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• 1. [Article] Carbon Dynamics in the Future Forest: the Importance of Long-Term Successional Legacy and Climate–Fire Interactions

  Understanding how climate change may influence forest carbon (C) budgets requires knowledge of forest growth relationships with regional climate, long-term forest succession, and past and future disturbances, ...

  Citation × Citation Citation Abstract Copy Title: Carbon Dynamics in the Future Forest: the Importance of Long-Term Successional Legacy and Climate–Fire Interactions Author: Loudermilk, E. Louise, Scheller, Robert M., Weisberg, Peter J., Yang, Jian, Dilts, Thomas E., Karam, Sarah L., Skinner, Carl Year: 2013 Understanding how climate change may influence forest carbon (C) budgets requires knowledge of forest growth relationships with regional climate, long-term forest succession, and past and future disturbances, such as wildfires and timber harvesting events. We used a landscape-scale model of forest succession, wildfire, and C dynamics (LANDIS-II) to evaluate the effects of a changing climate (A2 and B1 IPCC emissions; Geophysical Fluid Dynamics Laboratory General Circulation Models) on total forest C, tree species composition, and wildfire dynamics in the Lake Tahoe Basin, California, and Nevada. The independent effects of temperature and precipitation were assessed within and among climate models. Results highlight the importance of modeling forest succession and stand development processes at the landscape scale for understanding the C cycle. Due primarily to landscape legacy effects of historic logging of the Comstock Era in the late 1880s, C sequestration may continue throughout the current century, and the forest will remain a C sink (Net Ecosystem Carbon Balance > 0), regardless of climate regime. Climate change caused increases in temperatures limited simulated C sequestration potential because of augmented fire activity and reduced establishment ability of subalpine and upper montane trees. Higher temperatures influenced forest response more than reduced precipitation. As the forest reached its potential steady state, the forest could become C neutral or a C source, and climate change could accelerate this transition. The future of forest ecosystem C cycling in many forested systems worldwide may depend more on major disturbances and landscape legacies related to land use than on projected climate change alone. Title: Carbon Dynamics in the Future Forest: the Importance of Long-Term Successional Legacy and Climate–Fire Interactions Author: Loudermilk, E. Louise, Scheller, Robert M., Weisberg, Peter J., Yang, Jian, Dilts, Thomas E., Karam, Sarah L., Skinner, Carl Year: 2013 Understanding how climate change may influence forest carbon (C) budgets requires knowledge of forest growth relationships with regional climate, long-term forest succession, and past and future disturbances, such as wildfires and timber harvesting events. We used a landscape-scale model of forest succession, wildfire, and C dynamics (LANDIS-II) to evaluate the effects of a changing climate (A2 and B1 IPCC emissions; Geophysical Fluid Dynamics Laboratory General Circulation Models) on total forest C, tree species composition, and wildfire dynamics in the Lake Tahoe Basin, California, and Nevada. The independent effects of temperature and precipitation were assessed within and among climate models. Results highlight the importance of modeling forest succession and stand development processes at the landscape scale for understanding the C cycle. Due primarily to landscape legacy effects of historic logging of the Comstock Era in the late 1880s, C sequestration may continue throughout the current century, and the forest will remain a C sink (Net Ecosystem Carbon Balance > 0), regardless of climate regime. Climate change caused increases in temperatures limited simulated C sequestration potential because of augmented fire activity and reduced establishment ability of subalpine and upper montane trees. Higher temperatures influenced forest response more than reduced precipitation. As the forest reached its potential steady state, the forest could become C neutral or a C source, and climate change could accelerate this transition. The future of forest ecosystem C cycling in many forested systems worldwide may depend more on major disturbances and landscape legacies related to land use than on projected climate change alone. Close

• 2. [Article] Effectiveness of Fuel Treatments for Mitigating Wildfire Risk and Sequestering Forest Carbon: A Case Study in the Lake Tahoe Basin

  Fuel-reduction treatments are used extensively to reduce wildfire risk and restore forest diversity and function. In the near future, increasing regulation of carbon (C) emissions may force forest managers ...

  Citation × Citation Citation Abstract Copy Title: Effectiveness of Fuel Treatments for Mitigating Wildfire Risk and Sequestering Forest Carbon: A Case Study in the Lake Tahoe Basin Author: Loudermilk, E. Louise, Stanton, Alison, Scheller, Robert M., Dilts, Thomas E., Weisberg, Peter J., Skinner, Carl, Yang, Jian Year: 2014 Fuel-reduction treatments are used extensively to reduce wildfire risk and restore forest diversity and function. In the near future, increasing regulation of carbon (C) emissions may force forest managers to balance the use of fuel treatments for reducing wildfire risk against an alternative goal of C sequestration. The objective of this study was to evaluate how long-term fuel treatments mitigate wildfires and affect forest C. For the Lake Tahoe Basin in the central Sierra Nevada, USA, fuel treatment efficiency was explored with a landscape-scale simulation model, LANDIS-II, using five fuel treatment scenarios and two (contemporary and potential future) fire regimes. Treatment scenarios included applying a combination of light (hand) and moderate (mechanical) forest thinning continuously through time and transitioning from these prescriptions to a more mid-seral thinning prescription, both on a 15 and 30 year rotation interval. In the last scenario, fuel treatments were isolated to around the lake shore (nearby urban settlement) to simulate a low investment alternative were future resources may be limited. Results indicated that the forest will remain a C sink regardless of treatment or fire regime simulated, due to the landscape legacy of historic logging. Achievement of a net C gain required decades with intensive treatment and depended on wildfire activity: Fuel treatments were more effective in a more active fire environment, where the interface between wildfires and treatment areas increased and caused net C gain earlier than as compared to our scenarios with less wildfire activity. Fuel treatments were most effective when continuously applied and strategically placed in high ignition areas. Treatment type and re-application interval were less influential at the landscape scale, but had notable effects on species dynamics within management units. Treatments created more diverse forest conditions by shifting dominance patterns to a more mixed conifer system, with a higher proportion of fire-tolerant species. We demonstrated that a small amount of wildfire on the landscape resulted in significant changes in the C pool, and that strategically placed fuel treatments substantially reduced wildfire risk, increased fire resiliency of the forest, and is beneficial for long-term C management. Implications for landscape management included consideration for prioritization of treatment areas and creating ideal re-entry schedules that meet logistic, safety, and conservation goals. In forests with a concentrated wildland urban interface, fuel treatments may be vital for ensuring human welfare and enhancing forest integrity in a fire-prone future. Title: Effectiveness of Fuel Treatments for Mitigating Wildfire Risk and Sequestering Forest Carbon: A Case Study in the Lake Tahoe Basin Author: Loudermilk, E. Louise, Stanton, Alison, Scheller, Robert M., Dilts, Thomas E., Weisberg, Peter J., Skinner, Carl, Yang, Jian Year: 2014 Fuel-reduction treatments are used extensively to reduce wildfire risk and restore forest diversity and function. In the near future, increasing regulation of carbon (C) emissions may force forest managers to balance the use of fuel treatments for reducing wildfire risk against an alternative goal of C sequestration. The objective of this study was to evaluate how long-term fuel treatments mitigate wildfires and affect forest C. For the Lake Tahoe Basin in the central Sierra Nevada, USA, fuel treatment efficiency was explored with a landscape-scale simulation model, LANDIS-II, using five fuel treatment scenarios and two (contemporary and potential future) fire regimes. Treatment scenarios included applying a combination of light (hand) and moderate (mechanical) forest thinning continuously through time and transitioning from these prescriptions to a more mid-seral thinning prescription, both on a 15 and 30 year rotation interval. In the last scenario, fuel treatments were isolated to around the lake shore (nearby urban settlement) to simulate a low investment alternative were future resources may be limited. Results indicated that the forest will remain a C sink regardless of treatment or fire regime simulated, due to the landscape legacy of historic logging. Achievement of a net C gain required decades with intensive treatment and depended on wildfire activity: Fuel treatments were more effective in a more active fire environment, where the interface between wildfires and treatment areas increased and caused net C gain earlier than as compared to our scenarios with less wildfire activity. Fuel treatments were most effective when continuously applied and strategically placed in high ignition areas. Treatment type and re-application interval were less influential at the landscape scale, but had notable effects on species dynamics within management units. Treatments created more diverse forest conditions by shifting dominance patterns to a more mixed conifer system, with a higher proportion of fire-tolerant species. We demonstrated that a small amount of wildfire on the landscape resulted in significant changes in the C pool, and that strategically placed fuel treatments substantially reduced wildfire risk, increased fire resiliency of the forest, and is beneficial for long-term C management. Implications for landscape management included consideration for prioritization of treatment areas and creating ideal re-entry schedules that meet logistic, safety, and conservation goals. In forests with a concentrated wildland urban interface, fuel treatments may be vital for ensuring human welfare and enhancing forest integrity in a fire-prone future. Close

• 3. [Article] Climate and Bark Beetle Effects on Forest Productivity — Linking Dendroecology with Forest Landscape Modeling

  In forested systems throughout the world, climate influences tree growth and aboveground net primary productivity (ANPP). The effects of extreme climate events (i.e., drought) on ANPP can be compounded ...

  Citation × Citation Citation Abstract Copy Title: Climate and Bark Beetle Effects on Forest Productivity — Linking Dendroecology with Forest Landscape Modeling Author: Kretchun, Alec M., Loudermilk, E. Louise, Scheller, Robert M., Hurteau, Matthew, Belmecher, Soumaya Year: 2016 In forested systems throughout the world, climate influences tree growth and aboveground net primary productivity (ANPP). The effects of extreme climate events (i.e., drought) on ANPP can be compounded by biotic factors (e.g., insect outbreaks). Understanding the contribution of each of these influences on growth requires information at multiple spatial scales and is essential for understanding regional forest response to changing climate. The mixed conifer forests of the Lake Tahoe Basin, California and Nevada, provide an opportunity to analyze biotic and abiotic influences on ANPP. Our objective was to evaluate the influence of moisture stress (climatic water deficit, CWD) and bark beetles on basin-wide ANPP from 1987 to 2006, estimated through tree core increments and a landscape simulation model (LANDIS-II). Tree ring data revealed that ANPP increased throughout this period and had a nonlinear relationship to water demand. Simulation model results showed that despite increased complexity, simulations that include moderate moisture sensitivity and bark beetle outbreaks most closely approximated the field-derived ANPP∼CWD relationship. Although bark beetle outbreaks and episodic drought-induced mortality events are often correlated, decoupling them within a simulation model offers insight into assessing model performance, as well as examining how each contributes to total declines in productivity. Title: Climate and Bark Beetle Effects on Forest Productivity — Linking Dendroecology with Forest Landscape Modeling Author: Kretchun, Alec M., Loudermilk, E. Louise, Scheller, Robert M., Hurteau, Matthew, Belmecher, Soumaya Year: 2016 In forested systems throughout the world, climate influences tree growth and aboveground net primary productivity (ANPP). The effects of extreme climate events (i.e., drought) on ANPP can be compounded by biotic factors (e.g., insect outbreaks). Understanding the contribution of each of these influences on growth requires information at multiple spatial scales and is essential for understanding regional forest response to changing climate. The mixed conifer forests of the Lake Tahoe Basin, California and Nevada, provide an opportunity to analyze biotic and abiotic influences on ANPP. Our objective was to evaluate the influence of moisture stress (climatic water deficit, CWD) and bark beetles on basin-wide ANPP from 1987 to 2006, estimated through tree core increments and a landscape simulation model (LANDIS-II). Tree ring data revealed that ANPP increased throughout this period and had a nonlinear relationship to water demand. Simulation model results showed that despite increased complexity, simulations that include moderate moisture sensitivity and bark beetle outbreaks most closely approximated the field-derived ANPP∼CWD relationship. Although bark beetle outbreaks and episodic drought-induced mortality events are often correlated, decoupling them within a simulation model offers insight into assessing model performance, as well as examining how each contributes to total declines in productivity. Close

• 4. [Article] Interactions Among Fuel Management, Species Composition, Bark Beetles, and Climate Change and the Potential Effects on Forests of the Lake Tahoe Basin

  Climate-driven increases in wildfires, drought conditions, and insect outbreaks are critical threats to forest carbon stores. In particular, bark beetles are important disturbance agents although their ...

  Citation × Citation Citation Abstract Copy Title: Interactions Among Fuel Management, Species Composition, Bark Beetles, and Climate Change and the Potential Effects on Forests of the Lake Tahoe Basin Author: Scheller, Robert M., Kretchun, Alec M., Loudermilk, E. Louise, Hurteau, Matthew D., Weisberg, Peter J., Skinner, Carl Year: 2017 Climate-driven increases in wildfires, drought conditions, and insect outbreaks are critical threats to forest carbon stores. In particular, bark beetles are important disturbance agents although their long-term interactions with future climate change are poorly understood. Droughts and the associated moisture deficit contribute to the onset of bark beetle outbreaks although outbreak extent and severity is dependent upon the density of host trees, wildfire, and forest management. Our objective was to estimate the effects of climate change and bark beetle outbreaks on ecosystem carbon dynamics over the next century in a western US forest. Specifically, we hypothesized that (a) bark beetle outbreaks under climate change would reduce net ecosystem carbon balance (NECB) and increase uncertainty and (b) these effects could be ameliorated by fuels management. We also examined the specific tree species dynamics—competition and release—that determined NECB response to bark beetle outbreaks. Our study area was the Lake Tahoe Basin (LTB), CA and NV, USA, an area of diverse forest types encompassing steep elevation and climatic gradients and representative of mixed-conifer forests throughout the western United States. We simulated climate change, bark beetles, wildfire, and fuels management using a landscape-scale stochastic model of disturbance and succession. We simulated the period 2010–2100 using downscaled climate projections. Recurring droughts generated conditions conducive to large-scale outbreaks; the resulting large and sustained outbreaks significantly increased the probability of LTB forests becoming C sources over decadal time scales, with slower-than-anticipated landscape-scale recovery. Tree species composition was substantially altered with a reduction in functional redundancy and productivity. Results indicate heightened uncertainty due to the synergistic influences of climate change and interacting disturbances. Our results further indicate that current fuel management practices will not be effective at reducing landscape-scale outbreak mortality. Our results provide critical insights into the interaction of drivers (bark beetles, wildfire, fuel management) that increase the risk of C loss and shifting community composition if bark beetle outbreaks become more frequent. Title: Interactions Among Fuel Management, Species Composition, Bark Beetles, and Climate Change and the Potential Effects on Forests of the Lake Tahoe Basin Author: Scheller, Robert M., Kretchun, Alec M., Loudermilk, E. Louise, Hurteau, Matthew D., Weisberg, Peter J., Skinner, Carl Year: 2017 Climate-driven increases in wildfires, drought conditions, and insect outbreaks are critical threats to forest carbon stores. In particular, bark beetles are important disturbance agents although their long-term interactions with future climate change are poorly understood. Droughts and the associated moisture deficit contribute to the onset of bark beetle outbreaks although outbreak extent and severity is dependent upon the density of host trees, wildfire, and forest management. Our objective was to estimate the effects of climate change and bark beetle outbreaks on ecosystem carbon dynamics over the next century in a western US forest. Specifically, we hypothesized that (a) bark beetle outbreaks under climate change would reduce net ecosystem carbon balance (NECB) and increase uncertainty and (b) these effects could be ameliorated by fuels management. We also examined the specific tree species dynamics—competition and release—that determined NECB response to bark beetle outbreaks. Our study area was the Lake Tahoe Basin (LTB), CA and NV, USA, an area of diverse forest types encompassing steep elevation and climatic gradients and representative of mixed-conifer forests throughout the western United States. We simulated climate change, bark beetles, wildfire, and fuels management using a landscape-scale stochastic model of disturbance and succession. We simulated the period 2010–2100 using downscaled climate projections. Recurring droughts generated conditions conducive to large-scale outbreaks; the resulting large and sustained outbreaks significantly increased the probability of LTB forests becoming C sources over decadal time scales, with slower-than-anticipated landscape-scale recovery. Tree species composition was substantially altered with a reduction in functional redundancy and productivity. Results indicate heightened uncertainty due to the synergistic influences of climate change and interacting disturbances. Our results further indicate that current fuel management practices will not be effective at reducing landscape-scale outbreak mortality. Our results provide critical insights into the interaction of drivers (bark beetles, wildfire, fuel management) that increase the risk of C loss and shifting community composition if bark beetle outbreaks become more frequent. Close

• 5. [Article] Bending the Carbon Curve: Fire Management for Carbon Resilience under Climate Change

  Context: Forest landscapes are increasingly managed for fire resilience, particularly in the western US which has recently experienced drought and widespread, high-severity wildfires. Fuel reduction treatments ...

  Citation × Citation Citation Abstract Copy Title: Bending the Carbon Curve: Fire Management for Carbon Resilience under Climate Change Author: Loudermilk, E. Louise, Scheller, Robert M., Weisberg, Peter J., Kretchun, Alec M. Year: 2017 Context: Forest landscapes are increasingly managed for fire resilience, particularly in the western US which has recently experienced drought and widespread, high-severity wildfires. Fuel reduction treatments have been effective where fires coincide with treated areas. Fuel treatments also have the potential to reduce drought-mortality if tree density is uncharacteristically high, and to increase long-term carbon storage by reducing high-severity fire probability. Objective: Assess whether fuel treatments reduce fire intensity and spread and increase carbon storage under climate change. Methods: We used a simulation modeling approach that couples a landscape model of forest disturbance and succession with an ecosystem model of carbon dynamics (Century), to quantify the interacting effects of climate change, fuel treatments and wildfire for carbon storage potential in a mixed-conifer forest in the western USA. Results: Our results suggest that fuel treatments have the potential to ‘bend the C curve’, maintaining carbon resilience despite climate change and climate-related changes to the fire regime. Simulated fuel treatments resulted in reduced fire spread and severity. There was partial compensation of C lost during fuel treatments with increased growth of residual stock due to greater available soil water, as well as a shift in species composition to more drought- and fire-tolerant Pinus jeffreyi at the expense of shade-tolerant, fire-susceptible Abies concolor. Conclusions: Forest resilience to global change can be achieved through management that reduces drought stress and supports the establishment and dominance of tree species that are more fire- and drought-resistant, however, achieving a net C gain from fuel treatments may take decades. Title: Bending the Carbon Curve: Fire Management for Carbon Resilience under Climate Change Author: Loudermilk, E. Louise, Scheller, Robert M., Weisberg, Peter J., Kretchun, Alec M. Year: 2017 Context: Forest landscapes are increasingly managed for fire resilience, particularly in the western US which has recently experienced drought and widespread, high-severity wildfires. Fuel reduction treatments have been effective where fires coincide with treated areas. Fuel treatments also have the potential to reduce drought-mortality if tree density is uncharacteristically high, and to increase long-term carbon storage by reducing high-severity fire probability. Objective: Assess whether fuel treatments reduce fire intensity and spread and increase carbon storage under climate change. Methods: We used a simulation modeling approach that couples a landscape model of forest disturbance and succession with an ecosystem model of carbon dynamics (Century), to quantify the interacting effects of climate change, fuel treatments and wildfire for carbon storage potential in a mixed-conifer forest in the western USA. Results: Our results suggest that fuel treatments have the potential to ‘bend the C curve’, maintaining carbon resilience despite climate change and climate-related changes to the fire regime. Simulated fuel treatments resulted in reduced fire spread and severity. There was partial compensation of C lost during fuel treatments with increased growth of residual stock due to greater available soil water, as well as a shift in species composition to more drought- and fire-tolerant Pinus jeffreyi at the expense of shade-tolerant, fire-susceptible Abies concolor. Conclusions: Forest resilience to global change can be achieved through management that reduces drought stress and supports the establishment and dominance of tree species that are more fire- and drought-resistant, however, achieving a net C gain from fuel treatments may take decades. Close

• 6. [Article] Restoring Surface Fire Stabilizes Forest Carbon Under Extreme Fire Weather in the Sierra Nevada

  Climate change in the western United States has increased the frequency of extreme fire weather events and is projected to increase the area burned by wildfire in the coming decades. This changing fire regime, ...

  Citation × Citation Citation Abstract Copy Title: Restoring Surface Fire Stabilizes Forest Carbon Under Extreme Fire Weather in the Sierra Nevada Author: Krofcheck, Daniel J., Hurteau, Matthew D., Scheller, Robert M., Loudermilk, E. Louise Year: 2017 Climate change in the western United States has increased the frequency of extreme fire weather events and is projected to increase the area burned by wildfire in the coming decades. This changing fire regime, coupled with increased high-severity fire risk from a legacy of fire exclusion, could destabilize forest carbon (C), decrease net ecosystem exchange (NEE), and consequently reduce the ability of forests to regulate climate through C sequestration. While management options for minimizing the risk of high-severity fire exist, little is known about the longer-term carbon consequences of these actions in the context of continued extreme fire weather events. Our goal was to compare the impacts of extreme wildfire events on carbon stocks and fluxes in a watershed in the Sierra National Forest. We ran simulations to model wildfire under contemporary and extreme fire weather conditions, and test how three management scenarios (no-management, thin-only, thin and maintenance burning) influence fire severity, forest C stocks and fluxes, and wildfire C emissions. We found that the effects of treatment on wildfire under contemporary fire weather were minimal, and management conferred neither significant reduction in fire severity nor increases in C stocks. However, under extreme fire weather, the thin and maintenance burning scenario decreased mean fire severity by 25%, showed significantly greater C stability, and unlike the no-management and thin-only management options, the thin and maintenance burning scenario showed no decrease in NEE relative to the contemporary fire weather scenarios. Further, under extreme fire weather conditions, wildfire C emissions were lowest in the thin and maintenance burning scenario, (reduction of 13.7 Mg C/ha over the simulation period) even when taking into account the C costs associated with prescribed burning. Including prescribed burning in thinning operations may be critical to maintaining C stocks and reducing C emissions in the future where extreme fire weather events are more frequent. Title: Restoring Surface Fire Stabilizes Forest Carbon Under Extreme Fire Weather in the Sierra Nevada Author: Krofcheck, Daniel J., Hurteau, Matthew D., Scheller, Robert M., Loudermilk, E. Louise Year: 2017 Climate change in the western United States has increased the frequency of extreme fire weather events and is projected to increase the area burned by wildfire in the coming decades. This changing fire regime, coupled with increased high-severity fire risk from a legacy of fire exclusion, could destabilize forest carbon (C), decrease net ecosystem exchange (NEE), and consequently reduce the ability of forests to regulate climate through C sequestration. While management options for minimizing the risk of high-severity fire exist, little is known about the longer-term carbon consequences of these actions in the context of continued extreme fire weather events. Our goal was to compare the impacts of extreme wildfire events on carbon stocks and fluxes in a watershed in the Sierra National Forest. We ran simulations to model wildfire under contemporary and extreme fire weather conditions, and test how three management scenarios (no-management, thin-only, thin and maintenance burning) influence fire severity, forest C stocks and fluxes, and wildfire C emissions. We found that the effects of treatment on wildfire under contemporary fire weather were minimal, and management conferred neither significant reduction in fire severity nor increases in C stocks. However, under extreme fire weather, the thin and maintenance burning scenario decreased mean fire severity by 25%, showed significantly greater C stability, and unlike the no-management and thin-only management options, the thin and maintenance burning scenario showed no decrease in NEE relative to the contemporary fire weather scenarios. Further, under extreme fire weather conditions, wildfire C emissions were lowest in the thin and maintenance burning scenario, (reduction of 13.7 Mg C/ha over the simulation period) even when taking into account the C costs associated with prescribed burning. Including prescribed burning in thinning operations may be critical to maintaining C stocks and reducing C emissions in the future where extreme fire weather events are more frequent. Close