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  • This is the publisher’s final pdf. The published article is copyrighted by the author(s) and published by the Public Library of Science. The published article can be found at: http://www.plosone.org/.
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  • This is the authors’ post-peer review version of the final article. The final published version is copyrighted by Springer and can be found at: http://www.springer.com/life+sciences/evolutionary+%26+de...
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