regression models and multivariate ordination, were used during this study to evaluate factors thought to influence the upriver movements of the fish. In spite of the inherent differences, both methods provided strikingly similar results, indicating that the study findings were not dependent on the approach used, and suggesting that the results were plausible based on the information available and the weight of evidence. Both analytical methods had advantages, and provided complementary information. With hierarchical linear models, it was possible to simultaneously evaluate a wide range of explanatory variables (in our case, both biological and environmental), which provided standardized comparisons and simplified the interpretation of the results. Since both fixed and random effects were incorporated in the models, it was possible to account for sources of variation when insufficient information was available to identify the underlining factors – an important consideration since few field studies provide comprehensive data. With multivariate ordination, separate analyzes were needed to examine the relationships between the migration rates and the biotic and physical variables. In addition to being cumbersome, this limitation made it more difficult to compare the relative influence of the different factors and interactions between factors. However, ordination was very useful as an exploratory tool. Although compartmentalized by stock, across fish comparisons were simple and relatively straightforward. Because the explanatory variables were evaluated separately in relation to the ordination score assigned to the fish, it was possible to examine and compare highly correlated variables. Ordination was also able to identify overall patterns within the data and assess the relative importance. While this can be accomplished within the framework of linear regression using mixture models to determine whether multiple distributions exist within the data, the process is much simpler with ordination. The migratory patterns of the fish were influenced by a wide range of factors, with evidentiary support for complex, multi-faceted relationships. Physical features of the basin demonstrated stronger explanatory power, accounting for over 70% of the observed variation in migration rate compared to 18% for the biological characteristics of the fish. Parameter estimates associated with the steepness of the migratory route and remaining distance the fish had to travel to reach their natal rivers were most strongly correlated with migration rate, with consistent relationships observed across stocks. Migration rates were also noticeably slower in extensively braided reaches of the basin. The weaker relationships between migration rate and biotic factors may reflect stabilizing selection on long-distance migrants. Smaller fish exhibited minimally faster swimming speeds on average than larger individuals. This relationship was stronger in highly braided reaches. Run timing was positively related to migration rate for most stocks. Surprisingly, upper basin stocks traveling farther upriver displayed progressively negative relationships, suggesting that late-run fish were moving slower. Ancillary information suggests that this decline may relate to deteriorating fish condition later in the season. Close

• 2824. [Article] Dead wood dynamics and relationships to biophysical factors, forest history, ownership, and management practices in the Coastal Province of Oregon, USA

  Dead wood patterns and dynamics vary with biophysical factors, disturbance history, ownership, and management practices. Through field and modeling studies, I examined the current and potential future ...

  Citation × Citation Citation Abstract Copy Title: Dead wood dynamics and relationships to biophysical factors, forest history, ownership, and management practices in the Coastal Province of Oregon, USA Author: Kennedy, Rebecca S.H. Dead wood patterns and dynamics vary with biophysical factors, disturbance history, ownership, and management practices. Through field and modeling studies, I examined the current and potential future amounts of dead wood in two landscapes and region-wide in the Coastal Province of Oregon. The objectives of the first study were to (1) determine whether two landscapes with different recent disturbance histories differ in the amount and characteristics of dead wood; and (2) explore relationships between patterns of dead wood in each landscape to potentially related factors including topography. The objectives of the second study were to (1) describe current regional amounts of dead wood; (2) compare dead wood amounts across ownerships; (3) determine relationships between current dead wood amounts and ownership, current and past vegetation conditions, climate, topography, and soils; and (4) evaluate whether the factors related to dead wood patterns differed according to the scale of analysis. The objectives of the third study were to (1) characterize the projected future change in dead wood amounts in a multi-ownership Province; (2) determine the longevity of present-day dead wood of different types and sizes in relation to amendments from management and stand development; and (3) evaluate differences in management approaches in transitional dynamics and long-term patterns of dead wood. In the first study, I sampled logs and snags at four topographic positions (streams, lower slopes, middle slopes, upper slopes) in the Tillamook State Forest and the Siuslaw National Forest. These two landscapes experienced catastrophic fire at different points in recent history. I developed statistical models relating various attributes of dead wood abundance to biophysical variables related to climate, topography, historical vegetation, current vegetation, soils, and ecoregion. I found that the type and timing of disturbance was important to dead wood amounts and characteristics, and that potential source and sink areas for dead wood were related to topographic position. In particular, lower slopes had higher amounts of logs, and upper slopes had higher basal areas of potential source wood, in the form of snags and legacy (pre-fire) stumps. Climatic factors were of greater relative importance to overall gradients of dead wood in the landscape in which fire occurred less recently. In the second study, I analyzed dead wood data from a region-wide systematic grid of field plots according to ownership and biophysical variables at multiple scales of resolution including plots, subwatersheds. Dead wood abundance and types varied greatly among ownerships, with public lands (Forest Service, Bureau of Land Management, State of Oregon) typically having higher amounts of dead wood and more dead wood in the larger size classes than the private lands (forest industry, non-industrial private). I found that the relative influence of ownership, topography, current and historical vegetation, and climate varied with scale of resolution. Current vegetation was of greater relative importance at finer scales of plots and subwatersheds, whereas climate, topography, and historical vegetation were of greater relative importance at coarser scales of watersheds and subbasins. Ownership was important to overall dead wood gradients at all scales considered. In the third study, by simulating stand development and dead wood dynamics under various forest management scenarios over a 300-year period, I was able to examine the long-term effects of management on dead wood abundance in the Coastal Province. I estimated potential upper bounds for future dead wood amounts. Dead wood amounts increased over time on average across the Province, mainly because of policies on public lands, especially the federal lands under the Northwest Forest Plan. Forest industry, under the Oregon Forest Practices Act and assuming retention of all snags at harvest and thinning, maintained amounts of dead wood that were similar to present-day levels, but size classes shifted toward the smaller sizes as existing large legacy dead wood decomposed. Non-industrial private lands showed increases from very low present-day amounts of dead wood. Across the Province, legacy logs and snags remained present for over a century of the simulation period, and buffered effects of intensive management to dead wood amounts. Variation across landscapes in starting conditions meant that contrasting management approaches had differential effects on long-term dead wood dynamics depending on where they were applied. Current amounts of dead wood and live vegetation patterns in the Province resulted from historical fire and logging. Results of this simulation study indicate that recently established policies oriented toward dead wood production and retention, in the absence of fire or other large- or mid-scale disturbances, are likely to result in increases in dead wood amounts that greatly exceed present-day levels. My results suggest that dead wood patterns of abundance will continue to diverge according to land ownership and that management practices that foster dead wood creation are of increasing importance to the long-term abundance of large dead wood as legacy dead wood is lost through decomposition. Title: Dead wood dynamics and relationships to biophysical factors, forest history, ownership, and management practices in the Coastal Province of Oregon, USA Author: Kennedy, Rebecca S.H. Dead wood patterns and dynamics vary with biophysical factors, disturbance history, ownership, and management practices. Through field and modeling studies, I examined the current and potential future amounts of dead wood in two landscapes and region-wide in the Coastal Province of Oregon. The objectives of the first study were to (1) determine whether two landscapes with different recent disturbance histories differ in the amount and characteristics of dead wood; and (2) explore relationships between patterns of dead wood in each landscape to potentially related factors including topography. The objectives of the second study were to (1) describe current regional amounts of dead wood; (2) compare dead wood amounts across ownerships; (3) determine relationships between current dead wood amounts and ownership, current and past vegetation conditions, climate, topography, and soils; and (4) evaluate whether the factors related to dead wood patterns differed according to the scale of analysis. The objectives of the third study were to (1) characterize the projected future change in dead wood amounts in a multi-ownership Province; (2) determine the longevity of present-day dead wood of different types and sizes in relation to amendments from management and stand development; and (3) evaluate differences in management approaches in transitional dynamics and long-term patterns of dead wood. In the first study, I sampled logs and snags at four topographic positions (streams, lower slopes, middle slopes, upper slopes) in the Tillamook State Forest and the Siuslaw National Forest. These two landscapes experienced catastrophic fire at different points in recent history. I developed statistical models relating various attributes of dead wood abundance to biophysical variables related to climate, topography, historical vegetation, current vegetation, soils, and ecoregion. I found that the type and timing of disturbance was important to dead wood amounts and characteristics, and that potential source and sink areas for dead wood were related to topographic position. In particular, lower slopes had higher amounts of logs, and upper slopes had higher basal areas of potential source wood, in the form of snags and legacy (pre-fire) stumps. Climatic factors were of greater relative importance to overall gradients of dead wood in the landscape in which fire occurred less recently. In the second study, I analyzed dead wood data from a region-wide systematic grid of field plots according to ownership and biophysical variables at multiple scales of resolution including plots, subwatersheds. Dead wood abundance and types varied greatly among ownerships, with public lands (Forest Service, Bureau of Land Management, State of Oregon) typically having higher amounts of dead wood and more dead wood in the larger size classes than the private lands (forest industry, non-industrial private). I found that the relative influence of ownership, topography, current and historical vegetation, and climate varied with scale of resolution. Current vegetation was of greater relative importance at finer scales of plots and subwatersheds, whereas climate, topography, and historical vegetation were of greater relative importance at coarser scales of watersheds and subbasins. Ownership was important to overall dead wood gradients at all scales considered. In the third study, by simulating stand development and dead wood dynamics under various forest management scenarios over a 300-year period, I was able to examine the long-term effects of management on dead wood abundance in the Coastal Province. I estimated potential upper bounds for future dead wood amounts. Dead wood amounts increased over time on average across the Province, mainly because of policies on public lands, especially the federal lands under the Northwest Forest Plan. Forest industry, under the Oregon Forest Practices Act and assuming retention of all snags at harvest and thinning, maintained amounts of dead wood that were similar to present-day levels, but size classes shifted toward the smaller sizes as existing large legacy dead wood decomposed. Non-industrial private lands showed increases from very low present-day amounts of dead wood. Across the Province, legacy logs and snags remained present for over a century of the simulation period, and buffered effects of intensive management to dead wood amounts. Variation across landscapes in starting conditions meant that contrasting management approaches had differential effects on long-term dead wood dynamics depending on where they were applied. Current amounts of dead wood and live vegetation patterns in the Province resulted from historical fire and logging. Results of this simulation study indicate that recently established policies oriented toward dead wood production and retention, in the absence of fire or other large- or mid-scale disturbances, are likely to result in increases in dead wood amounts that greatly exceed present-day levels. My results suggest that dead wood patterns of abundance will continue to diverge according to land ownership and that management practices that foster dead wood creation are of increasing importance to the long-term abundance of large dead wood as legacy dead wood is lost through decomposition. Close

• 2825. [Article] The influence of climate change and restoration on stream temperature

  Water temperature is an essential property of a stream. Temperature regulates physical and biochemical processes in aquatic habitats. Various factors related to climatic conditions, landscape characteristics, ...

  Citation × Citation Citation Abstract Copy Title: The influence of climate change and restoration on stream temperature Author: Diabat, Mousa Water temperature is an essential property of a stream. Temperature regulates physical and biochemical processes in aquatic habitats. Various factors related to climatic conditions, landscape characteristics, and channel structure directly influence stream temperature. Numerous studies indicate that increased average air temperature during the past century has led to stream warming across the world. The trend of stream warming was also present in spring-fed watersheds, where summer flow has decreased. In addition, anthropogenic practices that alter the natural landscape and channel structure, such as forest management, agriculture, and mining contributed to stream warming. For example, deforested and unshaded stream reaches or dredged channels were warmer than shaded reaches and meandering streams. Stream temperatures in North American lotic habitats are of a specific concern due to their significant economic, cultural, and ecological value. With climate projections indicating that air temperature will only continue to rise throughout the 21st century, cold- or cool-water organisms, especially fishes, will be affected. Therefore, there is a strong need to better understand the impacts of changing climate, riparian landscape, and channel structure on a stream's heat budget. This may assist in restoring the historic thermal regime in impacted sites and mitigating the impacts of future climate change. This study looks into the relative influences of the different factors on a stream's heat budget with three manuscripts: one on stream temperature response to diel timing of air warming, one on stream temperature response to changes in air temperature, flow, and riparian vegetation, and one on stream temperature response to air warming and channel reconstruction. I used the software Heat Source version 8.05 to simulate stream temperature for all three analyses along the Middle Fork John Day River, Oregon USA. Two of the manuscripts were applied to an upper 37 km section of the Middle Fork John Day River (presented in chapter 2 and 3), where the third manuscript was applied to a 1.5-km section. The sensitivity analysis of stream temperature response to diel timing of air warming (Chapter 2: Diel Timing of Warmer Air under Climate Change Affects Magnitude, Timing, and Duration of Stream Temperature Change) was based on scenarios representing uniform air warming over the diel period, daytime warming, and nighttime warming. Uniform warming of air temperature is a simple representation of increases in the average daily or monthly temperatures generated by the 'delta method'. The delta method relies on adding a constant value to the air temperature time-series data. This constant value is the difference (delta) between base case average air temperatures and the projected one. Scenarios of daytime or nighttime warming represent conditions under which most of the warming of the air occurs during the daytime or the nighttime, respectively. I simulated the stream temperature response to warmer air conditions of +2 °C and +4 °C in daily average for all three cases of air warming conditions. The three cases of different diel distributions of air warming generated 7-day average daily maximum stream temperature (7DADM) increases of approximately +1.8 °C ± 0.1 °C at the downstream end of the study section relative to the base case. In most parts of the reach, the three distributions of air warming generated different ranges of stream temperatures, different 7DADM values, different durations of stream temperature changes, and different average daily temperatures. Changes of stream temperature were out of phase with imposed changes of air temperature. Therefore, nighttime warming of air temperatures would cause the greatest increase in maximum daily stream temperature, which typically occurs during the daytime. The sensitivity analysis of the relative influences of changes in air temperature, stream flow, and riparian vegetation on stream temperature (Chapter 3: Assessing Stream Temperature Response to Cumulative Influence of Changing Air Temperature, Flow, and Riparian Vegetation). This study summarized stream temperature simulation in 36 scenarios representing possible manifestations of 21st century climate conditions and land management strategies. In addition to existing conditions (base case) of flow, air temperature, and riparian vegetation, scenarios consisted of: two air temperature increases of 2 °C and 4 °C, two stream flow variations of +30% and -30%, three spatially uniform riparian vegetation conditions that create averages of effective shade 7%, 34%, and 79%, in addition to 14% for base case conditions. Results suggest that variation in riparian vegetation was the dominant factor influencing stream temperature because it regulates incoming shortwave radiation, the largest heat input to the stream, while variation in stream flow has a negligible influence. Results indicated that increasing the effective shade along the study section, particularly in the currently unshaded sections, could mitigate the influence of increasing air temperature, and would reduce stream temperature maxima below current values even under future climate conditions of warmer air. With the small influence it had, increasing stream flow reduced the 7DADM under low shade conditions. However, increasing stream flow showed counterintuitive results as it contributed to increasing stream temperature maxima when the stream was heavily shaded. The applied study examined the stream temperature response to restoration practices and their potential to mitigate the influence of warmer air conditions (Chapter 4: Estimating Stream Temperature Response to Restoring Channel and Riparian Vegetation and the Potential to Mitigate Warmer Air Conditions). This study focused on a 1.5 km section along the upper part of the Middle Fork John Day River that was modified due to past anthropogenic activities of mining for gold and timber harvest. Currently, the riparian vegetation of the study site is mostly shrubs and stands of short trees. Restoration designs call for the restoration of both the channel structure and replanting the riparian vegetation. Simulation results showed that the 7DADM was higher in the restored channel than the existing channel with both conditions of low and high effective shade conditions. However, a combined restoration practice of channel reconstruction and medium effective shade conditions reduced stream temperature maxima more than restoring riparian vegetation alone. In addition, results showed that restoring riparian vegetation was sufficient to mitigate the influence of warmer air on stream temperature, while restoring the channel alone is not. Heat budget analysis showed that heat accumulation during the daytime increased in the restored channel, which was longer, narrower, and deeper than the existing channel. It is important to emphasize that stream temperature is one of many goals that restoration activities aim to improve. Furthermore, differences in 7DADM among the different scenarios of restoration are negligible. Such small differences could hardly be measure. While this study examined a short section of 1.5 km, longer stream sections may increase the differences in 7DADM. Primary conclusions of this study are: 1) daily maxima of stream temperature will increase in response to increased air temperature regardless of the distribution of air warming during the diel cycle; 2) nighttime air warming caused a greater increase in stream temperature maximum than daytime warming; 3) riparian vegetation was the dominant factor on stream's heat budget, more than air temperature or stream flow; 4) restoring riparian vegetation mitigated the influence of warmer air; 5) restoring channel structure alone was not sufficient to lower temperature maxima; and 6) restoration project was most successful in improving degraded stream temperature when combined with channel reconstruction and improved riparian shade. Title: The influence of climate change and restoration on stream temperature Author: Diabat, Mousa Water temperature is an essential property of a stream. Temperature regulates physical and biochemical processes in aquatic habitats. Various factors related to climatic conditions, landscape characteristics, and channel structure directly influence stream temperature. Numerous studies indicate that increased average air temperature during the past century has led to stream warming across the world. The trend of stream warming was also present in spring-fed watersheds, where summer flow has decreased. In addition, anthropogenic practices that alter the natural landscape and channel structure, such as forest management, agriculture, and mining contributed to stream warming. For example, deforested and unshaded stream reaches or dredged channels were warmer than shaded reaches and meandering streams. Stream temperatures in North American lotic habitats are of a specific concern due to their significant economic, cultural, and ecological value. With climate projections indicating that air temperature will only continue to rise throughout the 21st century, cold- or cool-water organisms, especially fishes, will be affected. Therefore, there is a strong need to better understand the impacts of changing climate, riparian landscape, and channel structure on a stream's heat budget. This may assist in restoring the historic thermal regime in impacted sites and mitigating the impacts of future climate change. This study looks into the relative influences of the different factors on a stream's heat budget with three manuscripts: one on stream temperature response to diel timing of air warming, one on stream temperature response to changes in air temperature, flow, and riparian vegetation, and one on stream temperature response to air warming and channel reconstruction. I used the software Heat Source version 8.05 to simulate stream temperature for all three analyses along the Middle Fork John Day River, Oregon USA. Two of the manuscripts were applied to an upper 37 km section of the Middle Fork John Day River (presented in chapter 2 and 3), where the third manuscript was applied to a 1.5-km section. The sensitivity analysis of stream temperature response to diel timing of air warming (Chapter 2: Diel Timing of Warmer Air under Climate Change Affects Magnitude, Timing, and Duration of Stream Temperature Change) was based on scenarios representing uniform air warming over the diel period, daytime warming, and nighttime warming. Uniform warming of air temperature is a simple representation of increases in the average daily or monthly temperatures generated by the 'delta method'. The delta method relies on adding a constant value to the air temperature time-series data. This constant value is the difference (delta) between base case average air temperatures and the projected one. Scenarios of daytime or nighttime warming represent conditions under which most of the warming of the air occurs during the daytime or the nighttime, respectively. I simulated the stream temperature response to warmer air conditions of +2 °C and +4 °C in daily average for all three cases of air warming conditions. The three cases of different diel distributions of air warming generated 7-day average daily maximum stream temperature (7DADM) increases of approximately +1.8 °C ± 0.1 °C at the downstream end of the study section relative to the base case. In most parts of the reach, the three distributions of air warming generated different ranges of stream temperatures, different 7DADM values, different durations of stream temperature changes, and different average daily temperatures. Changes of stream temperature were out of phase with imposed changes of air temperature. Therefore, nighttime warming of air temperatures would cause the greatest increase in maximum daily stream temperature, which typically occurs during the daytime. The sensitivity analysis of the relative influences of changes in air temperature, stream flow, and riparian vegetation on stream temperature (Chapter 3: Assessing Stream Temperature Response to Cumulative Influence of Changing Air Temperature, Flow, and Riparian Vegetation). This study summarized stream temperature simulation in 36 scenarios representing possible manifestations of 21st century climate conditions and land management strategies. In addition to existing conditions (base case) of flow, air temperature, and riparian vegetation, scenarios consisted of: two air temperature increases of 2 °C and 4 °C, two stream flow variations of +30% and -30%, three spatially uniform riparian vegetation conditions that create averages of effective shade 7%, 34%, and 79%, in addition to 14% for base case conditions. Results suggest that variation in riparian vegetation was the dominant factor influencing stream temperature because it regulates incoming shortwave radiation, the largest heat input to the stream, while variation in stream flow has a negligible influence. Results indicated that increasing the effective shade along the study section, particularly in the currently unshaded sections, could mitigate the influence of increasing air temperature, and would reduce stream temperature maxima below current values even under future climate conditions of warmer air. With the small influence it had, increasing stream flow reduced the 7DADM under low shade conditions. However, increasing stream flow showed counterintuitive results as it contributed to increasing stream temperature maxima when the stream was heavily shaded. The applied study examined the stream temperature response to restoration practices and their potential to mitigate the influence of warmer air conditions (Chapter 4: Estimating Stream Temperature Response to Restoring Channel and Riparian Vegetation and the Potential to Mitigate Warmer Air Conditions). This study focused on a 1.5 km section along the upper part of the Middle Fork John Day River that was modified due to past anthropogenic activities of mining for gold and timber harvest. Currently, the riparian vegetation of the study site is mostly shrubs and stands of short trees. Restoration designs call for the restoration of both the channel structure and replanting the riparian vegetation. Simulation results showed that the 7DADM was higher in the restored channel than the existing channel with both conditions of low and high effective shade conditions. However, a combined restoration practice of channel reconstruction and medium effective shade conditions reduced stream temperature maxima more than restoring riparian vegetation alone. In addition, results showed that restoring riparian vegetation was sufficient to mitigate the influence of warmer air on stream temperature, while restoring the channel alone is not. Heat budget analysis showed that heat accumulation during the daytime increased in the restored channel, which was longer, narrower, and deeper than the existing channel. It is important to emphasize that stream temperature is one of many goals that restoration activities aim to improve. Furthermore, differences in 7DADM among the different scenarios of restoration are negligible. Such small differences could hardly be measure. While this study examined a short section of 1.5 km, longer stream sections may increase the differences in 7DADM. Primary conclusions of this study are: 1) daily maxima of stream temperature will increase in response to increased air temperature regardless of the distribution of air warming during the diel cycle; 2) nighttime air warming caused a greater increase in stream temperature maximum than daytime warming; 3) riparian vegetation was the dominant factor on stream's heat budget, more than air temperature or stream flow; 4) restoring riparian vegetation mitigated the influence of warmer air; 5) restoring channel structure alone was not sufficient to lower temperature maxima; and 6) restoration project was most successful in improving degraded stream temperature when combined with channel reconstruction and improved riparian shade. Close

• 2826. [Article] Hood River Bull Trout Abundance, Life History, and Habitat Connectivity, 2007 Progress Reports 2007

  Abstract -- Hood River bull trout are thought to exist as two independent reproductive units (USFWS 2004), known as local populations (Rieman and McIntyre 1995). The Clear Branch local population is isolated ...

  Citation × Citation Citation Abstract Copy Title: Hood River Bull Trout Abundance, Life History, and Habitat Connectivity, 2007 Progress Reports 2007 Abstract -- Hood River bull trout are thought to exist as two independent reproductive units (USFWS 2004), known as local populations (Rieman and McIntyre 1995). The Clear Branch local population is isolated above Clear Branch Dam, which provides limited downstream fish passage during infrequent and sporadic periods of spill and no upstream passage. Bull trout in this population inhabit Laurance Lake Reservoir and tributaries upstream of Clear Branch Dam. The Hood River local population occurs in the mainstem Hood River and Middle Fork Hood River downstream of the Clear Branch Dam and a small number of adult bull trout migrate each year into the Hood River from the Columbia River (Figure 1). The status of both populations is extremely precarious. The Clear Branch population is at risk of a random extinction event due to low numbers, negative interactions with non-native smallmouth bass, isolation and limited spawning habitat (USFWS, 1998). The Hood River population also appears to be small and is threatened by passage barriers, unscreened irrigation systems, impaired water quality and periodic siltation of spawning substrate by glacial outbursts. Clear Branch bull trout spawn in Clear Branch and Pinnacle Creek. After rearing in these two natal streams for an unknown time period, most are believed to migrate downstream to Laurance Lake Reservoir. Clear Branch bull trout have been documented passing over the dam spillway during high water events (Pribyl et al. 1996) and may provide a recruitment source for the Hood River local population. Adult bull trout tagged at Powerdale Dam have been observed at Coe Branch irrigation diversion and in a trap at the base of Clear Branch dam. These fish may have been attempting to reach spawning areas located upstream of the dam. However, the success of bull trout migrating downstream via the spillway or the possibility of successfully navigating through the diversion network has never been determined. Depending on the water year, the Middle Fork Irrigation District (MFID) may not spill at all, or the timing of the spill may not coincide with the timing of downstream migration, which is currently unknown (East Fork Hood River and Middle Fork Hood River Watershed analysis). Smallmouth bass were discovered in Lake Laurance Reservoir in the 1990s. Creel surveys have shown that large adult bass are caught occasionally in the reservoir and schools of bass fry have been seen by district fish biologist (Rod French, ODFW, personal communication), suggesting that they are spawning successfully. This illegal introduction poses a potential threat to the Clear Branch bull trout population, but its magnitude is unknown because the bass population size and degree of interaction between the two species are unknown. Bull trout and smallmouth bass have significantly different temperature preferences and tolerances, with bull trout being one of the most sensitive coldwater species and bass being a warm water species. Lake Laurance, a relatively high-altitude reservoir at 890 m (2,920 feet), does not provide ideal bass habitat so these two species may have largely non-overlapping distributions or differing activity periods (Terry Shrader, ODFW warmwater fish biologist, personal communication). However, based on past reservoir temperature data (Berger et al. 2005), there are periods in the reservoir when there is potential for bull trout and bass interaction: periods when bull trout are susceptible to bass predation and when juvenile fish might compete for resources. Spawning activity of the Hood River local population has been observed in a few locations within the Middle Fork of Hood River (Figure 1). Although consistent and extensive spawning areas for this population are not known, some of the locations where juvenile rearing or potential bull trout redds have been observed include the Middle Fork Hood River and some of its tributaries: Bear Creek, Compass Creek and Coe Branch (USFWS 2004). However, Coe Branch, Compass Creek, and the Middle Fork are glacial streams with a high volume of sand and silt which may compromise spawning success. No bull trout spawning or rearing has been observed on the East and West Forks of Hood River. The Middle Fork and mainstem Hood River provide foraging, migration and overwintering habitat. Hood River bull trout are also known to migrate into the Columbia River. Two bull trout tagged at Powerdale Dam (RK 7.2 of mainstem Hood River) were recovered near Drano Lake in Washington State; and one was captured 11 kilometers downstream of the confluence of the Hood and Columbia Rivers (USFWS 2004). Every year (usually between May and July), adult bull trout, presumably migrating upstream from the Columbia River, are captured and anchor tagged at Powerdale Dam. Although some of these tagged fish have been observed upstream (one in Coe Branch and three below Clear Branch dam), the spawning destination of fluvial adults within the Hood River basin is largely unknown. Dispersing juvenile bull trout and migrating adults in this local population are threatened by flow diversions with inadequate screening and passage facilities. Several structures are suspected to impede upstream migration or entrain juvenile and adult bull trout into irrigation works (Pribyl et al. 1996, HRWG 1999). These structures include: the diversion at Clear Branch Dam (passage and screening), Coe Branch (passage and screening), and the Farmers Irrigation District diversion (screening) on the mainstem Hood River (HRWG 1999). However, little research has been conducted to assess the impacts of these structures on migrating bull trout. Beyond a general knowledge of the distribution of Hood River bull trout and the nature of anthropogenic factors that potentially restrict their life history and habitat connectivity, little is known about this recovery unit. Baseline information about adult abundance is lacking for both local populations, the potential of a source (Clear Branch) and sink (Hood River) relationship between the two local populations has not been explored, and the migratory life history of adult fish caught at Powerdale Dam is unknown. The degree to which irrigation and hydropower diversions hamper connectivity within the Hood River basin is also poorly understood. Migratory life histories have been viewed as key to species persistence (Rieman and McIntyre 1995; Dunham and Rieman 1999), and understanding movement patterns and associated habitat requirements are critical to maintaining those migratory forms (Muhlfeld and Morotz 2005; Hostettler 2005). Gaining this information is also critical to evaluating bull trout recovery in the Hood River Subbasin (Coccoli 2004). The Oregon Department of Fish and Wildlife (ODFW) initiated a study in 2006 to improve our understanding of the abundance, life history, and potential limiting factors of the bull trout in this recovery unit. This report describes findings for the first two years of the study (2006-2007). Specific study objectives for the first two years were: 1. Determine the migratory life history of Hood River bull trout and assess the potential impacts of flow diversions and two new falls on the Middle Fork Hood River (scoured by the November 2006 glacial outburst) on bull trout migrations. 2. Determine current distribution of bull trout reproduction and early rearing in historical and potential bull trout streams in the Hood River Subbasin. 3. Determine the juvenile and adult life history the Clear Branch local population and develop a statistically reliable and cost-effective protocol for monitoring the abundance of adult Clear Branch bull trout. 4. Assess the potential impact of smallmouth bass on bull trout in Laurance Lake Reservoir. Title: Hood River Bull Trout Abundance, Life History, and Habitat Connectivity, 2007 Progress Reports 2007 Abstract -- Hood River bull trout are thought to exist as two independent reproductive units (USFWS 2004), known as local populations (Rieman and McIntyre 1995). The Clear Branch local population is isolated above Clear Branch Dam, which provides limited downstream fish passage during infrequent and sporadic periods of spill and no upstream passage. Bull trout in this population inhabit Laurance Lake Reservoir and tributaries upstream of Clear Branch Dam. The Hood River local population occurs in the mainstem Hood River and Middle Fork Hood River downstream of the Clear Branch Dam and a small number of adult bull trout migrate each year into the Hood River from the Columbia River (Figure 1). The status of both populations is extremely precarious. The Clear Branch population is at risk of a random extinction event due to low numbers, negative interactions with non-native smallmouth bass, isolation and limited spawning habitat (USFWS, 1998). The Hood River population also appears to be small and is threatened by passage barriers, unscreened irrigation systems, impaired water quality and periodic siltation of spawning substrate by glacial outbursts. Clear Branch bull trout spawn in Clear Branch and Pinnacle Creek. After rearing in these two natal streams for an unknown time period, most are believed to migrate downstream to Laurance Lake Reservoir. Clear Branch bull trout have been documented passing over the dam spillway during high water events (Pribyl et al. 1996) and may provide a recruitment source for the Hood River local population. Adult bull trout tagged at Powerdale Dam have been observed at Coe Branch irrigation diversion and in a trap at the base of Clear Branch dam. These fish may have been attempting to reach spawning areas located upstream of the dam. However, the success of bull trout migrating downstream via the spillway or the possibility of successfully navigating through the diversion network has never been determined. Depending on the water year, the Middle Fork Irrigation District (MFID) may not spill at all, or the timing of the spill may not coincide with the timing of downstream migration, which is currently unknown (East Fork Hood River and Middle Fork Hood River Watershed analysis). Smallmouth bass were discovered in Lake Laurance Reservoir in the 1990s. Creel surveys have shown that large adult bass are caught occasionally in the reservoir and schools of bass fry have been seen by district fish biologist (Rod French, ODFW, personal communication), suggesting that they are spawning successfully. This illegal introduction poses a potential threat to the Clear Branch bull trout population, but its magnitude is unknown because the bass population size and degree of interaction between the two species are unknown. Bull trout and smallmouth bass have significantly different temperature preferences and tolerances, with bull trout being one of the most sensitive coldwater species and bass being a warm water species. Lake Laurance, a relatively high-altitude reservoir at 890 m (2,920 feet), does not provide ideal bass habitat so these two species may have largely non-overlapping distributions or differing activity periods (Terry Shrader, ODFW warmwater fish biologist, personal communication). However, based on past reservoir temperature data (Berger et al. 2005), there are periods in the reservoir when there is potential for bull trout and bass interaction: periods when bull trout are susceptible to bass predation and when juvenile fish might compete for resources. Spawning activity of the Hood River local population has been observed in a few locations within the Middle Fork of Hood River (Figure 1). Although consistent and extensive spawning areas for this population are not known, some of the locations where juvenile rearing or potential bull trout redds have been observed include the Middle Fork Hood River and some of its tributaries: Bear Creek, Compass Creek and Coe Branch (USFWS 2004). However, Coe Branch, Compass Creek, and the Middle Fork are glacial streams with a high volume of sand and silt which may compromise spawning success. No bull trout spawning or rearing has been observed on the East and West Forks of Hood River. The Middle Fork and mainstem Hood River provide foraging, migration and overwintering habitat. Hood River bull trout are also known to migrate into the Columbia River. Two bull trout tagged at Powerdale Dam (RK 7.2 of mainstem Hood River) were recovered near Drano Lake in Washington State; and one was captured 11 kilometers downstream of the confluence of the Hood and Columbia Rivers (USFWS 2004). Every year (usually between May and July), adult bull trout, presumably migrating upstream from the Columbia River, are captured and anchor tagged at Powerdale Dam. Although some of these tagged fish have been observed upstream (one in Coe Branch and three below Clear Branch dam), the spawning destination of fluvial adults within the Hood River basin is largely unknown. Dispersing juvenile bull trout and migrating adults in this local population are threatened by flow diversions with inadequate screening and passage facilities. Several structures are suspected to impede upstream migration or entrain juvenile and adult bull trout into irrigation works (Pribyl et al. 1996, HRWG 1999). These structures include: the diversion at Clear Branch Dam (passage and screening), Coe Branch (passage and screening), and the Farmers Irrigation District diversion (screening) on the mainstem Hood River (HRWG 1999). However, little research has been conducted to assess the impacts of these structures on migrating bull trout. Beyond a general knowledge of the distribution of Hood River bull trout and the nature of anthropogenic factors that potentially restrict their life history and habitat connectivity, little is known about this recovery unit. Baseline information about adult abundance is lacking for both local populations, the potential of a source (Clear Branch) and sink (Hood River) relationship between the two local populations has not been explored, and the migratory life history of adult fish caught at Powerdale Dam is unknown. The degree to which irrigation and hydropower diversions hamper connectivity within the Hood River basin is also poorly understood. Migratory life histories have been viewed as key to species persistence (Rieman and McIntyre 1995; Dunham and Rieman 1999), and understanding movement patterns and associated habitat requirements are critical to maintaining those migratory forms (Muhlfeld and Morotz 2005; Hostettler 2005). Gaining this information is also critical to evaluating bull trout recovery in the Hood River Subbasin (Coccoli 2004). The Oregon Department of Fish and Wildlife (ODFW) initiated a study in 2006 to improve our understanding of the abundance, life history, and potential limiting factors of the bull trout in this recovery unit. This report describes findings for the first two years of the study (2006-2007). Specific study objectives for the first two years were: 1. Determine the migratory life history of Hood River bull trout and assess the potential impacts of flow diversions and two new falls on the Middle Fork Hood River (scoured by the November 2006 glacial outburst) on bull trout migrations. 2. Determine current distribution of bull trout reproduction and early rearing in historical and potential bull trout streams in the Hood River Subbasin. 3. Determine the juvenile and adult life history the Clear Branch local population and develop a statistically reliable and cost-effective protocol for monitoring the abundance of adult Clear Branch bull trout. 4. Assess the potential impact of smallmouth bass on bull trout in Laurance Lake Reservoir. Close