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• 1. [Article] Restoring oak habitats in the Southern Willamette Valley, Oregon: A multi-objective tradeoffs analysis for landowners and managers

  xvii, 160 p. : ill. (some col.) A print copy of this thesis is available through the UO Libraries. Search the library catalog for the location and call number.

  Citation × Citation Citation Abstract Copy Title: Restoring oak habitats in the Southern Willamette Valley, Oregon: A multi-objective tradeoffs analysis for landowners and managers Author: Ulrich, Nathan D., 1977- Year: 2011, 2010 xvii, 160 p. : ill. (some col.) A print copy of this thesis is available through the UO Libraries. Search the library catalog for the location and call number. Title: Restoring oak habitats in the Southern Willamette Valley, Oregon: A multi-objective tradeoffs analysis for landowners and managers Author: Ulrich, Nathan D., 1977- Year: 2011, 2010 xvii, 160 p. : ill. (some col.) A print copy of this thesis is available through the UO Libraries. Search the library catalog for the location and call number. Close

• 2. [Article] Final report: Predicting field establishment rates from standardized plant traits - Year two of: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management

  Willamette Valley wetland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species (Wilson et al. 1993, Noss et al. 1995, USFWS 2000, ...

  Citation × Citation Citation Abstract Copy Title: Final report: Predicting field establishment rates from standardized plant traits - Year two of: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management Author: Wilson, Mark V., Roberts, Rachel E., Clark, Deborah Louise, 1950- Willamette Valley wetland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species (Wilson et al. 1993, Noss et al. 1995, USFWS 2000, Schultz et al. 2003). A rigorous program of ecosystem restoration can protect and enhance wetland plants, animals, and services (Pywell and Putwain 1996). The restoration projects of the West Eugene Wetlands Program (WEWP) comprise one of the few large-scale and long-term integrated restoration programs in the world. Successful ecosystem restoration requires establishing and maintaining native plants. In turn, plant establishment hinges on having suitable environmental conditions, using species with adequate germination and growth rates, and reducing competitive pressure from non-native plants (Figure 1). In year one of this project, we synthesized the wealth of plant establishment data during wetland restoration in the West Eugene Wetlands Program (Wilson 2004). In year two, our objectives were to expand on these results in several important ways: • Generalize these results through the investigation of plant traits (Table 1) that consistently correspond to the patterns of establishment and vigor (Figure 1). • Systematically compile the results into a trait database. This database includes findings from similar ecosystems, both in the Willamette Valley and elsewhere. • Testing the effect of habitat variation on the relationship between traits and seedling establishment patterns. Our goal is to predict key aspects of prairie restoration performance, in this case establishment rates, based on species traits and habitat. These predictions can then be applied as management recommendations, such as which species to sow to maximize native plant abundance at a given site, even if the species have not been field tested. The two components of our project–plant traits and the database–are crucial to this goal. • Without the generalization that traits allow, understanding of wetland restoration increases slowly and expensively, one case study at a time. • The organization of the database will increase the power and efficiency of revealing the relationships between plant traits and plant performance. Perhaps even more important is the role of the database in developing a Web-based expert system for managers wishing to plan wetland restorations. Title: Final report: Predicting field establishment rates from standardized plant traits - Year two of: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management Author: Wilson, Mark V., Roberts, Rachel E., Clark, Deborah Louise, 1950- Willamette Valley wetland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species (Wilson et al. 1993, Noss et al. 1995, USFWS 2000, Schultz et al. 2003). A rigorous program of ecosystem restoration can protect and enhance wetland plants, animals, and services (Pywell and Putwain 1996). The restoration projects of the West Eugene Wetlands Program (WEWP) comprise one of the few large-scale and long-term integrated restoration programs in the world. Successful ecosystem restoration requires establishing and maintaining native plants. In turn, plant establishment hinges on having suitable environmental conditions, using species with adequate germination and growth rates, and reducing competitive pressure from non-native plants (Figure 1). In year one of this project, we synthesized the wealth of plant establishment data during wetland restoration in the West Eugene Wetlands Program (Wilson 2004). In year two, our objectives were to expand on these results in several important ways: • Generalize these results through the investigation of plant traits (Table 1) that consistently correspond to the patterns of establishment and vigor (Figure 1). • Systematically compile the results into a trait database. This database includes findings from similar ecosystems, both in the Willamette Valley and elsewhere. • Testing the effect of habitat variation on the relationship between traits and seedling establishment patterns. Our goal is to predict key aspects of prairie restoration performance, in this case establishment rates, based on species traits and habitat. These predictions can then be applied as management recommendations, such as which species to sow to maximize native plant abundance at a given site, even if the species have not been field tested. The two components of our project–plant traits and the database–are crucial to this goal. • Without the generalization that traits allow, understanding of wetland restoration increases slowly and expensively, one case study at a time. • The organization of the database will increase the power and efficiency of revealing the relationships between plant traits and plant performance. Perhaps even more important is the role of the database in developing a Web-based expert system for managers wishing to plan wetland restorations. Close

• 3. [Article] Progress report II: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management (Year two)

  Willamette Valley wetland and upland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species. Because these are dynamic systems, ...

  Citation × Citation Citation Abstract Copy Title: Progress report II: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management (Year two) Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Willamette Valley wetland and upland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species. Because these are dynamic systems, subject to ecological succession and invasion by aggressive non-native species, prairies require active management to maintain management goals. This is true of all Willamette Valley prairies, whether intact, degraded, or reestablished. Yet restoration and conservation efforts often lack experimental validation. Our overall objective for this project is to synthesize the scientific information now available relevant to the prairie restoration efforts of the West Eugene Wetlands project and, where there is sufficient support, develop concrete and defensible management recommendations Successful ecosystem restoration requires establishing and maintaining native plants. In turn, plant establishment hinges on having suitable environmental conditions, using species with adequate germination rates, and reducing competitive pressure from nonnative plants. In year one of this project, we synthesized the wealth of data in the West Eugene Wetlands establishment data set on plant abundance after sowing native species during wetland restoration (Wilson 2004). In year two, we will build and expand on these results in several important ways: • We are generalizing these results through the investigation of plant traits that consistently correspond to the patterns of establishment and vigor. • We are systematically compiling the results from year one and year two of this project into a public database. We are adding to this database findings from similar ecosystems, both in the Willamette Valley and elsewhere. • We are considering further the role of microsite variability on seedling establishment patterns. • We are synthesizing these results into scientific conclusions. • We are integrating these results, where there is sufficient support, into concrete and defensible management recommendations Our goal is to develop an ability to predict key aspects of prairie restoration performance, such as establishment rates, based on knowing species traits, site conditions, and maintenance. These predictions can then be converted into management recommendations, such as which species to sow and which site preparation and maintenance regimes should be followed to maximize native plant abundance and minimize non-native plant abundance at a given site. The two components of our project –plant traits and the database– are crucial to this goal. • Without the generalization that traits allow, understanding of wetland restoration increases slowly and expensively, one case study at a time. • The organization of the database will increase the power and efficiency of revealing the relationships between plant traits and plant performance. Perhaps even more important is the role of the database as a first step in developing a Web-based expert system for managers wishing to plan wetland restorations Title: Progress report II: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management (Year two) Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Willamette Valley wetland and upland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species. Because these are dynamic systems, subject to ecological succession and invasion by aggressive non-native species, prairies require active management to maintain management goals. This is true of all Willamette Valley prairies, whether intact, degraded, or reestablished. Yet restoration and conservation efforts often lack experimental validation. Our overall objective for this project is to synthesize the scientific information now available relevant to the prairie restoration efforts of the West Eugene Wetlands project and, where there is sufficient support, develop concrete and defensible management recommendations Successful ecosystem restoration requires establishing and maintaining native plants. In turn, plant establishment hinges on having suitable environmental conditions, using species with adequate germination rates, and reducing competitive pressure from nonnative plants. In year one of this project, we synthesized the wealth of data in the West Eugene Wetlands establishment data set on plant abundance after sowing native species during wetland restoration (Wilson 2004). In year two, we will build and expand on these results in several important ways: • We are generalizing these results through the investigation of plant traits that consistently correspond to the patterns of establishment and vigor. • We are systematically compiling the results from year one and year two of this project into a public database. We are adding to this database findings from similar ecosystems, both in the Willamette Valley and elsewhere. • We are considering further the role of microsite variability on seedling establishment patterns. • We are synthesizing these results into scientific conclusions. • We are integrating these results, where there is sufficient support, into concrete and defensible management recommendations Our goal is to develop an ability to predict key aspects of prairie restoration performance, such as establishment rates, based on knowing species traits, site conditions, and maintenance. These predictions can then be converted into management recommendations, such as which species to sow and which site preparation and maintenance regimes should be followed to maximize native plant abundance and minimize non-native plant abundance at a given site. The two components of our project –plant traits and the database– are crucial to this goal. • Without the generalization that traits allow, understanding of wetland restoration increases slowly and expensively, one case study at a time. • The organization of the database will increase the power and efficiency of revealing the relationships between plant traits and plant performance. Perhaps even more important is the role of the database as a first step in developing a Web-based expert system for managers wishing to plan wetland restorations Close

• 4. [Article] Progress report [I]: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management (Year two)

  Willamette Valley wetland and upland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species. These are dynamic systems, subject ...

  Citation × Citation Citation Abstract Copy Title: Progress report [I]: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management (Year two) Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Willamette Valley wetland and upland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species. These are dynamic systems, subject to ecological succession and invasion by aggressive non-native species, and require active management to maintain management goals. This is true of all Willamette Valley prairies, whether intact, degraded, or reestablished. Yet restoration and conservation efforts often lack experimental validation. Our overall objective is to synthesize the scientific information now available pertinent to the prairie restoration efforts of the West Eugene Wetlands Program (WEWP) and, where there is sufficient support, develop concrete and defensible management recommendations. Successful ecosystem restoration requires establishing and maintaining native plants. In turn, plant establishment hinges on having suitable environmental conditions, using species with adequate germination and growth rates, and reducing competitive pressure from non-native plants (Figure 1). In year one of this project, we synthesized the wealth of plant establishment data during wetland restoration in the West Eugene Wetlands Program. In year two, we are building and expanding on these results in several important ways: • We are generalizing these results through the investigation of plant traits (Table 1) that consistently correspond to the patterns of establishment and vigor (Figure 1). • We are systematically compiling the results from year one and year two of this project into a public database. We are adding to this database findings from similar ecosystems, both in the Willamette Valley and elsewhere. • We are considering further the role of microsite variability on seedling establishment patterns. • We are synthesizing these results into scientific conclusions. • We are integrating these results, where there is sufficient support, into concrete and defensible management recommendations Our goal is to develop an ability to predict key aspects of prairie restoration performance, such as establishment rates, based on species traits, site conditions, and maintenance. These predictions can then be restated as management recommendations, such as which species to sow and site preparation and maintenance regimes to follow to maximize native plant abundance and minimize non-native plant abundance at a given site. The two components of our project–plant traits and the database–are crucial to this goal. • Without the generalization that traits allow, understanding of wetland restoration increases slowly and expensively, one case study at a time. • The organization of the database will increase the power and efficiency of revealing the relationships between plant traits and plant performance. Perhaps even more important is the role of the database as a first step in developing a Web-based expert system for managers wishing to plan wetland restorations. Title: Progress report [I]: Restoring prairies: A synthesis of studies on vegetation and invasive species in support of effective management (Year two) Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Willamette Valley wetland and upland prairies are some of the most endangered ecosystems in the United States, and provide habitat for many federally listed species. These are dynamic systems, subject to ecological succession and invasion by aggressive non-native species, and require active management to maintain management goals. This is true of all Willamette Valley prairies, whether intact, degraded, or reestablished. Yet restoration and conservation efforts often lack experimental validation. Our overall objective is to synthesize the scientific information now available pertinent to the prairie restoration efforts of the West Eugene Wetlands Program (WEWP) and, where there is sufficient support, develop concrete and defensible management recommendations. Successful ecosystem restoration requires establishing and maintaining native plants. In turn, plant establishment hinges on having suitable environmental conditions, using species with adequate germination and growth rates, and reducing competitive pressure from non-native plants (Figure 1). In year one of this project, we synthesized the wealth of plant establishment data during wetland restoration in the West Eugene Wetlands Program. In year two, we are building and expanding on these results in several important ways: • We are generalizing these results through the investigation of plant traits (Table 1) that consistently correspond to the patterns of establishment and vigor (Figure 1). • We are systematically compiling the results from year one and year two of this project into a public database. We are adding to this database findings from similar ecosystems, both in the Willamette Valley and elsewhere. • We are considering further the role of microsite variability on seedling establishment patterns. • We are synthesizing these results into scientific conclusions. • We are integrating these results, where there is sufficient support, into concrete and defensible management recommendations Our goal is to develop an ability to predict key aspects of prairie restoration performance, such as establishment rates, based on species traits, site conditions, and maintenance. These predictions can then be restated as management recommendations, such as which species to sow and site preparation and maintenance regimes to follow to maximize native plant abundance and minimize non-native plant abundance at a given site. The two components of our project–plant traits and the database–are crucial to this goal. • Without the generalization that traits allow, understanding of wetland restoration increases slowly and expensively, one case study at a time. • The organization of the database will increase the power and efficiency of revealing the relationships between plant traits and plant performance. Perhaps even more important is the role of the database as a first step in developing a Web-based expert system for managers wishing to plan wetland restorations. Close

• 5. [Article] Traits, neighbors, and species performance in prairie restoration

  Questions: Are traits related to the performance of plant species in restoration? Are the relationships between traits and performance consistent across the functional groups of annual forbs, perennial ...

  Citation × Citation Citation Abstract Copy Title: Traits, neighbors, and species performance in prairie restoration Author: Roberts, R. E., Wilson, Mark V., Clark, Deborah Louise, 1950- Questions: Are traits related to the performance of plant species in restoration? Are the relationships between traits and performance consistent across the functional groups of annual forbs, perennial forbs, and grasses? Do the relationships between traits and performance depend on neighboring functional groups? Location: A former agricultural field, being restored to native upland prairie, in the Willamette Valley of western Oregon USA. Methods: Twenty-eight native species, representing three functional groups, were sown in seven different combinations. The following summer the performance of each species was measured. Eleven functional traits were measured from plants in the laboratory and in the field. Correlations between individual traits and performance variables were measured and stepwise regression techniques were used to determine which sets of traits were most strongly related to performance. Results: Sets of traits explained up to 56% of variation in cover, and up to 48% of variation in establishment frequency. The relationships between traits and performance were influenced by functional group identity, and the functional group identity of neighboring species also influenced species’ cover and the relationships between traits and cover. Species’ establishment rate in monoculture was the trait most strongly correlated to both establishment and cover in mixtures. In multi-trait models, annual forb functional group identity was strongly related to establishment in mixtures, and height, leaf weight ratio at 7 days, and seed mass were strongly related to cover. Conclusions: Based on the strength of the statistical models, multiple-trait models should be a useful way of predicting the performance of species prior to sowing in restoration. The functional group identity of each species as well as the other species being sown may need to be taken into account when making predictions. Title: Traits, neighbors, and species performance in prairie restoration Author: Roberts, R. E., Wilson, Mark V., Clark, Deborah Louise, 1950- Questions: Are traits related to the performance of plant species in restoration? Are the relationships between traits and performance consistent across the functional groups of annual forbs, perennial forbs, and grasses? Do the relationships between traits and performance depend on neighboring functional groups? Location: A former agricultural field, being restored to native upland prairie, in the Willamette Valley of western Oregon USA. Methods: Twenty-eight native species, representing three functional groups, were sown in seven different combinations. The following summer the performance of each species was measured. Eleven functional traits were measured from plants in the laboratory and in the field. Correlations between individual traits and performance variables were measured and stepwise regression techniques were used to determine which sets of traits were most strongly related to performance. Results: Sets of traits explained up to 56% of variation in cover, and up to 48% of variation in establishment frequency. The relationships between traits and performance were influenced by functional group identity, and the functional group identity of neighboring species also influenced species’ cover and the relationships between traits and cover. Species’ establishment rate in monoculture was the trait most strongly correlated to both establishment and cover in mixtures. In multi-trait models, annual forb functional group identity was strongly related to establishment in mixtures, and height, leaf weight ratio at 7 days, and seed mass were strongly related to cover. Conclusions: Based on the strength of the statistical models, multiple-trait models should be a useful way of predicting the performance of species prior to sowing in restoration. The functional group identity of each species as well as the other species being sown may need to be taken into account when making predictions. Close

• 6. [Article] Final Report 1994-1997: Effects of Fire, Mowing, and Mowing with Herbicide on Native Prairie of Baskett Butte, Baskett Slough NWR

  We report results from a field experiment evaluating the effectiveness of mowing, mowing and herbicides, and burning on woody plants, key native and non-native grasses and forbs, and the Fender’s blue ...

  Citation × Citation Citation Abstract Copy Title: Final Report 1994-1997: Effects of Fire, Mowing, and Mowing with Herbicide on Native Prairie of Baskett Butte, Baskett Slough NWR Author: Wilson, Mark V., Clark, Deborah Louise, 1950- We report results from a field experiment evaluating the effectiveness of mowing, mowing and herbicides, and burning on woody plants, key native and non-native grasses and forbs, and the Fender’s blue butterfly. The goal was to find a technique that controlled woody pest plants without harming native species or promoting non-native species. Three years after the initial manipulations, all treatments significantly reduced woody plant cover. Mowing promoted the group of measured native grasses without promoting non-native species. In contrast, burning had mixed effects on herbaceous species, promoting growth for some species and decreasing growth for others. Each treatment, particularly mowing, caused local increases in numbers of Fender’s blue butterfly egg masses, probably because lupines invigorated by treatments attracted ovipositing females. Although prescribed burning seemed to kill larvae, treated areas still accumulated more eggs masses than in controls. Three years after the start of manipulations, mowing was the most effective overall treatment for controlling woody plants, increasing native species abundance without promoting non-native species, and increasing Fender’s blue butterfly numbers. We recommend mowing for restoring similar habitats. Prescribed burning would also be an effective choice for habitat restoration. Title: Final Report 1994-1997: Effects of Fire, Mowing, and Mowing with Herbicide on Native Prairie of Baskett Butte, Baskett Slough NWR Author: Wilson, Mark V., Clark, Deborah Louise, 1950- We report results from a field experiment evaluating the effectiveness of mowing, mowing and herbicides, and burning on woody plants, key native and non-native grasses and forbs, and the Fender’s blue butterfly. The goal was to find a technique that controlled woody pest plants without harming native species or promoting non-native species. Three years after the initial manipulations, all treatments significantly reduced woody plant cover. Mowing promoted the group of measured native grasses without promoting non-native species. In contrast, burning had mixed effects on herbaceous species, promoting growth for some species and decreasing growth for others. Each treatment, particularly mowing, caused local increases in numbers of Fender’s blue butterfly egg masses, probably because lupines invigorated by treatments attracted ovipositing females. Although prescribed burning seemed to kill larvae, treated areas still accumulated more eggs masses than in controls. Three years after the start of manipulations, mowing was the most effective overall treatment for controlling woody plants, increasing native species abundance without promoting non-native species, and increasing Fender’s blue butterfly numbers. We recommend mowing for restoring similar habitats. Prescribed burning would also be an effective choice for habitat restoration. Close

• 7. [Article] Plant traits – a tool for restoration?

  Question: Most results of restoration efforts are species-specific and/or site-specific and therefore are not general enough to be easily applied to other species and other sites. Our research addresses ...

  Citation × Citation Citation Abstract Copy Title: Plant traits – a tool for restoration? Author: Wilson, Mark V., Kaye, Thomas N., Roberts, Rachel, Dunwiddie, Peter W., Clark, Deborah Louise, 1950-, Stanley, Amanda Question: Most results of restoration efforts are species-specific and/or site-specific and therefore are not general enough to be easily applied to other species and other sites. Our research addresses the issue of species-specific results by investigating the feasibility of using plant traits instead of taxonomic species to characterize species responses to restoration treatments. Specifically, we test the explanatory power of plant traits, one necessary prerequisite for the development of predictive and general plant trait models. Location: Ten remnant bunchgrass prairie sites in the Pacific Northwest of North America (Oregon and Washington, USA; British Columbia, Canada). Methods: We developed two types of quantitative models for each of 10 prairie restoration sites: 1) plant trait models, which related plant traits to species field responses following restoration management treatments, and 2) species identity models, which related species taxonomic identity to species field responses following restoration management treatments. Species identity models determined the maximum amount of variability of field responses that can be explained by differences in individual species’ responses to management treatments. Plant trait models determined what proportion of this explanatory power can be attributed to plant traits. This approach contrasts with approaches often used in other plant trait studies that describe how traits vary with environmental conditions. Specifically, we used these two models to address the following questions: 1) How much of the variability in field responses of plants to restoration management treatments is explained by plant traits? 2) How well do plant traits explain the variability of field responses following restoration management treatments compared to models relating field responses to species identity? Our approach was to measure two aspects of explanatory power: R2 (variability explained) and AIC (a measure of model fit that accounts for parsimony, i.e., how well a model fits the data with relatively few explanatory variables). Results: 1) The plant trait models (relating plant traits with plant field responses) explained much of the variability within each of the ten restoration sites, with R2 values ranging between 31% and 69%. 2) The species identity models (relating species taxonomic identity with plant field responses) explained between 47% and 74% of variability in field performance (R2). Thus, the plant trait models explained nearly as much variability as the species identity models. In seven out of nine sites, the plant trait models were superior to the species identity models as measured by AIC; that is, the trait models did well at explaining variability with less model complexity (i.e., fewer explanatory variables). Conclusion: Development of general and predictive plant trait models is a multi-step process. Strong explanatory power by plant trait models, both on an absolute scale and as compared to species identity models, supports the feasibility of using plant traits instead of species taxonomic identity as a common language to characterize plant field responses to restoration treatments. Such high explanatory power is one necessary pre-requisite for the development of predictive and general plant trait models. Our results also indicate that the plant trait models are site-specific even though all sites were upland bunchgrass remnant prairies. We discuss the next steps in the development of more general and predictive models: incorporating environmental factors into the plant trait models to address the issue of site-specificity and testing the power of these models to predict vegetation responses. Title: Plant traits – a tool for restoration? Author: Wilson, Mark V., Kaye, Thomas N., Roberts, Rachel, Dunwiddie, Peter W., Clark, Deborah Louise, 1950-, Stanley, Amanda Question: Most results of restoration efforts are species-specific and/or site-specific and therefore are not general enough to be easily applied to other species and other sites. Our research addresses the issue of species-specific results by investigating the feasibility of using plant traits instead of taxonomic species to characterize species responses to restoration treatments. Specifically, we test the explanatory power of plant traits, one necessary prerequisite for the development of predictive and general plant trait models. Location: Ten remnant bunchgrass prairie sites in the Pacific Northwest of North America (Oregon and Washington, USA; British Columbia, Canada). Methods: We developed two types of quantitative models for each of 10 prairie restoration sites: 1) plant trait models, which related plant traits to species field responses following restoration management treatments, and 2) species identity models, which related species taxonomic identity to species field responses following restoration management treatments. Species identity models determined the maximum amount of variability of field responses that can be explained by differences in individual species’ responses to management treatments. Plant trait models determined what proportion of this explanatory power can be attributed to plant traits. This approach contrasts with approaches often used in other plant trait studies that describe how traits vary with environmental conditions. Specifically, we used these two models to address the following questions: 1) How much of the variability in field responses of plants to restoration management treatments is explained by plant traits? 2) How well do plant traits explain the variability of field responses following restoration management treatments compared to models relating field responses to species identity? Our approach was to measure two aspects of explanatory power: R2 (variability explained) and AIC (a measure of model fit that accounts for parsimony, i.e., how well a model fits the data with relatively few explanatory variables). Results: 1) The plant trait models (relating plant traits with plant field responses) explained much of the variability within each of the ten restoration sites, with R2 values ranging between 31% and 69%. 2) The species identity models (relating species taxonomic identity with plant field responses) explained between 47% and 74% of variability in field performance (R2). Thus, the plant trait models explained nearly as much variability as the species identity models. In seven out of nine sites, the plant trait models were superior to the species identity models as measured by AIC; that is, the trait models did well at explaining variability with less model complexity (i.e., fewer explanatory variables). Conclusion: Development of general and predictive plant trait models is a multi-step process. Strong explanatory power by plant trait models, both on an absolute scale and as compared to species identity models, supports the feasibility of using plant traits instead of species taxonomic identity as a common language to characterize plant field responses to restoration treatments. Such high explanatory power is one necessary pre-requisite for the development of predictive and general plant trait models. Our results also indicate that the plant trait models are site-specific even though all sites were upland bunchgrass remnant prairies. We discuss the next steps in the development of more general and predictive models: incorporating environmental factors into the plant trait models to address the issue of site-specificity and testing the power of these models to predict vegetation responses. Close

• 8. [Article] Native plant revegetation at Butterfly Meadows (Benton County), habitat for the endangered Fender's blue butterfly

  Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon's ecosystems and are in critical need of conservation. One of the largest remaining ...

  Citation × Citation Citation Abstract Copy Title: Native plant revegetation at Butterfly Meadows (Benton County), habitat for the endangered Fender's blue butterfly Author: Clark, Deborah Louise, 1950- Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon's ecosystems and are in critical need of conservation. One of the largest remaining parcels of native upland prairie, Butterfly Meadows (Benton County), is being invaded by Brachypodium sylvaticum (false brome). This site is one of three most important remaining habitats for the Fender's blue butterfly and Kincaid's lupine, federally listed as Endangered and Threatened respectively. Most of Butterfly Meadows is owned by Starker Forests (Corvallis) with a small portion owned by Oregon State University. A partnership was formed in 2002 of people and organizations interested in protecting and restoring Butterfly Meadows: Fred Pfund (Starker Forests) Matt Blakeley-Smith (Institute for Applied Ecology) Deborah Clark (Oregon State University, Biology Program) Paul Hammond (Entomologist, Private consultant) Debbie Johnson (Oregon State University, College Forests) Tom Kaye (Institute for Applied Ecology) Bruce Kelpsas (Helena Chemical Company) In fall 2002, the partnership submitted a proposal (Control of Brachypodium sylvaticum and Restoration of Rare Native Upland Prairie Habitat at Butterfly Meadows, Benton County) and received funding from the Oregon Department of Agriculture. The project goal was and is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Since receiving funding in 2002, the partnership has developed and implemented herbicide treatments, beginning with small scale experiments, which control Brachypodium sylvaticum without harming native prairie vegetation or the Fender's blue butterfly. We used the results of these small-scale experiments to develop strategies for restoration of larger areas of the Butterfly Meadows, which Starker Forests has implemented at its own expense. These strategies have been relatively successful in controlling false brome, but we are now at a time where we need to revegetate these areas where false brome has been controlled. These areas are too large to quickly revegetate with existing native herbaceous species, so we need to augment the site with additional seed. Transplanting greenhouse grown plants is also an option but is much more costly. We hope to apply for additional funding elsewhere for these transplants. Title: Native plant revegetation at Butterfly Meadows (Benton County), habitat for the endangered Fender's blue butterfly Author: Clark, Deborah Louise, 1950- Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon's ecosystems and are in critical need of conservation. One of the largest remaining parcels of native upland prairie, Butterfly Meadows (Benton County), is being invaded by Brachypodium sylvaticum (false brome). This site is one of three most important remaining habitats for the Fender's blue butterfly and Kincaid's lupine, federally listed as Endangered and Threatened respectively. Most of Butterfly Meadows is owned by Starker Forests (Corvallis) with a small portion owned by Oregon State University. A partnership was formed in 2002 of people and organizations interested in protecting and restoring Butterfly Meadows: Fred Pfund (Starker Forests) Matt Blakeley-Smith (Institute for Applied Ecology) Deborah Clark (Oregon State University, Biology Program) Paul Hammond (Entomologist, Private consultant) Debbie Johnson (Oregon State University, College Forests) Tom Kaye (Institute for Applied Ecology) Bruce Kelpsas (Helena Chemical Company) In fall 2002, the partnership submitted a proposal (Control of Brachypodium sylvaticum and Restoration of Rare Native Upland Prairie Habitat at Butterfly Meadows, Benton County) and received funding from the Oregon Department of Agriculture. The project goal was and is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Since receiving funding in 2002, the partnership has developed and implemented herbicide treatments, beginning with small scale experiments, which control Brachypodium sylvaticum without harming native prairie vegetation or the Fender's blue butterfly. We used the results of these small-scale experiments to develop strategies for restoration of larger areas of the Butterfly Meadows, which Starker Forests has implemented at its own expense. These strategies have been relatively successful in controlling false brome, but we are now at a time where we need to revegetate these areas where false brome has been controlled. These areas are too large to quickly revegetate with existing native herbaceous species, so we need to augment the site with additional seed. Transplanting greenhouse grown plants is also an option but is much more costly. We hope to apply for additional funding elsewhere for these transplants. Close

• 9. [Article] Final report, year two: Control of Brachypodium sylvaticum and restoration of rare native upland prairie habitat at Butterfly Meadows, Benton County ODA 1514 GR

  Our project goal is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Native ...

  Citation × Citation Citation Abstract Copy Title: Final report, year two: Control of Brachypodium sylvaticum and restoration of rare native upland prairie habitat at Butterfly Meadows, Benton County ODA 1514 GR Author: Johnson, Deborah, Pfund, Fred, Hammond, Paul, Blakeley-Smith, Matt, Vomocil, Marc, Kaye, Tom, Clark, Deborah Louise, 1950-, Kelpsas, Bruce, Wilson, Mark V. Our project goal is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. One of the largest remaining parcels of native upland prairie, Butterfly Meadows (Benton County), is being invaded by Brachypodium sylvaticum. This site is one of the three most important remaining habitats for the Fender’s blue butterfly and Kincaid’s lupine, listed as Endangered and Threatened respectively. We propose to develop and implement herbicide treatments that control Brachypodium sylvaticum without harming native prairie vegetation. We will also develop and implement measures to reestablish native species from seed after removal of Brachypodium sylvaticum. Conifers and mature shrubs that have encroached on the meadow, both on the edge and in the complex, will be removed by mechanical means or girdling with herbicides. We propose to construct a buffer zone between the boundary of Butterfly Meadows and the neighboring intact forest and recently clear-cut areas, which are continuing sources of seed of Brachypodium sylvaticum invading Butterfly Meadows. This buffer zone will extend into both Starker Forest owned portions and OSU owned portions. To determine the success of control of Brachypodium sylvaticum and woody species and restoration of native vegetation, we will monitor changes in abundance of Brachypodium sylvaticum, woody species, and native vegetation. We will use monitoring results to adjust future Brachypodium sylvaticum control measures and native vegetation restoration measures as needed. Title: Final report, year two: Control of Brachypodium sylvaticum and restoration of rare native upland prairie habitat at Butterfly Meadows, Benton County ODA 1514 GR Author: Johnson, Deborah, Pfund, Fred, Hammond, Paul, Blakeley-Smith, Matt, Vomocil, Marc, Kaye, Tom, Clark, Deborah Louise, 1950-, Kelpsas, Bruce, Wilson, Mark V. Our project goal is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. One of the largest remaining parcels of native upland prairie, Butterfly Meadows (Benton County), is being invaded by Brachypodium sylvaticum. This site is one of the three most important remaining habitats for the Fender’s blue butterfly and Kincaid’s lupine, listed as Endangered and Threatened respectively. We propose to develop and implement herbicide treatments that control Brachypodium sylvaticum without harming native prairie vegetation. We will also develop and implement measures to reestablish native species from seed after removal of Brachypodium sylvaticum. Conifers and mature shrubs that have encroached on the meadow, both on the edge and in the complex, will be removed by mechanical means or girdling with herbicides. We propose to construct a buffer zone between the boundary of Butterfly Meadows and the neighboring intact forest and recently clear-cut areas, which are continuing sources of seed of Brachypodium sylvaticum invading Butterfly Meadows. This buffer zone will extend into both Starker Forest owned portions and OSU owned portions. To determine the success of control of Brachypodium sylvaticum and woody species and restoration of native vegetation, we will monitor changes in abundance of Brachypodium sylvaticum, woody species, and native vegetation. We will use monitoring results to adjust future Brachypodium sylvaticum control measures and native vegetation restoration measures as needed. Close

• 10. [Article] Final report [year one]: Control of Brachypodium sylvaticum and restoration of rare native upland prairie habitat at Butterfly Meadows, Benton County ODA 1342 GR

  Our project goal is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Native ...

  Citation × Citation Citation Abstract Copy Title: Final report [year one]: Control of Brachypodium sylvaticum and restoration of rare native upland prairie habitat at Butterfly Meadows, Benton County ODA 1342 GR Author: Johnson, Deborah, Pfund, Fred, Hammond, Paul, Wilson, Mark V., Vomocil, Marc, Kaye, Tom, Clark, Deborah Louise, 1950-, Kelpsas, Bruce Our project goal is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. One of the largest remaining parcels of native upland prairie, Butterfly Meadows (Benton County), is being invaded by Brachypodium sylvaticum. This site is one of the three most important remaining habitats for the Fender’s blue butterfly and Kincaid’s lupine, listed as Endangered and Threatened respectively. We propose to develop and implement herbicide treatments that control Brachypodium sylvaticum without harming native prairie vegetation. We will also develop and implement measures to reestablish native species from seed after removal of Brachypodium sylvaticum. Conifers and mature shrubs that have encroached on the meadow, both on the edge and in the complex, will be removed by mechanical means or girdling. We propose to construct a buffer zone between the boundary of Butterfly Meadows and the neighboring intact forest and recently clear-cut areas, which are continuing sources of seed of Brachypodium sylvaticum invading Butterfly Meadows. This buffer zone will extend into both Starker Forest owned portions and OSU owned portions. To determine the success of control of Brachypodium sylvaticum and woody species and restoration of native vegetation, we will monitor changes in abundance of Brachypodium sylvaticum, woody species, and native vegetation. We will use monitoring results to adjust future Brachypodium sylvaticum control measures and native vegetation restoration measures as needed. Title: Final report [year one]: Control of Brachypodium sylvaticum and restoration of rare native upland prairie habitat at Butterfly Meadows, Benton County ODA 1342 GR Author: Johnson, Deborah, Pfund, Fred, Hammond, Paul, Wilson, Mark V., Vomocil, Marc, Kaye, Tom, Clark, Deborah Louise, 1950-, Kelpsas, Bruce Our project goal is to protect and restore rare Willamette Valley upland prairie habitat at Butterfly Meadows (Benton County) from invasion by the noxious weed Brachypodium sylvaticum (false brome). Native prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. One of the largest remaining parcels of native upland prairie, Butterfly Meadows (Benton County), is being invaded by Brachypodium sylvaticum. This site is one of the three most important remaining habitats for the Fender’s blue butterfly and Kincaid’s lupine, listed as Endangered and Threatened respectively. We propose to develop and implement herbicide treatments that control Brachypodium sylvaticum without harming native prairie vegetation. We will also develop and implement measures to reestablish native species from seed after removal of Brachypodium sylvaticum. Conifers and mature shrubs that have encroached on the meadow, both on the edge and in the complex, will be removed by mechanical means or girdling. We propose to construct a buffer zone between the boundary of Butterfly Meadows and the neighboring intact forest and recently clear-cut areas, which are continuing sources of seed of Brachypodium sylvaticum invading Butterfly Meadows. This buffer zone will extend into both Starker Forest owned portions and OSU owned portions. To determine the success of control of Brachypodium sylvaticum and woody species and restoration of native vegetation, we will monitor changes in abundance of Brachypodium sylvaticum, woody species, and native vegetation. We will use monitoring results to adjust future Brachypodium sylvaticum control measures and native vegetation restoration measures as needed. Close

• 11. [Article] Final Report: Controlling Tall Oatgrass in Fender’s Blue Butterfly Habitats

  We report final results from a five-year field experiment evaluating the effects of different mowing regimes on groups of native and non-native grasses and forbs. The study site was a degraded prairie ...

  Citation × Citation Citation Abstract Copy Title: Final Report: Controlling Tall Oatgrass in Fender’s Blue Butterfly Habitats Author: Wilson, Mark V., Clark, Deborah Louise, 1950- We report final results from a five-year field experiment evaluating the effects of different mowing regimes on groups of native and non-native grasses and forbs. The study site was a degraded prairie at Baskett Slough National Wildlife Refuge dominated by the pest plant Arrhenatherum elatius (tall oatgrass). Mowing treatments differed in time of year they were applied, mowing height, and removal of cut material. Treatments started in 1994 and continued through 1998. Title: Final Report: Controlling Tall Oatgrass in Fender’s Blue Butterfly Habitats Author: Wilson, Mark V., Clark, Deborah Louise, 1950- We report final results from a five-year field experiment evaluating the effects of different mowing regimes on groups of native and non-native grasses and forbs. The study site was a degraded prairie at Baskett Slough National Wildlife Refuge dominated by the pest plant Arrhenatherum elatius (tall oatgrass). Mowing treatments differed in time of year they were applied, mowing height, and removal of cut material. Treatments started in 1994 and continued through 1998. Close

• 12. [Article] Final report: Restoration of Fender’s blue butterfly and its prairie ecosystem: Management applications of fire to Baskett Slough NWR

  We tested the recolonization ability of the Fender’s blue butterfly and monitored the vegetation response to a management-scale prescribed fire at the Baskett Slough National Wildlife Refuge, western Oregon. ...

  Citation × Citation Citation Abstract Copy Title: Final report: Restoration of Fender’s blue butterfly and its prairie ecosystem: Management applications of fire to Baskett Slough NWR Author: Wilson, Mark V., Clark, Deborah Louise, 1950- We tested the recolonization ability of the Fender’s blue butterfly and monitored the vegetation response to a management-scale prescribed fire at the Baskett Slough National Wildlife Refuge, western Oregon. The prescribed fire successfully cut shrub cover in half, but also reduced the size of Festuca roemeri, a dominant native grass. Although the fire killed all Fender’s blue butterfly larvae, as expected, adult females were able to recolonize the burned area from a nearby, unburned patch. These results support predictions that the Fender’s blue butterfly can recolonize burned areas for a distance of at least 100 m. Title: Final report: Restoration of Fender’s blue butterfly and its prairie ecosystem: Management applications of fire to Baskett Slough NWR Author: Wilson, Mark V., Clark, Deborah Louise, 1950- We tested the recolonization ability of the Fender’s blue butterfly and monitored the vegetation response to a management-scale prescribed fire at the Baskett Slough National Wildlife Refuge, western Oregon. The prescribed fire successfully cut shrub cover in half, but also reduced the size of Festuca roemeri, a dominant native grass. Although the fire killed all Fender’s blue butterfly larvae, as expected, adult females were able to recolonize the burned area from a nearby, unburned patch. These results support predictions that the Fender’s blue butterfly can recolonize burned areas for a distance of at least 100 m. Close

• 13. [Article] Restoration of native upland prairies: Habitat for Fender’s blue butterfly (Icaricia icarioides fenderi)

  Our project goal is to promote the abundance of the endangered Fender’s blue butterfly (Icaricia icarioides fenderi) by recreating upland prairie habitat in former agricultural land at Finley National ...

  Citation × Citation Citation Abstract Copy Title: Restoration of native upland prairies: Habitat for Fender’s blue butterfly (Icaricia icarioides fenderi) Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Our project goal is to promote the abundance of the endangered Fender’s blue butterfly (Icaricia icarioides fenderi) by recreating upland prairie habitat in former agricultural land at Finley National Wildlife Refuge, Corvallis, Oregon. To achieve this goal we initiated the following three studies. Study One: Experimental Investigation on the Effectiveness of Restoration Treatments Our approach was to set up experimental field plots in a former agricultural field into which we sowed seeds of native grasses and dicots in the fall for two years, 2003 and 2004. Objective one: Investigate the effect of carbon banding on abundance of sowed native species and non-native species. Carbon banding is a treatment used by Willamette Valley grass seed farmers to promote seedling establishment of agricultural grasses and inhibit seedling establishment of weeds. Objective two: Investigate the effect of the sowing sequence of native dicot and grass species on the abundance and species richness of the sowed species. Our experiments were designed to test the hypothesis that our native grasses are more aggressive than the target native dicots in the initial stages of restoration. Objective three: Investigate the effect of the sowing sequence of the native dicot and grass species on the abundance of non-native species. Study Two: Monoculture Sowing of Native Species Compare the cover of non-native species and the cover of native species one and two years after sowing native species in monocultures. Study Three: Buffer Sowing of Native Grasses Monitor the abundance of the native grasses sowed in the buffer areas outside the experimental plots. Carbon banding had no significant effect on cover of non-native species or on cover and species richness of sowed native dicots or grasses after both the first and second growing seasons. However, carbon banding tended to reduce the cover of the annual Clarkia rhomboidea after the first growing season. Our results supported the hypothesis that our native grasses are more aggressive than the target native dicots in the initial stages of restoration. The sowing sequences tested did not affect cover and species richness of grasses but did affect cover and species richness of dicot species. Both dicot cover and dicot species richness were greater when sowed a year ahead of the grasses. Eriophyllum lanatum had significantly greater cover when sowed a year before the grasses. Average cover of sowed dicots was significantly greater when dicots were sowed with grasses compared to when dicots were sowed into a year-old stand of grasses. In particular, Madia gracilis and Prunella vulgaris var. lanceolata had less cover when sowed into the existing stand of grass. Species richness of all native species (grasses + dicots) was significantly less when dicots were sowed into a year-old stand of grasses. However, total cover and species richness of sowed native species (grasses + dicots) were not significantly different whether dicots were sowed a year ahead of the grasses or whether dicots and grasses were sowed together. Average cover of sowed native dicots after two years was always greater than the cover of sowed native grasses except when dicots were sowed a year after the grasses. After two growing seasons, non-native cover did not significantly vary among the three sowing sequences of sowing grasses and dicots at the same time, sowing grasses into existing dicot vegetation, and sowing grasses into existing dicot vegetation. The first year after sowing the monocultures (Study two), Collinsia grandiflora had the highest cover, greatly surpassing the next top performers, Sanguisorba occidentalis and Madia gracilis, all of which are annual species. In general, the cover of individual species sowed in monocultures decreased the second year after sowing. Only seven species, all of which were perennials, out of the 26 sowed species increased in cover. Two of these species Eriophyllum lanatum and Prunella vulgaris var. lanceolata were the top performers for 2005, but their cover did not reach the levels of the top performers in 2004. Title: Restoration of native upland prairies: Habitat for Fender’s blue butterfly (Icaricia icarioides fenderi) Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Our project goal is to promote the abundance of the endangered Fender’s blue butterfly (Icaricia icarioides fenderi) by recreating upland prairie habitat in former agricultural land at Finley National Wildlife Refuge, Corvallis, Oregon. To achieve this goal we initiated the following three studies. Study One: Experimental Investigation on the Effectiveness of Restoration Treatments Our approach was to set up experimental field plots in a former agricultural field into which we sowed seeds of native grasses and dicots in the fall for two years, 2003 and 2004. Objective one: Investigate the effect of carbon banding on abundance of sowed native species and non-native species. Carbon banding is a treatment used by Willamette Valley grass seed farmers to promote seedling establishment of agricultural grasses and inhibit seedling establishment of weeds. Objective two: Investigate the effect of the sowing sequence of native dicot and grass species on the abundance and species richness of the sowed species. Our experiments were designed to test the hypothesis that our native grasses are more aggressive than the target native dicots in the initial stages of restoration. Objective three: Investigate the effect of the sowing sequence of the native dicot and grass species on the abundance of non-native species. Study Two: Monoculture Sowing of Native Species Compare the cover of non-native species and the cover of native species one and two years after sowing native species in monocultures. Study Three: Buffer Sowing of Native Grasses Monitor the abundance of the native grasses sowed in the buffer areas outside the experimental plots. Carbon banding had no significant effect on cover of non-native species or on cover and species richness of sowed native dicots or grasses after both the first and second growing seasons. However, carbon banding tended to reduce the cover of the annual Clarkia rhomboidea after the first growing season. Our results supported the hypothesis that our native grasses are more aggressive than the target native dicots in the initial stages of restoration. The sowing sequences tested did not affect cover and species richness of grasses but did affect cover and species richness of dicot species. Both dicot cover and dicot species richness were greater when sowed a year ahead of the grasses. Eriophyllum lanatum had significantly greater cover when sowed a year before the grasses. Average cover of sowed dicots was significantly greater when dicots were sowed with grasses compared to when dicots were sowed into a year-old stand of grasses. In particular, Madia gracilis and Prunella vulgaris var. lanceolata had less cover when sowed into the existing stand of grass. Species richness of all native species (grasses + dicots) was significantly less when dicots were sowed into a year-old stand of grasses. However, total cover and species richness of sowed native species (grasses + dicots) were not significantly different whether dicots were sowed a year ahead of the grasses or whether dicots and grasses were sowed together. Average cover of sowed native dicots after two years was always greater than the cover of sowed native grasses except when dicots were sowed a year after the grasses. After two growing seasons, non-native cover did not significantly vary among the three sowing sequences of sowing grasses and dicots at the same time, sowing grasses into existing dicot vegetation, and sowing grasses into existing dicot vegetation. The first year after sowing the monocultures (Study two), Collinsia grandiflora had the highest cover, greatly surpassing the next top performers, Sanguisorba occidentalis and Madia gracilis, all of which are annual species. In general, the cover of individual species sowed in monocultures decreased the second year after sowing. Only seven species, all of which were perennials, out of the 26 sowed species increased in cover. Two of these species Eriophyllum lanatum and Prunella vulgaris var. lanceolata were the top performers for 2005, but their cover did not reach the levels of the top performers in 2004. Close

• 14. [Article] Effects of Soil Nutrients and Fire on Seedling Establishment of Native Prairie Species

  We sought to determine 1) the relationship between seedling establishment rates of target native prairie species and field levels of three soil nutrients (phosphate, nitrate, and ammonium) recorded in ...

  Citation × Citation Citation Abstract Copy Title: Effects of Soil Nutrients and Fire on Seedling Establishment of Native Prairie Species Author: Clark, Deborah Louise, 1950- We sought to determine 1) the relationship between seedling establishment rates of target native prairie species and field levels of three soil nutrients (phosphate, nitrate, and ammonium) recorded in fall, winter, and summer, 2) the effect of fire on these three nutrients, and 3) the overall effect of fire on seedling establishment rates of the target native prairie species. Seedling establishment rates were not significantly correlated with soil nitrate or ammonium. Phosphate was significantly correlated with seedling establishment rates but the direction and strength of the relationship depended on the species, with three species showing positive relationships and three species showing negative relationships. Soil phosphate increased significantly in the burned plots compared to the unburned plots, soil ammonium decreased significantly in the burned plots, and soil nitrate showed no significant differences. These nutrient patterns successfully predicted the overall fire effects on seedling establishment in a separate experiment. The three species with positive correlations between seedling establishment and phosphate had greater seedling establishment rates in the burned plots compared to the unburned plots (two significantly, P≤0.1), and the three species with negative correlations had fewer seeds establish in the burned plots (two significantly (P ≤ 0.1). Title: Effects of Soil Nutrients and Fire on Seedling Establishment of Native Prairie Species Author: Clark, Deborah Louise, 1950- We sought to determine 1) the relationship between seedling establishment rates of target native prairie species and field levels of three soil nutrients (phosphate, nitrate, and ammonium) recorded in fall, winter, and summer, 2) the effect of fire on these three nutrients, and 3) the overall effect of fire on seedling establishment rates of the target native prairie species. Seedling establishment rates were not significantly correlated with soil nitrate or ammonium. Phosphate was significantly correlated with seedling establishment rates but the direction and strength of the relationship depended on the species, with three species showing positive relationships and three species showing negative relationships. Soil phosphate increased significantly in the burned plots compared to the unburned plots, soil ammonium decreased significantly in the burned plots, and soil nitrate showed no significant differences. These nutrient patterns successfully predicted the overall fire effects on seedling establishment in a separate experiment. The three species with positive correlations between seedling establishment and phosphate had greater seedling establishment rates in the burned plots compared to the unburned plots (two significantly, P≤0.1), and the three species with negative correlations had fewer seeds establish in the burned plots (two significantly (P ≤ 0.1). Close

• 15. [Article] Recommendations for Control of Tall Oatgrass, Poison Oak, and Rose in Willamette Valley Upland Prairies

  Native upland prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon's ecosystems. Even though only remnants remain today, they harbor many rare ...

  Citation × Citation Citation Abstract Copy Title: Recommendations for Control of Tall Oatgrass, Poison Oak, and Rose in Willamette Valley Upland Prairies Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Native upland prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon's ecosystems. Even though only remnants remain today, they harbor many rare and endangered species such as the Fender's blue butterfly and its host plant, Kincaid’s lupine. Invasion of remnant prairies by woody plants and weedy non-native herbaceous species is of great concern to agencies and managers responsible for conserving these native ecosystems. Available methods for the control of these pests include herbicides, biological control, prescribed burning, and mowing. This report first reviews the literature on the control of tall oatgrass and the shrubs poison oak and rose. Each of these species is an important threat to Willamette Valley upland prairies. We then presents recommendations. For control of tall oatgrass, we recommend for most situations mowing in late spring or early summer. Mowing height should be just above most of the foliage of native plants to be protected (typically 10 cm - 15 cm). In areas with Kincaid’s lupine and Fender’s blue butterfly, mowing should be delayed until July, after the flowering and flight seasons. For control of prairie shrubs, we recommend further studies to evaluate herbicide effectiveness against the target shrubs and safety for non-target native plants and animals. If herbicide use is unavailable, we recommend burning or mowing. All burning or mowing treatments should be repeated, perhaps indefinitely, to maintain pest plant control. Title: Recommendations for Control of Tall Oatgrass, Poison Oak, and Rose in Willamette Valley Upland Prairies Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Native upland prairies, which once dominated the landscape of the Willamette Valley, are considered among the rarest of Oregon's ecosystems. Even though only remnants remain today, they harbor many rare and endangered species such as the Fender's blue butterfly and its host plant, Kincaid’s lupine. Invasion of remnant prairies by woody plants and weedy non-native herbaceous species is of great concern to agencies and managers responsible for conserving these native ecosystems. Available methods for the control of these pests include herbicides, biological control, prescribed burning, and mowing. This report first reviews the literature on the control of tall oatgrass and the shrubs poison oak and rose. Each of these species is an important threat to Willamette Valley upland prairies. We then presents recommendations. For control of tall oatgrass, we recommend for most situations mowing in late spring or early summer. Mowing height should be just above most of the foliage of native plants to be protected (typically 10 cm - 15 cm). In areas with Kincaid’s lupine and Fender’s blue butterfly, mowing should be delayed until July, after the flowering and flight seasons. For control of prairie shrubs, we recommend further studies to evaluate herbicide effectiveness against the target shrubs and safety for non-target native plants and animals. If herbicide use is unavailable, we recommend burning or mowing. All burning or mowing treatments should be repeated, perhaps indefinitely, to maintain pest plant control. Close

• 16. [Article] Fire effects on wetland prairie species: final report

  Project No. 14-48-0001-96749

  Citation × Citation Citation Abstract Copy Title: Fire effects on wetland prairie species: final report Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Project No. 14-48-0001-96749 Title: Fire effects on wetland prairie species: final report Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Project No. 14-48-0001-96749 Close

• 17. [Article] Controlling woody vegetation in wetland prairies, 1994-1999

  Wetland prairies of the Willamette Valley, among the rarest of Oregon’s ecosystems, are threatened by invasion of woody species and non-native pest species. Because fire has been important in maintaining ...

  Citation × Citation Citation Abstract Copy Title: Controlling woody vegetation in wetland prairies, 1994-1999 Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Wetland prairies of the Willamette Valley, among the rarest of Oregon’s ecosystems, are threatened by invasion of woody species and non-native pest species. Because fire has been important in maintaining Willamette Valley prairies for at least 1000 years, prescribed burning is a top choice of managers for preventing encroachment of woody species. However, the effects of prescribed burning on present day wetland prairie communities, with their mix of native and non-native species, are not clear. Moreover, because strict governmental requirements for smoke management hinder prescribed burning, natural-area managers are considering alternatives to fire, like mowing and hand removal of woody plants. The present study explored the ecological interactions of a wetland prairie by experimentally testing the responses of burning, hand-removal of woody species, and mowing with removal of cut material on woody plants and key native and non-native grasses and forbs. We evaluated these perturbations for their effectiveness in meeting management objectives of reducing abundance of woody species, reducing or preventing an increase in abundance of nonnative pest species, and increasing or at least maintaining native species’ abundance. After three treatments periods (1994, 1996, and 1998) the following patterns emerged: • Woody species Burning and hand-removal consistently caused the greatest reduction in cover of woody species. Mowing with removal of cut material did not reduce the abundance of woody species compared to the control. As woody plant cover decreased, plant mortality increased, indicating that treatments influenced woody plant cover at least partially through changes in survival. • Native herbaceous species All management treatments consistently failed to significantly increase the cover of Deschampsia cespitosa, while burning decreased the number of inflorescences. As a group, native graminoid flowering consistently showed no significant responses to the management treatments. Although burning and hand-removal significantly increased the cover of native forbs as group after the second treatment period in 1997, native forbs showed no significant responses to treatments in 1999. • Non-native herbaceous species Response of non-native forb cover to management treatments has been inconsistent during the five year study period with no significant differences in 1995, significant reductions by burning and hand-removal in 1997, and increases with mowing in 1997. Mowing consistently increased flowering of nonnative graminoids as a group. Overall no treatment was clearly superior in fulfilling restoration objectives. Burning was effective in reducing woody cover and did not promote non-native herbaceous, but it reduced the flowering the dominant native grass, Deschampsia cespitosa. Hand-removal woody species was also effective at reducing woody cover and sometimes promoted the abundance of some native species at different times, but it also at times promoted non-native herbaceous species. Because mowing with removal of cut material was ineffective in reducing woody cover and tended to promote non-native herbaceous species, this treatment is not a recommended management strategy. Title: Controlling woody vegetation in wetland prairies, 1994-1999 Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Wetland prairies of the Willamette Valley, among the rarest of Oregon’s ecosystems, are threatened by invasion of woody species and non-native pest species. Because fire has been important in maintaining Willamette Valley prairies for at least 1000 years, prescribed burning is a top choice of managers for preventing encroachment of woody species. However, the effects of prescribed burning on present day wetland prairie communities, with their mix of native and non-native species, are not clear. Moreover, because strict governmental requirements for smoke management hinder prescribed burning, natural-area managers are considering alternatives to fire, like mowing and hand removal of woody plants. The present study explored the ecological interactions of a wetland prairie by experimentally testing the responses of burning, hand-removal of woody species, and mowing with removal of cut material on woody plants and key native and non-native grasses and forbs. We evaluated these perturbations for their effectiveness in meeting management objectives of reducing abundance of woody species, reducing or preventing an increase in abundance of nonnative pest species, and increasing or at least maintaining native species’ abundance. After three treatments periods (1994, 1996, and 1998) the following patterns emerged: • Woody species Burning and hand-removal consistently caused the greatest reduction in cover of woody species. Mowing with removal of cut material did not reduce the abundance of woody species compared to the control. As woody plant cover decreased, plant mortality increased, indicating that treatments influenced woody plant cover at least partially through changes in survival. • Native herbaceous species All management treatments consistently failed to significantly increase the cover of Deschampsia cespitosa, while burning decreased the number of inflorescences. As a group, native graminoid flowering consistently showed no significant responses to the management treatments. Although burning and hand-removal significantly increased the cover of native forbs as group after the second treatment period in 1997, native forbs showed no significant responses to treatments in 1999. • Non-native herbaceous species Response of non-native forb cover to management treatments has been inconsistent during the five year study period with no significant differences in 1995, significant reductions by burning and hand-removal in 1997, and increases with mowing in 1997. Mowing consistently increased flowering of nonnative graminoids as a group. Overall no treatment was clearly superior in fulfilling restoration objectives. Burning was effective in reducing woody cover and did not promote non-native herbaceous, but it reduced the flowering the dominant native grass, Deschampsia cespitosa. Hand-removal woody species was also effective at reducing woody cover and sometimes promoted the abundance of some native species at different times, but it also at times promoted non-native herbaceous species. Because mowing with removal of cut material was ineffective in reducing woody cover and tended to promote non-native herbaceous species, this treatment is not a recommended management strategy. Close

• 18. [Article] Controlling woody vegetation in wetland prairies, 1994-2001

  Wetland prairies of the Willamette Valley, among the rarest of Oregon’s ecosystems, are threatened by invasion of woody species and non-native pest species. Because fire has been important in maintaining ...

  Citation × Citation Citation Abstract Copy Title: Controlling woody vegetation in wetland prairies, 1994-2001 Author: Clark, Deborah Louise, 1950- Wetland prairies of the Willamette Valley, among the rarest of Oregon’s ecosystems, are threatened by invasion of woody species and non-native pest species. Because fire has been important in maintaining Willamette Valley prairies for at least 1000 years, prescribed burning is a top choice of managers for preventing encroachment of woody species. However, the effects of prescribed burning on present day wetland prairie communities, with their mix of native and non-native species, are not clear. Moreover, because strict governmental requirements for smoke management hinder prescribed burning, natural-area managers are considering alternatives to fire, like mowing and hand-removal of woody plants. The present study explored the ecological interactions of a wetland prairie by experimentally testing the responses of burning, hand-removal of woody species, and mowing with removal of cut material on woody plants and key native and non-native grasses and forbs. We evaluated these perturbations for their effectiveness in meeting management objectives of reducing abundance of woody species, reducing or preventing an increase in abundance of nonnative pest species, and increasing or at least maintaining native species’ abundance. After four treatments periods (1994, 1996, 1998, and 2000) the following patterns emerged: • Woody species Burning and hand-removal of woody plants every two years consistently decreased the cover of woody species, even with a gradual increase in the controls to over double 1994 levels. The change in woody cover after mowing with removal of cut material was not significantly different from that in the control. As woody plant cover decreased, plant mortality increased, indicating that treatments influenced woody plant cover at least partially through changes in survival. • Native herbaceous species None of the treatments had a significant effect after four treatment periods on the proportional change in cover or in number of inflorescences of Deschampsia cespitosa, although the change in number of inflorescences was consistently less than that in the control, including a significant decrease with burning in 1997. As a group, native graminoid species showed no significant differences in flowering response to treatments during any of the four treatment periods. However, burning and mowing consistently promoted flowering of Juncus tenuis. The cover of native forbs as a group was not significantly different from that of the control, with the exception of 1997, where cover increased with burning and hand-removal. Even though native forbs as a group have generally not shown significant differences, individual native forb species have shown positive responses to treatments: Sidalcea cusickii, spp. purpurea, Eriophyllum lanatum, and Sisyrinchium idahoensis. • Non-native herbaceous species Both hand-removal and mowing promoted flowering of non-native graminoids as a group. In contrast, burning maintained levels of flowering of non-native graminoids similar to that of the controls. Cover of non-native forbs as a group showed no significant treatment differences except for 1997, in which significant reductions in cover occurred after burning and hand-removal. The results to date show that prescribed burning is the best option of the three tested management treatments for achieving the restoration goals for native wetland prairies. Because mowing with removal of cut material was ineffective in reducing woody cover and tended to promote non-native herbaceous species, this treatment should be avoided. Title: Controlling woody vegetation in wetland prairies, 1994-2001 Author: Clark, Deborah Louise, 1950- Wetland prairies of the Willamette Valley, among the rarest of Oregon’s ecosystems, are threatened by invasion of woody species and non-native pest species. Because fire has been important in maintaining Willamette Valley prairies for at least 1000 years, prescribed burning is a top choice of managers for preventing encroachment of woody species. However, the effects of prescribed burning on present day wetland prairie communities, with their mix of native and non-native species, are not clear. Moreover, because strict governmental requirements for smoke management hinder prescribed burning, natural-area managers are considering alternatives to fire, like mowing and hand-removal of woody plants. The present study explored the ecological interactions of a wetland prairie by experimentally testing the responses of burning, hand-removal of woody species, and mowing with removal of cut material on woody plants and key native and non-native grasses and forbs. We evaluated these perturbations for their effectiveness in meeting management objectives of reducing abundance of woody species, reducing or preventing an increase in abundance of nonnative pest species, and increasing or at least maintaining native species’ abundance. After four treatments periods (1994, 1996, 1998, and 2000) the following patterns emerged: • Woody species Burning and hand-removal of woody plants every two years consistently decreased the cover of woody species, even with a gradual increase in the controls to over double 1994 levels. The change in woody cover after mowing with removal of cut material was not significantly different from that in the control. As woody plant cover decreased, plant mortality increased, indicating that treatments influenced woody plant cover at least partially through changes in survival. • Native herbaceous species None of the treatments had a significant effect after four treatment periods on the proportional change in cover or in number of inflorescences of Deschampsia cespitosa, although the change in number of inflorescences was consistently less than that in the control, including a significant decrease with burning in 1997. As a group, native graminoid species showed no significant differences in flowering response to treatments during any of the four treatment periods. However, burning and mowing consistently promoted flowering of Juncus tenuis. The cover of native forbs as a group was not significantly different from that of the control, with the exception of 1997, where cover increased with burning and hand-removal. Even though native forbs as a group have generally not shown significant differences, individual native forb species have shown positive responses to treatments: Sidalcea cusickii, spp. purpurea, Eriophyllum lanatum, and Sisyrinchium idahoensis. • Non-native herbaceous species Both hand-removal and mowing promoted flowering of non-native graminoids as a group. In contrast, burning maintained levels of flowering of non-native graminoids similar to that of the controls. Cover of non-native forbs as a group showed no significant treatment differences except for 1997, in which significant reductions in cover occurred after burning and hand-removal. The results to date show that prescribed burning is the best option of the three tested management treatments for achieving the restoration goals for native wetland prairies. Because mowing with removal of cut material was ineffective in reducing woody cover and tended to promote non-native herbaceous species, this treatment should be avoided. 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• 19. [Article] Promoting regeneration of native species In Willamette Valley upland prairies

  Native prairies of the Willamette Valley are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. Management strategies for increasing the abundance of native species ...

  Citation × Citation Citation Abstract Copy Title: Promoting regeneration of native species In Willamette Valley upland prairies Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Native prairies of the Willamette Valley are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. Management strategies for increasing the abundance of native species are urgently needed, particularly those strategies that promote the regeneration of native species from seed. The focus of this investigation was to address this need. The general approach was to sow seeds of native species into experimental field plots, which had been burned once, burned repeatedly, mowed, or unmanaged at three sites at Baskett Slough National Wildlife Refuge. Seedling numbers were then counted one growing season after sowing. To investigate mechanisms for the regeneration responses of species to management treatments, selected abiotic and biotic factors potentially important for seed regeneration were measured in the field plots. Seedling establishment rates in unmanaged plots were relatively small, averaging 9.7% for all three sites. Only one species, Danthonia californica, showed a positive response to the mowing treatment. Of the 13 species sowed in the burn plots, six species (Bromus carinatus, Elymus glacus, Festuca roemeri/rubra, Clarkia quadrivulera, Prunella vulgaris, Sidalcea campestris) had significant more seedlings in the burn plots compared to the unmanaged plots. Seedling establishment of two species, Danthonia californica and Potentilla gracilis, was significantly lower with burn treatments. The other five species showed no significant responses to the burn treatments: Carex tumicola, Calochortus tolmiei, Zygadensus venenosus, Balsamorhiza deltoidea, and Agoseris grandilflora. The strongest fire effects were generally in plots that had been burned only once, rather than plots burned more than once. Abundance of litter and vegetation were the factors that were most often related, both positively and negatively, with seedling establishment rates. Based on the results of this investigation, prescribed burning, rather than mowing, is a more viable management option to promote regeneration of native species from seed in upland prairies. However, prescribed burning must be used with caution as it can reduce seedling establishment rates for some species. Sowing seeds in unmanaged sites is an option, but sowing rates need to be high to compensate for the small seedling establishment rates in unmanaged prairies. Title: Promoting regeneration of native species In Willamette Valley upland prairies Author: Wilson, Mark V., Clark, Deborah Louise, 1950- Native prairies of the Willamette Valley are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. Management strategies for increasing the abundance of native species are urgently needed, particularly those strategies that promote the regeneration of native species from seed. The focus of this investigation was to address this need. The general approach was to sow seeds of native species into experimental field plots, which had been burned once, burned repeatedly, mowed, or unmanaged at three sites at Baskett Slough National Wildlife Refuge. Seedling numbers were then counted one growing season after sowing. To investigate mechanisms for the regeneration responses of species to management treatments, selected abiotic and biotic factors potentially important for seed regeneration were measured in the field plots. Seedling establishment rates in unmanaged plots were relatively small, averaging 9.7% for all three sites. Only one species, Danthonia californica, showed a positive response to the mowing treatment. Of the 13 species sowed in the burn plots, six species (Bromus carinatus, Elymus glacus, Festuca roemeri/rubra, Clarkia quadrivulera, Prunella vulgaris, Sidalcea campestris) had significant more seedlings in the burn plots compared to the unmanaged plots. Seedling establishment of two species, Danthonia californica and Potentilla gracilis, was significantly lower with burn treatments. The other five species showed no significant responses to the burn treatments: Carex tumicola, Calochortus tolmiei, Zygadensus venenosus, Balsamorhiza deltoidea, and Agoseris grandilflora. The strongest fire effects were generally in plots that had been burned only once, rather than plots burned more than once. Abundance of litter and vegetation were the factors that were most often related, both positively and negatively, with seedling establishment rates. Based on the results of this investigation, prescribed burning, rather than mowing, is a more viable management option to promote regeneration of native species from seed in upland prairies. However, prescribed burning must be used with caution as it can reduce seedling establishment rates for some species. Sowing seeds in unmanaged sites is an option, but sowing rates need to be high to compensate for the small seedling establishment rates in unmanaged prairies. Close

• 20. [Article] Increasing the abundance of rare native wetland prairie species

  Native prairies of the Willamette Valley are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. Management strategies for increasing the abundance of native species ...

  Citation × Citation Citation Abstract Copy Title: Increasing the abundance of rare native wetland prairie species Author: Goodridge, Jennifer, Wilson, Mark V., Clark, Deborah Louise, 1950- Native prairies of the Willamette Valley are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. Management strategies for increasing the abundance of native species are urgently needed, particularly those strategies that promote the regeneration of native species from seed. Fire may be an important factor in promoting regeneration of native species from seed because of its historical role in maintaining the prairie landscape. The study objectives were (1) to determine the effect of prescribed burning on regeneration of native species from seed, and (2) to establish predictive relationships between easily measured plant traits and seedling establishment rates in the field. The general approach was to sow seeds of target wetland prairie species during the fall into experimental field plots already established at the Danebo Wetland, Eugene, OR. Seedling establishment rates were then compared between burned and unburned plots the following spring. To establish predictive relationships these seedling establishment rates were related to selected seed and seedling traits measured under laboratory conditions. Overall seedling establishment rates showed no significant differences between burned (7.0%) and unburned plots (8.7%), although for the seven species showing positive responses to prescribed burning, the increase was approximately doubled in the burned plots. Burning significantly increased the seedling establishment rates of three species, Wyethia angustifolia, Grindelia integrifolia, and Danthonia californica. Seedling establishment significantly decreased with burning for one species, Sidalcea campestris. For three of the four endangered, threatened or rare species, Aster curtus, Horkelia congesta, and Sidalcea cusickii var. purpurea, seedling establishment rates were smaller in the burned plots compared to the unburned plots, although the differences were not significant. Lomatium bradshawii had no seedlings establish in either the burned or the unburned plots. Plant weight 7 days after germination was the best trait at predicting field seedling establishment rates for both the burned plots and the unburned plots, explaining a significant amount of variation in establishment rates: 70% for the burned plots and 45% for the unburned plots. This model could be used by managers to choose species for sowing in burned and unburned prairies or to estimate seeding rates at burned and unburned sites. Title: Increasing the abundance of rare native wetland prairie species Author: Goodridge, Jennifer, Wilson, Mark V., Clark, Deborah Louise, 1950- Native prairies of the Willamette Valley are considered among the rarest of Oregon’s ecosystems and are in critical need of conservation. Management strategies for increasing the abundance of native species are urgently needed, particularly those strategies that promote the regeneration of native species from seed. Fire may be an important factor in promoting regeneration of native species from seed because of its historical role in maintaining the prairie landscape. The study objectives were (1) to determine the effect of prescribed burning on regeneration of native species from seed, and (2) to establish predictive relationships between easily measured plant traits and seedling establishment rates in the field. The general approach was to sow seeds of target wetland prairie species during the fall into experimental field plots already established at the Danebo Wetland, Eugene, OR. Seedling establishment rates were then compared between burned and unburned plots the following spring. To establish predictive relationships these seedling establishment rates were related to selected seed and seedling traits measured under laboratory conditions. Overall seedling establishment rates showed no significant differences between burned (7.0%) and unburned plots (8.7%), although for the seven species showing positive responses to prescribed burning, the increase was approximately doubled in the burned plots. Burning significantly increased the seedling establishment rates of three species, Wyethia angustifolia, Grindelia integrifolia, and Danthonia californica. Seedling establishment significantly decreased with burning for one species, Sidalcea campestris. For three of the four endangered, threatened or rare species, Aster curtus, Horkelia congesta, and Sidalcea cusickii var. purpurea, seedling establishment rates were smaller in the burned plots compared to the unburned plots, although the differences were not significant. Lomatium bradshawii had no seedlings establish in either the burned or the unburned plots. Plant weight 7 days after germination was the best trait at predicting field seedling establishment rates for both the burned plots and the unburned plots, explaining a significant amount of variation in establishment rates: 70% for the burned plots and 45% for the unburned plots. This model could be used by managers to choose species for sowing in burned and unburned prairies or to estimate seeding rates at burned and unburned sites. Close