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• 1. [Article] Invertebrate Community Composition Across Inundation Regimes and Its Potential to Reduce Plant Stress

  Appreciation of the ecological and economic values associated with healthy salt marshes has led to a recent rise in the number of marshes that are being targeted for restoration by dike removal. The success ...

  Citation × Citation Citation Abstract Copy Title: Invertebrate Community Composition Across Inundation Regimes and Its Potential to Reduce Plant Stress Author: Lawson, Inez Ilicia Year: 2017 Appreciation of the ecological and economic values associated with healthy salt marshes has led to a recent rise in the number of marshes that are being targeted for restoration by dike removal. The success of restoration is often measured by the return of marsh plants, though this overlooks a key component of salt marshes, that of the invertebrate community within marsh sediments. To evaluate the short-term recovery of these invertebrates, sediment cores were collected across an elevational gradient in a recent dike removal marsh, one and two years post removal, and a nearby reference marsh. Abundance, richness and diversity as well as morphospecies community composition were compared across treatment groups (Reference, Removal) and elevation zone (High Marsh, Low Marsh). Morphospecies richness, abundance and diversity were significantly higher in Low Marsh samples than in High Marsh samples, though no statistically significant differences were found across treatments of the same elevation (e.g., Reference Low Marsh versus Removal Low Marsh). Pair-wise ANOSIM results found significant differences between community compositions across treatments, specifically Reference Low Marsh and Removal Low Marsh. The marsh edge, the lowest point of vascular plant growth before transitioning to tide flats, is considered a high stress environment for emergent vegetation. Plant establishment and survival in this low elevation zone is limited by the tolerance to inundation duration and frequency and anoxic sediments. Bioturbation and burrowing by macroinvertebrates increases the surface area exposed to surface water for gas exchange, increasing the depth of the redox potential discontinuity layer. Crabs that make stable, maintained burrows have been shown to increase oxygen penetration into sediment, improving plant productivity. Such crabs are not found in salt marshes of the Pacific Northwest of North America. However, other burrowing invertebrates may have a positive impact on plant health in these areas by reducing abiotic stress due to anoxic sediments, thereby allowing plants to establish and survive lower in the intertidal zone. To assess this potential relationship, study plots of Distichlis spicata were selected at equivalent elevations at the lowest point of plant establishment at the marsh edge. Focal plant rhizomes were severed from upland ramets and assigned an invertebrate abundance treatment based on a visual burrow count surrounding each plant (9 cm diameter). Focal plants were visited monthly from July to September 2016, plant health variables of chlorophyll content and chlorophyll fluorescence (photosynthetic efficiency), and sediment ORP readings were collected. Plant survivorship was significantly higher in plots with invertebrates, 96% of plants in "With Invertebrate" plots and 50% of plants in "No Invertebrates" plots survived the duration of the study. Plant health (chlorophyll content and chlorophyll fluorescence) generally increased with increased invertebrate presence though, not statistically significant. There may be potential for improved plant productivity and resilience to plants at the marsh edge due to invertebrate burrowing activity. This benefit could help mitigate projected losses in plant productivity due to sea level rise, though more research is needed to investigate the mechanism by which these invertebrates confer a health benefit to plants at the marsh edge. Title: Invertebrate Community Composition Across Inundation Regimes and Its Potential to Reduce Plant Stress Author: Lawson, Inez Ilicia Year: 2017 Appreciation of the ecological and economic values associated with healthy salt marshes has led to a recent rise in the number of marshes that are being targeted for restoration by dike removal. The success of restoration is often measured by the return of marsh plants, though this overlooks a key component of salt marshes, that of the invertebrate community within marsh sediments. To evaluate the short-term recovery of these invertebrates, sediment cores were collected across an elevational gradient in a recent dike removal marsh, one and two years post removal, and a nearby reference marsh. Abundance, richness and diversity as well as morphospecies community composition were compared across treatment groups (Reference, Removal) and elevation zone (High Marsh, Low Marsh). Morphospecies richness, abundance and diversity were significantly higher in Low Marsh samples than in High Marsh samples, though no statistically significant differences were found across treatments of the same elevation (e.g., Reference Low Marsh versus Removal Low Marsh). Pair-wise ANOSIM results found significant differences between community compositions across treatments, specifically Reference Low Marsh and Removal Low Marsh. The marsh edge, the lowest point of vascular plant growth before transitioning to tide flats, is considered a high stress environment for emergent vegetation. Plant establishment and survival in this low elevation zone is limited by the tolerance to inundation duration and frequency and anoxic sediments. Bioturbation and burrowing by macroinvertebrates increases the surface area exposed to surface water for gas exchange, increasing the depth of the redox potential discontinuity layer. Crabs that make stable, maintained burrows have been shown to increase oxygen penetration into sediment, improving plant productivity. Such crabs are not found in salt marshes of the Pacific Northwest of North America. However, other burrowing invertebrates may have a positive impact on plant health in these areas by reducing abiotic stress due to anoxic sediments, thereby allowing plants to establish and survive lower in the intertidal zone. To assess this potential relationship, study plots of Distichlis spicata were selected at equivalent elevations at the lowest point of plant establishment at the marsh edge. Focal plant rhizomes were severed from upland ramets and assigned an invertebrate abundance treatment based on a visual burrow count surrounding each plant (9 cm diameter). Focal plants were visited monthly from July to September 2016, plant health variables of chlorophyll content and chlorophyll fluorescence (photosynthetic efficiency), and sediment ORP readings were collected. Plant survivorship was significantly higher in plots with invertebrates, 96% of plants in "With Invertebrate" plots and 50% of plants in "No Invertebrates" plots survived the duration of the study. Plant health (chlorophyll content and chlorophyll fluorescence) generally increased with increased invertebrate presence though, not statistically significant. There may be potential for improved plant productivity and resilience to plants at the marsh edge due to invertebrate burrowing activity. This benefit could help mitigate projected losses in plant productivity due to sea level rise, though more research is needed to investigate the mechanism by which these invertebrates confer a health benefit to plants at the marsh edge. Close

• 2. [Article] Investigations of the Air-Water Interface: A Structural Analysis of Metallic Surface Films and Aquatic Surface Films by Comparative Microscopy

  The air-water interface is an important natural boundary layer that has been neglected as an area of environmental field research. This study establishes that comparative microscopy can be an effective ...

  Citation × Citation Citation Abstract Copy Title: Investigations of the Air-Water Interface: A Structural Analysis of Metallic Surface Films and Aquatic Surface Films by Comparative Microscopy Author: Smith, Randall William Year: 2015 The air-water interface is an important natural boundary layer that has been neglected as an area of environmental field research. This study establishes that comparative microscopy can be an effective environmental method, and establishes that the term metallic surface films, is a more accurate descriptor than iron oxide surface films. This research shows that surface films are complex, often with layered structure, serve as habitat for significant biota, and act as a point of mineralization to several transition metal elements including manganese, iron, copper, nickel and zinc. This study demonstrates that surface films form under several conditions and can have diverse morphology. Activity of biota, microbes, particularly diatoms, suggests that bacteria and cyanobacteria integrate into the film often in patches, represented by forms and casts. Analytical imaging is used to document and compare film morphology and structures, using scanning electron microscopy, photoemission electron microscopy and transmission electron microscopy with elemental analysis by energy dispersive spectroscopy to confirm the hypothesis. Instrument parameters and strengths are reviewed. Component layers of a copper/zinc film were used to confirm metallic layers and elemental distribution. Bacterial casts were used to confirm film interaction, and to show entrainment and enrichment of the film to incorporate autochthonous and allochthonous materials into the films themselves. Most samples were from Oregon selected sites, with some samples from Maryland and Barbados. Title: Investigations of the Air-Water Interface: A Structural Analysis of Metallic Surface Films and Aquatic Surface Films by Comparative Microscopy Author: Smith, Randall William Year: 2015 The air-water interface is an important natural boundary layer that has been neglected as an area of environmental field research. This study establishes that comparative microscopy can be an effective environmental method, and establishes that the term metallic surface films, is a more accurate descriptor than iron oxide surface films. This research shows that surface films are complex, often with layered structure, serve as habitat for significant biota, and act as a point of mineralization to several transition metal elements including manganese, iron, copper, nickel and zinc. This study demonstrates that surface films form under several conditions and can have diverse morphology. Activity of biota, microbes, particularly diatoms, suggests that bacteria and cyanobacteria integrate into the film often in patches, represented by forms and casts. Analytical imaging is used to document and compare film morphology and structures, using scanning electron microscopy, photoemission electron microscopy and transmission electron microscopy with elemental analysis by energy dispersive spectroscopy to confirm the hypothesis. Instrument parameters and strengths are reviewed. Component layers of a copper/zinc film were used to confirm metallic layers and elemental distribution. Bacterial casts were used to confirm film interaction, and to show entrainment and enrichment of the film to incorporate autochthonous and allochthonous materials into the films themselves. Most samples were from Oregon selected sites, with some samples from Maryland and Barbados. Close

• 3. [Article] Hydrogeochemical Characterization of the Alvord Valley Known Geothermal Resources Area, Harney County, Oregon

  The Alvord Valley Known Geothermal Resources Area (KGRA) , located east of the Steens Mountain-Pueblo Mountains fault block in southeastern Oregon, is within the northern Basin and Range province. This ...

  Citation × Citation Citation Abstract Copy Title: Hydrogeochemical Characterization of the Alvord Valley Known Geothermal Resources Area, Harney County, Oregon Author: St. John, Anna Maria Year: 1993 The Alvord Valley Known Geothermal Resources Area (KGRA) , located east of the Steens Mountain-Pueblo Mountains fault block in southeastern Oregon, is within the northern Basin and Range province. This investigation focuses on three thermal areas in the Alvord Basin: Borax Lake and the hot springs north of Borax Lake, Alvord Hot Springs and Mickey Springs. Mickey Springs and the springs north of Borax Lake are boiling at the surface (94 and 95° C, respectively). Inflow temperatures to Borax Lake, measured at a depth of 30 m, are greater than 100° C. Surface temperatures for Alvord Hot Springs and a flowing well northeast of Borax Lake are 78 and 59° C, respectively. Thermal fluids issue from Quaternary lacustrine and alluvial deposits. While silica sinter deposits are present at all three thermal areas, sinter is not presently being deposited. Minor calcite is being deposited at the springs north of Borax Lake. The springs discharge from N to NEstriking, high-angle, basin-bounding faults along the base of Steens Mountain and Mickey Mountain and NE-striking intrabasinal faults south of Alvord Lake. The thermal waters are dilute sodium-bicarbonate waters with significant amounts of sulfate and chloride. Conservative element plots (B, F, and Li vs. Cl) indicate good correlation between Cl and the other conservative elements. These correlations could result from mixing of thermal water with a dilute cold water or fluid evolution due to increased fluid-rock interaction, evaporation, and steam loss due to boiling. The small variations in chloride concentrations of thermal fluids during the sampling period argues against mixing of thermal fluids with cold water. The geothermal system is a hot-water rather than a vapordominated system. The ỎD content of thermal fluids is similar to the ỎD content of local cold water wells, springs, basinal pore fluids at a depth of 4 to 5 m, and perennial streams. Similarities in ỎD values indicate recharge for geothermal fluids is precipitation from the Steens Mountain fault block. The Ỏ18 content of thermal fluids is shifted 2 to 3°/oo to the right of the world meteoric water line indicating fluid-rock interaction at elevated temperatures in the reservoir. Tritium contents indicate relatively long residence times and/or low-velocity circulation of meteoric water through basement rocks. Values range from 0 to 0.25 T.U. The application of two end-member models, which calculate fluid residence times, generate a minimum of 57 years and a maximum of greater than 10,000 years. Estimated reservoir temperatures based on cation and silica geothermometry are between 170 and 200°C. Oxygensulfate isotope geothermometer estimates indicate reservoir temperatures between 198 and 207' C for Borax Lake and Alvord Hot Springs. Mickey Springs and a flowing well northeast of Borax Lake yield temperature estimates of 168 and 150° C, respectively. These values indicate partial reequilibration of the isotopic system. The Ỏ13C contents of carbon dioxide and methane of gas discharges from the thermal areas are similar to geothermal fluids from other sites. The Ỏ13C of methane indicate "normal" geothermal methane for Alvord Hot Springs and Mickey Springs (-27.8 and -27.6, respectively). The Ỏ13C of CH4 for springs north of Borax Lake (-33.6) indicates a small amount of thermogenic methane may be contributed by thermal alteration of organics in basinal sediments. The Ỏ13C contents for C02 at Alvord Hot Springs and Borax Lake are within the range expected for atmospheric, fumarolic, or mantle derived C02 (-6.5 and -6.6, respectively). The Ỏ13C content of C02 from Mickey Springs is isotopically lighter than gas released from fumaroles or the mantle (-9.4). N2/Ar ratios for Mickey Springs and Borax Lake gases (39.2 and 40.8, respectively) indicate interaction with airsaturated ground water during flow through the the zone of aeration. Helium is enriched relative to Ar and N2 in gas discharges from Alvord Hot Springs, indicating longer fluid residence times and/or increased crustal interaction at high temperatures. Ratios of B/Cl indicate the fluid reservoir is hosted in volcanic rocks. The Li/Cs ratios for the Borax Lake thermal area are consistent with a reservoir located in rhyoli tic rocks. The 228Ra/226Ra content of Borax Lake thermal fluids (1.14 ± 0.13 dpm/kg) indicates interaction with volcanic rocks for Borax Lake. The 228Ra/226Ra content of thermal fluids from Alvord Hot Springs and Mickey Springs (0.38~0.02 and 0.17 ~ 0.09) are lower than those expected for volcanic rocks and may indicate local uranium accumulation in the reservoir or zones of upflow. The 87Sr / 86Sr values for thermal waters and stratigraphic uni ts indicate the fluid reservoir is located in volcanic rocks beneath Steens Basalt. Equilibration of fluids in these units argues for thermal water circulation depths of 2 to 2.5 km in the Borax Lake thermal area, greater than 3 km in the Alvord Hot Springs area and 1 to 2 km in the Mickey Springs area. Data presented in this study do not preclude a single large deep reservoir system discharging at these three thermal areas in the Alvord basin. Differences in the chemical and isotopic composition of discharge from the three thermal areas are produced during upf low from the reservoir. During upflow, thermal waters follow a complex pathway of vertical and lateral fractures which includes short residence times in shallow reservoirs before reaching the surface. Boiling, mixing with condensate, oxidation, mixing with 1-3% tritium-bearing, near-surface cold water, relative differences in flow rate and volume, and slow cooling without vigorous boiling are processes that modify fluid composition during upflow from the deep geothermal reservoir. Title: Hydrogeochemical Characterization of the Alvord Valley Known Geothermal Resources Area, Harney County, Oregon Author: St. John, Anna Maria Year: 1993 The Alvord Valley Known Geothermal Resources Area (KGRA) , located east of the Steens Mountain-Pueblo Mountains fault block in southeastern Oregon, is within the northern Basin and Range province. This investigation focuses on three thermal areas in the Alvord Basin: Borax Lake and the hot springs north of Borax Lake, Alvord Hot Springs and Mickey Springs. Mickey Springs and the springs north of Borax Lake are boiling at the surface (94 and 95° C, respectively). Inflow temperatures to Borax Lake, measured at a depth of 30 m, are greater than 100° C. Surface temperatures for Alvord Hot Springs and a flowing well northeast of Borax Lake are 78 and 59° C, respectively. Thermal fluids issue from Quaternary lacustrine and alluvial deposits. While silica sinter deposits are present at all three thermal areas, sinter is not presently being deposited. Minor calcite is being deposited at the springs north of Borax Lake. The springs discharge from N to NEstriking, high-angle, basin-bounding faults along the base of Steens Mountain and Mickey Mountain and NE-striking intrabasinal faults south of Alvord Lake. The thermal waters are dilute sodium-bicarbonate waters with significant amounts of sulfate and chloride. Conservative element plots (B, F, and Li vs. Cl) indicate good correlation between Cl and the other conservative elements. These correlations could result from mixing of thermal water with a dilute cold water or fluid evolution due to increased fluid-rock interaction, evaporation, and steam loss due to boiling. The small variations in chloride concentrations of thermal fluids during the sampling period argues against mixing of thermal fluids with cold water. The geothermal system is a hot-water rather than a vapordominated system. The ỎD content of thermal fluids is similar to the ỎD content of local cold water wells, springs, basinal pore fluids at a depth of 4 to 5 m, and perennial streams. Similarities in ỎD values indicate recharge for geothermal fluids is precipitation from the Steens Mountain fault block. The Ỏ18 content of thermal fluids is shifted 2 to 3°/oo to the right of the world meteoric water line indicating fluid-rock interaction at elevated temperatures in the reservoir. Tritium contents indicate relatively long residence times and/or low-velocity circulation of meteoric water through basement rocks. Values range from 0 to 0.25 T.U. The application of two end-member models, which calculate fluid residence times, generate a minimum of 57 years and a maximum of greater than 10,000 years. Estimated reservoir temperatures based on cation and silica geothermometry are between 170 and 200°C. Oxygensulfate isotope geothermometer estimates indicate reservoir temperatures between 198 and 207' C for Borax Lake and Alvord Hot Springs. Mickey Springs and a flowing well northeast of Borax Lake yield temperature estimates of 168 and 150° C, respectively. These values indicate partial reequilibration of the isotopic system. The Ỏ13C contents of carbon dioxide and methane of gas discharges from the thermal areas are similar to geothermal fluids from other sites. The Ỏ13C of methane indicate "normal" geothermal methane for Alvord Hot Springs and Mickey Springs (-27.8 and -27.6, respectively). The Ỏ13C of CH4 for springs north of Borax Lake (-33.6) indicates a small amount of thermogenic methane may be contributed by thermal alteration of organics in basinal sediments. The Ỏ13C contents for C02 at Alvord Hot Springs and Borax Lake are within the range expected for atmospheric, fumarolic, or mantle derived C02 (-6.5 and -6.6, respectively). The Ỏ13C content of C02 from Mickey Springs is isotopically lighter than gas released from fumaroles or the mantle (-9.4). N2/Ar ratios for Mickey Springs and Borax Lake gases (39.2 and 40.8, respectively) indicate interaction with airsaturated ground water during flow through the the zone of aeration. Helium is enriched relative to Ar and N2 in gas discharges from Alvord Hot Springs, indicating longer fluid residence times and/or increased crustal interaction at high temperatures. Ratios of B/Cl indicate the fluid reservoir is hosted in volcanic rocks. The Li/Cs ratios for the Borax Lake thermal area are consistent with a reservoir located in rhyoli tic rocks. The 228Ra/226Ra content of Borax Lake thermal fluids (1.14 ± 0.13 dpm/kg) indicates interaction with volcanic rocks for Borax Lake. The 228Ra/226Ra content of thermal fluids from Alvord Hot Springs and Mickey Springs (0.38~0.02 and 0.17 ~ 0.09) are lower than those expected for volcanic rocks and may indicate local uranium accumulation in the reservoir or zones of upflow. The 87Sr / 86Sr values for thermal waters and stratigraphic uni ts indicate the fluid reservoir is located in volcanic rocks beneath Steens Basalt. Equilibration of fluids in these units argues for thermal water circulation depths of 2 to 2.5 km in the Borax Lake thermal area, greater than 3 km in the Alvord Hot Springs area and 1 to 2 km in the Mickey Springs area. Data presented in this study do not preclude a single large deep reservoir system discharging at these three thermal areas in the Alvord basin. Differences in the chemical and isotopic composition of discharge from the three thermal areas are produced during upf low from the reservoir. During upflow, thermal waters follow a complex pathway of vertical and lateral fractures which includes short residence times in shallow reservoirs before reaching the surface. Boiling, mixing with condensate, oxidation, mixing with 1-3% tritium-bearing, near-surface cold water, relative differences in flow rate and volume, and slow cooling without vigorous boiling are processes that modify fluid composition during upflow from the deep geothermal reservoir. Close

• 4. [Article] Long-Term Exposure to Fluoxetine Reduces Growth and Reproductive Potential in the Dominant Rocky Intertidal Mussel, Mytilus californianus

  Environmental stressors shape community composition and ecosystem functioning. Contaminants such as pharmaceuticals are of increasing concern as an environmental stressor due to their persistence in surface ...

  Citation × Citation Citation Abstract Copy Title: Long-Term Exposure to Fluoxetine Reduces Growth and Reproductive Potential in the Dominant Rocky Intertidal Mussel, Mytilus californianus Author: Peters, Joseph R., Granek, Elise F. Year: 2016 Environmental stressors shape community composition and ecosystem functioning. Contaminants such as pharmaceuticals are of increasing concern as an environmental stressor due to their persistence in surface waters worldwide. Limited attention has been paid to the effects of pharmaceuticals on marine life, despite widespread detection of these contaminants in the marine environment. Of the existing studies, the majority assess the negative effects of pharmaceuticals over an exposure period of 30 days or less and focus on cellular and subcellular biomarkers. Longer studies are required to determine if chronic contaminant exposure poses risks to marine life at environmentally relevant concentrations; and examination of whole organism effects are necessary to identify potential community-level consequences in estuarine and marine ecosystems. We conducted a long-term exposure study (107 days) with the anti-depressant pharmaceutical, fluoxetine (the active constituent in Prozac®) to determine whether minimal concentrations affected whole organism metrics in the California mussel, Mytilus californianus. We measured algal clearance rates, mussel growth, and the gonadosomatic index, a measure of reproductive health. We found that fluoxetine negatively affects all measured characteristics, however many effects were mediated by length of exposure. Our results fill an important data gap, highlighting organism-level effects of chronic exposure periods; such data more explicitly identify the overall impacts of pharmaceuticals and other contaminants on marine communities and ecosystems. Title: Long-Term Exposure to Fluoxetine Reduces Growth and Reproductive Potential in the Dominant Rocky Intertidal Mussel, Mytilus californianus Author: Peters, Joseph R., Granek, Elise F. Year: 2016 Environmental stressors shape community composition and ecosystem functioning. Contaminants such as pharmaceuticals are of increasing concern as an environmental stressor due to their persistence in surface waters worldwide. Limited attention has been paid to the effects of pharmaceuticals on marine life, despite widespread detection of these contaminants in the marine environment. Of the existing studies, the majority assess the negative effects of pharmaceuticals over an exposure period of 30 days or less and focus on cellular and subcellular biomarkers. Longer studies are required to determine if chronic contaminant exposure poses risks to marine life at environmentally relevant concentrations; and examination of whole organism effects are necessary to identify potential community-level consequences in estuarine and marine ecosystems. We conducted a long-term exposure study (107 days) with the anti-depressant pharmaceutical, fluoxetine (the active constituent in Prozac®) to determine whether minimal concentrations affected whole organism metrics in the California mussel, Mytilus californianus. We measured algal clearance rates, mussel growth, and the gonadosomatic index, a measure of reproductive health. We found that fluoxetine negatively affects all measured characteristics, however many effects were mediated by length of exposure. Our results fill an important data gap, highlighting organism-level effects of chronic exposure periods; such data more explicitly identify the overall impacts of pharmaceuticals and other contaminants on marine communities and ecosystems. Close

• 5. [Article] Water Flow through Temperate Glaciers

  Understanding water movement through a glacier is fundamental to several critical issues in glaciology, including glacier dynamics, glacier-induced floods, and the prediction of runoff from glacierized ...

  Citation × Citation Citation Abstract Copy Title: Water Flow through Temperate Glaciers Author: Fountain, Andrew G., Walder, Joseph S. Year: 1998 Understanding water movement through a glacier is fundamental to several critical issues in glaciology, including glacier dynamics, glacier-induced floods, and the prediction of runoff from glacierized drainage basins. To this end we have synthesized a conceptual model of water movement through a temperate glacier from the surface to the outlet stream. Processes that regulate the rate and distribution of water input at the glacier surface and that regulate water movement from the surface to the bed play important but commonly neglected roles in glacier hydrology. Where a glacier is covered by a layer of porous, permeable firn (the accumulation zone), the flux of water to the glacier interior varies slowly because the firn temporarily stores water and thereby smooths out variations in the supply rate. In the firn-free ablation zone, in contrast, the flux of water into the glacier depends directly on the rate of surface melt or rainfall and therefore varies greatly in time. Water moves from the surface to the bed through an upward branching arborescent network consisting of both steeply inclined conduits, formed by the enlargement of intergranular veins, and gently inclined conduits, spawned by water flow along the bottoms of near-surface fractures (crevasses). Englacial drainage conduits deliver water to the glacier bed at a limited number of points, probably a long distance downglacier of where water enters the glacier. Englacial conduits supplied from the accumulation zone are quasi steady state features that convey the slowly varying water flux delivered via the firn. Their size adjusts so that they are usually full of water and flow is pressurized. In contrast, water flow in englacial conduits supplied from the ablation area is pressurized only near times of peak daily flow or during rainstorms; flow is otherwise in an open-channel configuration. The subglacial drainage system typically consists of several elements that are distinct both morphologically and hydrologically. An upglacier branching, arborescent network of channels incised into the basal ice conveys water rapidly. Much of the water flux to the bed probably enters directly into the arborescent channel network, which covers only a small fraction of the glacier bed. More extensive spatially is a nonarborescent network, which commonly includes cavities (gaps between the glacier sole and bed), channels incised into the bed, and a layer of permeable sediment. The nonarborescent network conveys water slowly and is usually poorly connected to the arborescent system. The arborescent channel network largely collapses during winter but reforms in the spring as the first flush of meltwater to the bed destabilizes the cavities within the nonarborescent network. The volume of water stored by a glacier varies diurnally and seasonally. Small, temperate alpine glaciers seem to attain a maximum seasonal water storage of approximately 200 mm of water averaged over the area of the glacier bed, with daily fluctuations of as much as 20-30 mm. The likely storage capacity of subglacial cavities is insufficient to account for estimated stored water volumes, so most water storage may actually occur englacially. Stored water may also be released abruptly and catastrophically in the form of outburst floods. Title: Water Flow through Temperate Glaciers Author: Fountain, Andrew G., Walder, Joseph S. Year: 1998 Understanding water movement through a glacier is fundamental to several critical issues in glaciology, including glacier dynamics, glacier-induced floods, and the prediction of runoff from glacierized drainage basins. To this end we have synthesized a conceptual model of water movement through a temperate glacier from the surface to the outlet stream. Processes that regulate the rate and distribution of water input at the glacier surface and that regulate water movement from the surface to the bed play important but commonly neglected roles in glacier hydrology. Where a glacier is covered by a layer of porous, permeable firn (the accumulation zone), the flux of water to the glacier interior varies slowly because the firn temporarily stores water and thereby smooths out variations in the supply rate. In the firn-free ablation zone, in contrast, the flux of water into the glacier depends directly on the rate of surface melt or rainfall and therefore varies greatly in time. Water moves from the surface to the bed through an upward branching arborescent network consisting of both steeply inclined conduits, formed by the enlargement of intergranular veins, and gently inclined conduits, spawned by water flow along the bottoms of near-surface fractures (crevasses). Englacial drainage conduits deliver water to the glacier bed at a limited number of points, probably a long distance downglacier of where water enters the glacier. Englacial conduits supplied from the accumulation zone are quasi steady state features that convey the slowly varying water flux delivered via the firn. Their size adjusts so that they are usually full of water and flow is pressurized. In contrast, water flow in englacial conduits supplied from the ablation area is pressurized only near times of peak daily flow or during rainstorms; flow is otherwise in an open-channel configuration. The subglacial drainage system typically consists of several elements that are distinct both morphologically and hydrologically. An upglacier branching, arborescent network of channels incised into the basal ice conveys water rapidly. Much of the water flux to the bed probably enters directly into the arborescent channel network, which covers only a small fraction of the glacier bed. More extensive spatially is a nonarborescent network, which commonly includes cavities (gaps between the glacier sole and bed), channels incised into the bed, and a layer of permeable sediment. The nonarborescent network conveys water slowly and is usually poorly connected to the arborescent system. The arborescent channel network largely collapses during winter but reforms in the spring as the first flush of meltwater to the bed destabilizes the cavities within the nonarborescent network. The volume of water stored by a glacier varies diurnally and seasonally. Small, temperate alpine glaciers seem to attain a maximum seasonal water storage of approximately 200 mm of water averaged over the area of the glacier bed, with daily fluctuations of as much as 20-30 mm. The likely storage capacity of subglacial cavities is insufficient to account for estimated stored water volumes, so most water storage may actually occur englacially. Stored water may also be released abruptly and catastrophically in the form of outburst floods. Close

• 6. [Article] Recreational Boats as Potential Vectors of Marine Organisms at an Invasion Hotspot

  With more than 200 aquatic nonindigenous species (NIS), San Francisco Bay (California, USA) is among the world’s most invaded harbors. Hard-substratum benthic (biofouling) organisms, which dominate NIS ...

  Citation × Citation Citation Abstract Copy Title: Recreational Boats as Potential Vectors of Marine Organisms at an Invasion Hotspot Author: Davidson, Ian C., Zabin, Chela J., Chang, Andrew L., Brown, Christopher W., Sytsma, Mark, Ruiz, Gregory M. Year: 2010 With more than 200 aquatic nonindigenous species (NIS), San Francisco Bay (California, USA) is among the world’s most invaded harbors. Hard-substratum benthic (biofouling) organisms, which dominate NIS richness, have arrived primarily as a result of shipping and aquaculture activity over past centuries. To date there has been no assessment of the leisure craft vector in the Bay. We aimed to characterize (1) biofouling on boats’ submerged surfaces and (2) boater behavior likely to affect the risk of NIS transfers. We used an underwater pole-cam, specimen collections, and a boater questionnaire to quantify the extent and composition of biofouling on recreational boats and to evaluate boater behavior at a subset of the Bay’s marinas. Several NIS, already established within the Bay, were recorded from vessel hulls, including the bryozoans Bugula neritina, Membranipora chesapeakensis and Watersipora sp., the ascidians Botrylloides violaceus, Styela clava and Ciona intestinalis, the polychaete Ficopomatus enigmaticus, and the sponge Clathria prolifera. Only 16% of questionnaire respondents had traveled to sites outside the Bay in the previous 12 mo. Frequency of hull painting and cleaning varied substantially, but we did not find strong patterns of biofouling extent associated with hull husbandry or boat usage. The potential for within-Bay and coastwise regional spread of NIS is high, and recreational boats probably interact in close proximity to other vectors (e.g. commercial ships), causing a ratchet effect of vector events; however, there remains a gap in understanding the levels and condition of biofouling on transient boats. Transient vessels from San Francisco Bay and other West Coast sites should be the focus of future studies to evaluate the extent to which organisms are being transferred among bays and how vector management could be applied to prevent NIS transfers and impacts. Title: Recreational Boats as Potential Vectors of Marine Organisms at an Invasion Hotspot Author: Davidson, Ian C., Zabin, Chela J., Chang, Andrew L., Brown, Christopher W., Sytsma, Mark, Ruiz, Gregory M. Year: 2010 With more than 200 aquatic nonindigenous species (NIS), San Francisco Bay (California, USA) is among the world’s most invaded harbors. Hard-substratum benthic (biofouling) organisms, which dominate NIS richness, have arrived primarily as a result of shipping and aquaculture activity over past centuries. To date there has been no assessment of the leisure craft vector in the Bay. We aimed to characterize (1) biofouling on boats’ submerged surfaces and (2) boater behavior likely to affect the risk of NIS transfers. We used an underwater pole-cam, specimen collections, and a boater questionnaire to quantify the extent and composition of biofouling on recreational boats and to evaluate boater behavior at a subset of the Bay’s marinas. Several NIS, already established within the Bay, were recorded from vessel hulls, including the bryozoans Bugula neritina, Membranipora chesapeakensis and Watersipora sp., the ascidians Botrylloides violaceus, Styela clava and Ciona intestinalis, the polychaete Ficopomatus enigmaticus, and the sponge Clathria prolifera. Only 16% of questionnaire respondents had traveled to sites outside the Bay in the previous 12 mo. Frequency of hull painting and cleaning varied substantially, but we did not find strong patterns of biofouling extent associated with hull husbandry or boat usage. The potential for within-Bay and coastwise regional spread of NIS is high, and recreational boats probably interact in close proximity to other vectors (e.g. commercial ships), causing a ratchet effect of vector events; however, there remains a gap in understanding the levels and condition of biofouling on transient boats. Transient vessels from San Francisco Bay and other West Coast sites should be the focus of future studies to evaluate the extent to which organisms are being transferred among bays and how vector management could be applied to prevent NIS transfers and impacts. Close

• 7. [Article] A Dynamic Physical Model for Soil Temperature and Water in Taylor Valley, Antarctica

  We developed a simulation model for terrestrial sites including sensible heat exchange between the atmosphere and ground surface, inter- and intra-layer heat conduction by rock and soil, and shortwave ...

  Citation × Citation Citation Abstract Copy Title: A Dynamic Physical Model for Soil Temperature and Water in Taylor Valley, Antarctica Author: Hunt, H. W., Fountain, Andrew G., Doran, Peter T., Basagic, Hassan J. Year: 2010 We developed a simulation model for terrestrial sites including sensible heat exchange between the atmosphere and ground surface, inter- and intra-layer heat conduction by rock and soil, and shortwave and longwave radiation. Water fluxes included snowmelt, freezing/thawing of soil water, soil capillary flow, and vapour flows among atmosphere, soil, and snow. The model accounted for 96-99% of variation in soil temperature data. No long-term temporal trends in soil temperature were apparent. Soil water vapour concentration in thawed surface soil in summer often was higher than in frozen deeper soils, leading to downward vapour fluxes. Katabatic winds caused a reversal of the usual winter pattern of upward vapour fluxes. The model exhibited a steady state depth distribution of soil water due to vapour flows and in the absence of capillary flows below the top 0.5 cm soil layer. Beginning with a completely saturated soil profile, soil water was lost rapidly, and within a few hundred years approached a steady state characterized by dry soil (,0.5% gravimetric) down to one metre depth and saturated soil below that. In contrast, it took 42,000 years to approach steady state beginning from a completely dry initial condition. Title: A Dynamic Physical Model for Soil Temperature and Water in Taylor Valley, Antarctica Author: Hunt, H. W., Fountain, Andrew G., Doran, Peter T., Basagic, Hassan J. Year: 2010 We developed a simulation model for terrestrial sites including sensible heat exchange between the atmosphere and ground surface, inter- and intra-layer heat conduction by rock and soil, and shortwave and longwave radiation. Water fluxes included snowmelt, freezing/thawing of soil water, soil capillary flow, and vapour flows among atmosphere, soil, and snow. The model accounted for 96-99% of variation in soil temperature data. No long-term temporal trends in soil temperature were apparent. Soil water vapour concentration in thawed surface soil in summer often was higher than in frozen deeper soils, leading to downward vapour fluxes. Katabatic winds caused a reversal of the usual winter pattern of upward vapour fluxes. The model exhibited a steady state depth distribution of soil water due to vapour flows and in the absence of capillary flows below the top 0.5 cm soil layer. Beginning with a completely saturated soil profile, soil water was lost rapidly, and within a few hundred years approached a steady state characterized by dry soil (,0.5% gravimetric) down to one metre depth and saturated soil below that. In contrast, it took 42,000 years to approach steady state beginning from a completely dry initial condition. Close

• 8. [Article] Convergent Surface Water Distributions in U.S. Cities

  Earth's surface is rapidly urbanizing, resulting in dramatic changes in the abundance, distribution and character of surface water features in urban landscapes. However,the scope and consequences of surface ...

  Citation × Citation Citation Abstract Copy Title: Convergent Surface Water Distributions in U.S. Cities Author: Steele, Meredith K., Heffernan, James B., Bettez, Neil D., Cavender-Bares, Jeannine, Groffman, Peter M., Grove, J. Morgan, Hall, Sharon J., Hobbie, Sarah E., Larson, Kelli L., Morse, Jennifer L., Neill, Christopher, Nelson, Kristen C., O'Neil-Dunne, Jarlath, Ogden, Laura A., Pataki, Diane E., Polsky, Colin, Chowdhury, Rinku Roy Year: 2014 Earth's surface is rapidly urbanizing, resulting in dramatic changes in the abundance, distribution and character of surface water features in urban landscapes. However,the scope and consequences of surface water redistribution at broad spatialscales are not well understood. We hypothesized that urbanization would lead to convergent surface water abundance and distribution: in other words, cities will gain or lose water such that they become more similar to each other than are their surrounding natural landscapes. Using a database of more than 1 million water bodies and 1 million km of streams, we compared the surface water of 100 US cities with their surrounding undeveloped land. We evaluated differences in areal ( A) and numeric densities ( N) of water bodies (lakes, wetlands, and so on), the morphological characteristics of water bodies (size), and the density ( D) of surface flow channels (that is, streams and rivers). The variance of urban A, N, and D across the 100 MSAs decreased, by 89, 25, and 71%, respectively, compared to undeveloped land. These data show that many cities are surface water poor relative to undeveloped land; however, in drier landscapesurbanization increases the occurrence of surface water. This convergence pattern strengthened with development intensity, such that high intensityurban development had an areal water body density 98% less than undeveloped lands. Urbanization appears to drive the convergence of hydrological features across the US, such that surface water distributions of cities are more similar to each other than to their surrounding landscapes. Title: Convergent Surface Water Distributions in U.S. Cities Author: Steele, Meredith K., Heffernan, James B., Bettez, Neil D., Cavender-Bares, Jeannine, Groffman, Peter M., Grove, J. Morgan, Hall, Sharon J., Hobbie, Sarah E., Larson, Kelli L., Morse, Jennifer L., Neill, Christopher, Nelson, Kristen C., O'Neil-Dunne, Jarlath, Ogden, Laura A., Pataki, Diane E., Polsky, Colin, Chowdhury, Rinku Roy Year: 2014 Earth's surface is rapidly urbanizing, resulting in dramatic changes in the abundance, distribution and character of surface water features in urban landscapes. However,the scope and consequences of surface water redistribution at broad spatialscales are not well understood. We hypothesized that urbanization would lead to convergent surface water abundance and distribution: in other words, cities will gain or lose water such that they become more similar to each other than are their surrounding natural landscapes. Using a database of more than 1 million water bodies and 1 million km of streams, we compared the surface water of 100 US cities with their surrounding undeveloped land. We evaluated differences in areal ( A) and numeric densities ( N) of water bodies (lakes, wetlands, and so on), the morphological characteristics of water bodies (size), and the density ( D) of surface flow channels (that is, streams and rivers). The variance of urban A, N, and D across the 100 MSAs decreased, by 89, 25, and 71%, respectively, compared to undeveloped land. These data show that many cities are surface water poor relative to undeveloped land; however, in drier landscapesurbanization increases the occurrence of surface water. This convergence pattern strengthened with development intensity, such that high intensityurban development had an areal water body density 98% less than undeveloped lands. Urbanization appears to drive the convergence of hydrological features across the US, such that surface water distributions of cities are more similar to each other than to their surrounding landscapes. Close

• 9. [Article] Introduction to Common Native and Potential Invasive Freshwater Plants in Alaska

  Freshwater aquatic plants are found in most lakes and rivers in Alaska. They range from tiny floating plants that can form mats on a lake surface to emergent plants growing two meters above the water. ...

  Citation × Citation Citation Abstract Copy Title: Introduction to Common Native and Potential Invasive Freshwater Plants in Alaska Author: Morgan, Vanessa Howard, Sytsma, Mark Year: 2009 Freshwater aquatic plants are found in most lakes and rivers in Alaska. They range from tiny floating plants that can form mats on a lake surface to emergent plants growing two meters above the water. Aquatic plants are an important part of freshwater environments. They provide food and shelter for a wide variety of insects, fish, mammals, and birds; stabilize shorelines; and form an important link in nutrient cycles. To date, Alaska has had relatively few problems with invasive, non-native aquatic plants. However, invasive aquatic plants pose an increasing threat to human safety and the integrity of native aquatic communities. Actions taken to detect and prevent the introduction and spread of invasive aquatic plants in Alaska are necessary in order to avoid the environmental and economic harm invasive plants have caused in other parts of the United States. The primary purposes of this manual are to aide the identification of freshwater aquatic plants that are likely to be seen in Alaska’s lakes, streams, bogs and marshes; and to help prevent the spread of harmful invasive plants. This manual was written for people who do not necessarily have a scientific background, but it also should be useful for professionals who need assistance in identifying common aquatic species. Title: Introduction to Common Native and Potential Invasive Freshwater Plants in Alaska Author: Morgan, Vanessa Howard, Sytsma, Mark Year: 2009 Freshwater aquatic plants are found in most lakes and rivers in Alaska. They range from tiny floating plants that can form mats on a lake surface to emergent plants growing two meters above the water. Aquatic plants are an important part of freshwater environments. They provide food and shelter for a wide variety of insects, fish, mammals, and birds; stabilize shorelines; and form an important link in nutrient cycles. To date, Alaska has had relatively few problems with invasive, non-native aquatic plants. However, invasive aquatic plants pose an increasing threat to human safety and the integrity of native aquatic communities. Actions taken to detect and prevent the introduction and spread of invasive aquatic plants in Alaska are necessary in order to avoid the environmental and economic harm invasive plants have caused in other parts of the United States. The primary purposes of this manual are to aide the identification of freshwater aquatic plants that are likely to be seen in Alaska’s lakes, streams, bogs and marshes; and to help prevent the spread of harmful invasive plants. This manual was written for people who do not necessarily have a scientific background, but it also should be useful for professionals who need assistance in identifying common aquatic species. Close

• 10. [Article] Surface Energy Balance and Melt Thresholds over 11 years at Taylor Glacier, Antarctica

  In the McMurdo Dry Valleys, Victoria Land, Antarctica, melting of glacial ice is the primary source of water to streams, lakes, and associated ecosystems. To understand geochemical fluxes and ecological ...

  Citation × Citation Citation Abstract Copy Title: Surface Energy Balance and Melt Thresholds over 11 years at Taylor Glacier, Antarctica Author: Hoffman, Matthew James, Fountain, Andrew G., Liston, Glen E. Year: 2008 In the McMurdo Dry Valleys, Victoria Land, Antarctica, melting of glacial ice is the primary source of water to streams, lakes, and associated ecosystems. To understand geochemical fluxes and ecological responses to past and future climates requires a physically based energy balance model. We applied a one-dimensional model to one site on Taylor Glacier using 11 years of daily meteorological data and seasonal ablation measurements. Inclusion of transmission of solar radiation into the ice was necessary to accurately model summer ablation and ice temperatures. Results showed good correspondence between calculated and measured ablation and ice temperatures over the 11 years. Ablation (18 cm a??) was dominated by sublimation with very few occurrences of melt (42 days during 11 years). Results also indicated that above freezing air temperatures did not necessarily result in melt and, in turn, melt occurred during subfreezing air temperatures under some conditions. For air temperatures near freezing, low wind speed was critically important for melt initiation. According to the model, subsurface melt, away from rocks and sediment in the ice, occurred three times more frequently than surface melt; occurs no deeper than 50 cm below the glacier surface; and was small, never exceeding 8% by mass. The magnitude of subsurface melting and the energy balance indicate that Taylor Title: Surface Energy Balance and Melt Thresholds over 11 years at Taylor Glacier, Antarctica Author: Hoffman, Matthew James, Fountain, Andrew G., Liston, Glen E. Year: 2008 In the McMurdo Dry Valleys, Victoria Land, Antarctica, melting of glacial ice is the primary source of water to streams, lakes, and associated ecosystems. To understand geochemical fluxes and ecological responses to past and future climates requires a physically based energy balance model. We applied a one-dimensional model to one site on Taylor Glacier using 11 years of daily meteorological data and seasonal ablation measurements. Inclusion of transmission of solar radiation into the ice was necessary to accurately model summer ablation and ice temperatures. Results showed good correspondence between calculated and measured ablation and ice temperatures over the 11 years. Ablation (18 cm a??) was dominated by sublimation with very few occurrences of melt (42 days during 11 years). Results also indicated that above freezing air temperatures did not necessarily result in melt and, in turn, melt occurred during subfreezing air temperatures under some conditions. For air temperatures near freezing, low wind speed was critically important for melt initiation. According to the model, subsurface melt, away from rocks and sediment in the ice, occurred three times more frequently than surface melt; occurs no deeper than 50 cm below the glacier surface; and was small, never exceeding 8% by mass. The magnitude of subsurface melting and the energy balance indicate that Taylor Close