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• 1. [Article] Resistance of Selected Wood-Based Materials to Fungal and Termite Attack in Non–Soil Contact Exposures

  The resistance of three naturally durable heartwood species and a stranded giant bamboo product to fungal and termite (Coptotermes formosanus) attack was evaluated at a test site located near Hilo, Hawaii. ...

  Citation × Citation Citation Abstract Copy Title: Resistance of Selected Wood-Based Materials to Fungal and Termite Attack in Non–Soil Contact Exposures Author: Morrell, Jeffrey J. The resistance of three naturally durable heartwood species and a stranded giant bamboo product to fungal and termite (Coptotermes formosanus) attack was evaluated at a test site located near Hilo, Hawaii. Merbau (Intsia bijuga or I. palembanica) and ipe (Tabebuia spp.) were both exceptionally resistant to fungal and termite attack, while western juniper (Juniperus occidentalis) heartwood was slightly less resistant to degradation. The presence of heartwood on western juniper samples had no noticeable effect on the performance of adjacent sapwood. The bamboo decking proved to the least durable of the materials and experienced substantial termite and fungal attack over the 32-month test period. Title: Resistance of Selected Wood-Based Materials to Fungal and Termite Attack in Non–Soil Contact Exposures Author: Morrell, Jeffrey J. The resistance of three naturally durable heartwood species and a stranded giant bamboo product to fungal and termite (Coptotermes formosanus) attack was evaluated at a test site located near Hilo, Hawaii. Merbau (Intsia bijuga or I. palembanica) and ipe (Tabebuia spp.) were both exceptionally resistant to fungal and termite attack, while western juniper (Juniperus occidentalis) heartwood was slightly less resistant to degradation. The presence of heartwood on western juniper samples had no noticeable effect on the performance of adjacent sapwood. The bamboo decking proved to the least durable of the materials and experienced substantial termite and fungal attack over the 32-month test period. Close

• 2. [Article] Vegetation and Small Mammal Responses to Western Juniper (Juniperus occidentalis) Control in Eastern Oregon

  Shrub-steppe ecosystems of western North America provide habitat for many wildlife species, are important components of public and private rangelands, and offer recreational opportunities for millions ...

  Citation × Citation Citation Abstract Copy Title: Vegetation and Small Mammal Responses to Western Juniper (Juniperus occidentalis) Control in Eastern Oregon Author: Morozumi, Corinne N. Shrub-steppe ecosystems of western North America provide habitat for many wildlife species, are important components of public and private rangelands, and offer recreational opportunities for millions of people. They are some of the most vulnerable ecosystems in the United States and have been altered by human activities such as livestock grazing, active fire suppression, conversion to agriculture, and urbanization of the west. Since the late 1800s woody encroachment of piñon-juniper species has also contributed to the loss of shrub-steppe habitat. Presently, land managers remove woody tree species in order to recover shrub-steppe although it is unclear how responses to these management activities differ due to site-specific conditions and existing woodland development. I studied post juniper-thinning responses in eastern Oregon at a wildlife area important as winter range for mule deer (Odocoileus hemionus). I investigated vegetative responses to western juniper (Juniperus occidentalis) thinning across a woodland development gradient and tested for interactive effects of juniper cutting and cattle exclusion. In addition, I explored plant and small mammal successional dynamics after juniper thinning and examined plant community responses within microhabitats created by the felled trees. I compared vegetative responses to juniper thinning as well as cattle exclusion among sites where juniper were subordinate (Phase I), co-dominant (Phase II), and dominant to shrubs and grasses (Phase III, sensu Miller et al. 2005). At the Phase I site, thinning did not increase herbaceous biomass while thinned plots at the Phase II site had 2.71 times more median herbaceous biomass (99.17% CI: 1.37 to 5.37 times more biomass) than unthinned plots. Conversely, herbaceous biomass at the Phase III juniper woodland site was 0.36 times lower in thinned plots when compared to unthinned plots (99.17% CI: 0.17 to 0.78 times less biomass). Unfortunately, many of the responses were driven by exotic species release. Where juniper were subordinate (Phase I), mean percent cover by exotic grass increased by 24.58 percentage points in thinned plots as compared to unthinned plots (98.30% CI: 0.27 to 48.90 percentage points higher). Thinning at the Phase II site increased mean exotic grass cover by an estimated 28.47 percentage points as compared to mean exotic cover in the same plots before treatment (98.30% CI: 4.15 to 52.79 percentage points higher cover). Median native bunchgrass cover at this site was 5.06 times greater after juniper treatment (99.7% CI: 1.78 to 14.35 times higher percent cover). I found few main or interactive effects of cattle exclosure after one year of treatment. Responses to grazing exclosure may take longer to develop. These results indicate that sites within the wildlife area respond differently to juniper management and that exotic grass control will be key to successful shrub-steppe recovery. In addition, I used a time-since-juniper thinning chronoseries consisting of plots cut in 2008, 2009, 2010, 2012, and an adjacent uncut control to explore how shrub-steppe flora and fauna are responding to juniper treatment through time. Shrub cover and seedling density were low in each plot. I recorded the highest seedling abundance (mean of 0.25 seedlings) in the uncut control plot. Median grass cover in the uncut control was 9.50% while in the most recently treated plot (2012) it was 26.75%. Small mammal relative abundance and diversity was low across all time-since-treatment plots though highest in the plot with the greatest time-since-treatment. Deer mouse (Peromyscus maniculatus) was the most abundant species in all plots and accounted for 70-95% of all unique captures. Least chipmunk (Tamias minimus) were present in the plot with greatest time-since-treatment indicating the potential recovery of key native shrubs since these small mammals perform an important seed dispersal role. I also investigated how potential microhabitats created by the felled juniper might support different plant communities. I assessed plant responses within zones created by 1) the felled tree (canopy zone), 2) the original duff zone, and 3) the between - tree interspace zone. The three zones had different plant community compositions as analyzed multivariately with non-metric multidimensional scaling (NMS). Canopy and duff zones were forb - dominated and had less exotic grass invasion while interspace zones were heavily invested with medusahead (Taeniatherum caput-medusae) and ventenata (Ventenata dubia). Exotic herbaceous productivity was lowest under the standing trees of the control plot and high in the interspace of all plots with juniper thinning. In the 2012 cut plot, grass cover was 27.90 percentage points less in the canopy zone compared to the interspace zone (98.75% CI: 17.64 to 38.16 percentage points less grass cover). In the uncut control, median native biomass was 2.08 times greater under the standing trees compared to the interspace (96.25% CI: 1.09 to 3.97 times more biomass). Results from the time-since-treatment chronoseries indicated that exotic grass dominance might be limiting shrub-steppe recovery. Active restoration of shrubs and native grasses may be necessary to address the dominance of exotic grass after juniper thinning. Positive signs of habitat recovery included small mammal responses though I was unable to calculate population estimates due to plot size. Title: Vegetation and Small Mammal Responses to Western Juniper (Juniperus occidentalis) Control in Eastern Oregon Author: Morozumi, Corinne N. Shrub-steppe ecosystems of western North America provide habitat for many wildlife species, are important components of public and private rangelands, and offer recreational opportunities for millions of people. They are some of the most vulnerable ecosystems in the United States and have been altered by human activities such as livestock grazing, active fire suppression, conversion to agriculture, and urbanization of the west. Since the late 1800s woody encroachment of piñon-juniper species has also contributed to the loss of shrub-steppe habitat. Presently, land managers remove woody tree species in order to recover shrub-steppe although it is unclear how responses to these management activities differ due to site-specific conditions and existing woodland development. I studied post juniper-thinning responses in eastern Oregon at a wildlife area important as winter range for mule deer (Odocoileus hemionus). I investigated vegetative responses to western juniper (Juniperus occidentalis) thinning across a woodland development gradient and tested for interactive effects of juniper cutting and cattle exclusion. In addition, I explored plant and small mammal successional dynamics after juniper thinning and examined plant community responses within microhabitats created by the felled trees. I compared vegetative responses to juniper thinning as well as cattle exclusion among sites where juniper were subordinate (Phase I), co-dominant (Phase II), and dominant to shrubs and grasses (Phase III, sensu Miller et al. 2005). At the Phase I site, thinning did not increase herbaceous biomass while thinned plots at the Phase II site had 2.71 times more median herbaceous biomass (99.17% CI: 1.37 to 5.37 times more biomass) than unthinned plots. Conversely, herbaceous biomass at the Phase III juniper woodland site was 0.36 times lower in thinned plots when compared to unthinned plots (99.17% CI: 0.17 to 0.78 times less biomass). Unfortunately, many of the responses were driven by exotic species release. Where juniper were subordinate (Phase I), mean percent cover by exotic grass increased by 24.58 percentage points in thinned plots as compared to unthinned plots (98.30% CI: 0.27 to 48.90 percentage points higher). Thinning at the Phase II site increased mean exotic grass cover by an estimated 28.47 percentage points as compared to mean exotic cover in the same plots before treatment (98.30% CI: 4.15 to 52.79 percentage points higher cover). Median native bunchgrass cover at this site was 5.06 times greater after juniper treatment (99.7% CI: 1.78 to 14.35 times higher percent cover). I found few main or interactive effects of cattle exclosure after one year of treatment. Responses to grazing exclosure may take longer to develop. These results indicate that sites within the wildlife area respond differently to juniper management and that exotic grass control will be key to successful shrub-steppe recovery. In addition, I used a time-since-juniper thinning chronoseries consisting of plots cut in 2008, 2009, 2010, 2012, and an adjacent uncut control to explore how shrub-steppe flora and fauna are responding to juniper treatment through time. Shrub cover and seedling density were low in each plot. I recorded the highest seedling abundance (mean of 0.25 seedlings) in the uncut control plot. Median grass cover in the uncut control was 9.50% while in the most recently treated plot (2012) it was 26.75%. Small mammal relative abundance and diversity was low across all time-since-treatment plots though highest in the plot with the greatest time-since-treatment. Deer mouse (Peromyscus maniculatus) was the most abundant species in all plots and accounted for 70-95% of all unique captures. Least chipmunk (Tamias minimus) were present in the plot with greatest time-since-treatment indicating the potential recovery of key native shrubs since these small mammals perform an important seed dispersal role. I also investigated how potential microhabitats created by the felled juniper might support different plant communities. I assessed plant responses within zones created by 1) the felled tree (canopy zone), 2) the original duff zone, and 3) the between - tree interspace zone. The three zones had different plant community compositions as analyzed multivariately with non-metric multidimensional scaling (NMS). Canopy and duff zones were forb - dominated and had less exotic grass invasion while interspace zones were heavily invested with medusahead (Taeniatherum caput-medusae) and ventenata (Ventenata dubia). Exotic herbaceous productivity was lowest under the standing trees of the control plot and high in the interspace of all plots with juniper thinning. In the 2012 cut plot, grass cover was 27.90 percentage points less in the canopy zone compared to the interspace zone (98.75% CI: 17.64 to 38.16 percentage points less grass cover). In the uncut control, median native biomass was 2.08 times greater under the standing trees compared to the interspace (96.25% CI: 1.09 to 3.97 times more biomass). Results from the time-since-treatment chronoseries indicated that exotic grass dominance might be limiting shrub-steppe recovery. Active restoration of shrubs and native grasses may be necessary to address the dominance of exotic grass after juniper thinning. Positive signs of habitat recovery included small mammal responses though I was unable to calculate population estimates due to plot size. Close

• 3. [Article] Climate-growth relationships of western juniper and ponderosa pine at the pine-woodland ecotone in southern Oregon

  Pinyon-juniper woodlands throughout the western U.S. have expanded rapidly following European settlement during the late 19th century. In central and eastern Oregon, western juniper (Juniperus occidentalis ...

  Citation × Citation Citation Abstract Copy Title: Climate-growth relationships of western juniper and ponderosa pine at the pine-woodland ecotone in southern Oregon Author: Knutson, Kevin C. Pinyon-juniper woodlands throughout the western U.S. have expanded rapidly following European settlement during the late 19th century. In central and eastern Oregon, western juniper (Juniperus occidentalis var. occidentalis Hook.) encroachment has been previously documented in the sagebrush steppe and upper elevation aspen communities. While these vegetation changes and dynamics have been extensively studied, the ability to accurately predict future effects on vegetation structure in response to climatic effects requires the identification of climatic factors that most directly regulate growth across variations in spatial features, particularly when future climate change is likely. I sought to address this problem at the pine-woodland ecotone using tree-ring data for western juniper and ponderosa pine (Pinus ponderosa Dougl. Ex Loud.) from the Fremont National Forest of central Oregon. Site and soil-derived growth chronologies for the period 1950 to 2000 were developed for both species using standard dendrochronological techniques at 17 sites within the Fremont National Forest stratified by elevation (1584 and 1865 meters). Analysis of the climatic relationship with site and soil-derived chronologies using correlation and linear regression revealed that, similar to previous studies in Oregon, radial growth in both species is highly dependent on winter-spring precipitation events that recharge growing-season soil water. Ordinations of site- and soil-derived chronology principal components (PCA) scores for both species identified distinct gradients in growth patterns related to elevation, slope, and soil infiltration class. Low elevation sites had reduced growth in the nine driest years during 1950-2000 compared to high elevation sites. Slope was also positively associated with the ratio of standardized growth in wet years to dry years (W:D), a measure of climate sensitivity. Tree growth of both species on coarse-textured soils with rapid infiltration rates was more climate-sensitive (higher values of W:D) and was also reduced during drought years when compared to other infiltration classes. These findings suggest potentially significant variation in growth responses to future climate changes across distinct local features at the pine-woodland ecotone. Juniper and pine radial growth at sites that feature poor water-holding capacities (i.e., low elevation, steep slopes, and coarse-textured soils) can be expected to be most sensitive to future drought and climate fluctuations in southern Oregon. Title: Climate-growth relationships of western juniper and ponderosa pine at the pine-woodland ecotone in southern Oregon Author: Knutson, Kevin C. Pinyon-juniper woodlands throughout the western U.S. have expanded rapidly following European settlement during the late 19th century. In central and eastern Oregon, western juniper (Juniperus occidentalis var. occidentalis Hook.) encroachment has been previously documented in the sagebrush steppe and upper elevation aspen communities. While these vegetation changes and dynamics have been extensively studied, the ability to accurately predict future effects on vegetation structure in response to climatic effects requires the identification of climatic factors that most directly regulate growth across variations in spatial features, particularly when future climate change is likely. I sought to address this problem at the pine-woodland ecotone using tree-ring data for western juniper and ponderosa pine (Pinus ponderosa Dougl. Ex Loud.) from the Fremont National Forest of central Oregon. Site and soil-derived growth chronologies for the period 1950 to 2000 were developed for both species using standard dendrochronological techniques at 17 sites within the Fremont National Forest stratified by elevation (1584 and 1865 meters). Analysis of the climatic relationship with site and soil-derived chronologies using correlation and linear regression revealed that, similar to previous studies in Oregon, radial growth in both species is highly dependent on winter-spring precipitation events that recharge growing-season soil water. Ordinations of site- and soil-derived chronology principal components (PCA) scores for both species identified distinct gradients in growth patterns related to elevation, slope, and soil infiltration class. Low elevation sites had reduced growth in the nine driest years during 1950-2000 compared to high elevation sites. Slope was also positively associated with the ratio of standardized growth in wet years to dry years (W:D), a measure of climate sensitivity. Tree growth of both species on coarse-textured soils with rapid infiltration rates was more climate-sensitive (higher values of W:D) and was also reduced during drought years when compared to other infiltration classes. These findings suggest potentially significant variation in growth responses to future climate changes across distinct local features at the pine-woodland ecotone. Juniper and pine radial growth at sites that feature poor water-holding capacities (i.e., low elevation, steep slopes, and coarse-textured soils) can be expected to be most sensitive to future drought and climate fluctuations in southern Oregon. Close