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• 1. [Article] Density, ages, and growth rates in old-growth and young-growth forests in coastal Oregon

  We studied the ages and diameter growth rates of trees in former Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) old-growth stands on 10 sites and compared them with young-growth stands (50-70 years ...

  Citation × Citation Citation Abstract Copy Title: Density, ages, and growth rates in old-growth and young-growth forests in coastal Oregon Author: Huffman, David W. (William), 1962-, Tappeiner, John C, Spies, Thomas A, Marshall, David, Bailey, John D We studied the ages and diameter growth rates of trees in former Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) old-growth stands on 10 sites and compared them with young-growth stands (50-70 years old, regenerated after timber harvest) in the Coast Range of western Oregon. The diameters and diameter growth rates for the first 100 years of trees in the old-growth stands were significantly greater than those in the young-growth stands. Growth rates in the old stands were comparable with those from long-term studies of young stands in which density is about 100-120 trees/ha; often young-growth stand density is well over 500 trees/ha. Ages of large trees in the old stands ranged from 100 to 420 years; ages in young stands varied by only about 5 to 10 years. Apparently, regeneration of old-growth stands on these sites occurred over a prolonged period, and trees grew at low density with little self-thinning; in contrast, after timber harvest, young stands may develop with high density of trees with similar ages and considerable self-thinning. The results suggest that thinning may be needed in dense young stands where the management objective is to speed development of old-growth characteristics. Title: Density, ages, and growth rates in old-growth and young-growth forests in coastal Oregon Author: Huffman, David W. (William), 1962-, Tappeiner, John C, Spies, Thomas A, Marshall, David, Bailey, John D We studied the ages and diameter growth rates of trees in former Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) old-growth stands on 10 sites and compared them with young-growth stands (50-70 years old, regenerated after timber harvest) in the Coast Range of western Oregon. The diameters and diameter growth rates for the first 100 years of trees in the old-growth stands were significantly greater than those in the young-growth stands. Growth rates in the old stands were comparable with those from long-term studies of young stands in which density is about 100-120 trees/ha; often young-growth stand density is well over 500 trees/ha. Ages of large trees in the old stands ranged from 100 to 420 years; ages in young stands varied by only about 5 to 10 years. Apparently, regeneration of old-growth stands on these sites occurred over a prolonged period, and trees grew at low density with little self-thinning; in contrast, after timber harvest, young stands may develop with high density of trees with similar ages and considerable self-thinning. The results suggest that thinning may be needed in dense young stands where the management objective is to speed development of old-growth characteristics. Close

• 2. [Article] Thinning : an important timber management tool

  Published August 1983. Facts and recommendations in this publication may no longer be valid. Please look for up-to-date information in the OSU Extension Catalog: http://extension.oregonstate.edu/catalo...

  Citation × Citation Citation Abstract Copy Title: Thinning : an important timber management tool Author: Elwood, Norman E. (Norman Eugene), Emmingham, William H., Oregon State University. Extension Service Published August 1983. Facts and recommendations in this publication may no longer be valid. Please look for up-to-date information in the OSU Extension Catalog: http://extension.oregonstate.edu/catalog Title: Thinning : an important timber management tool Author: Elwood, Norman E. (Norman Eugene), Emmingham, William H., Oregon State University. Extension Service Published August 1983. Facts and recommendations in this publication may no longer be valid. Please look for up-to-date information in the OSU Extension Catalog: http://extension.oregonstate.edu/catalog Close

• 3. [Article] Basic principles of forest fuel reduction treatments

  Successful fire exclusion in the 20th century has created severe fire problems across the West. Not every forest is at risk of uncharacteristically severe wildfire, but drier forests are in need of active ...

  Citation × Citation Citation Abstract Copy Title: Basic principles of forest fuel reduction treatments Author: Agee, James K., Skinner, Carl N. Successful fire exclusion in the 20th century has created severe fire problems across the West. Not every forest is at risk of uncharacteristically severe wildfire, but drier forests are in need of active management to mitigate fire hazard. We summarize a set of simple principles important to address in fuel reduction treatments: reduction of surface fuels, increasing the height to live crown, decreasing crown density, and retaining large trees of fire-resistant species. Thinning and prescribed fire can be useful tools to achieve these objectives. Low thinning will be more effective than crown or selection thinning, and management of surface fuels will increase the likelihood that the stand will survive a wildfire. Five empirical examples of such treatment are discussed: Hayfork fires, California, 1987; Tyee fire,Washington, 1994; Megram fire, California, 1999; Hayman fire, Colorado, 2002; and the Cone fire, California, 2002. Applying treatments at an appropriate landscape scale will be critical to the success of fuel reduction treatments in reducing wildfire losses in Western forests. Title: Basic principles of forest fuel reduction treatments Author: Agee, James K., Skinner, Carl N. Successful fire exclusion in the 20th century has created severe fire problems across the West. Not every forest is at risk of uncharacteristically severe wildfire, but drier forests are in need of active management to mitigate fire hazard. We summarize a set of simple principles important to address in fuel reduction treatments: reduction of surface fuels, increasing the height to live crown, decreasing crown density, and retaining large trees of fire-resistant species. Thinning and prescribed fire can be useful tools to achieve these objectives. Low thinning will be more effective than crown or selection thinning, and management of surface fuels will increase the likelihood that the stand will survive a wildfire. Five empirical examples of such treatment are discussed: Hayfork fires, California, 1987; Tyee fire,Washington, 1994; Megram fire, California, 1999; Hayman fire, Colorado, 2002; and the Cone fire, California, 2002. Applying treatments at an appropriate landscape scale will be critical to the success of fuel reduction treatments in reducing wildfire losses in Western forests. Close

• 4. [Article] Silvicultural Treatment Impacts on Understory Trees and 20-Year Understory Vegetation Dynamics in Mature Douglas-Fir Forests

  Policy regarding the management of public forests has undergone a drastic shift over the past couple of decades due to the loss old-growth forests in the Pacific Northwest (PNW) of the United States. For ...

  Citation × Citation Citation Abstract Copy Title: Silvicultural Treatment Impacts on Understory Trees and 20-Year Understory Vegetation Dynamics in Mature Douglas-Fir Forests Author: Priebe, Jim E. Policy regarding the management of public forests has undergone a drastic shift over the past couple of decades due to the loss old-growth forests in the Pacific Northwest (PNW) of the United States. For much of the 20th century, forest management on public lands emphasized timber production through the use of even-aged management practices. There has been increasing recognition, however, that traditional even-aged management approaches are unable to support species that rely on the complex, heterogeneous structures provided by old-growth forests. In response, public forest managers have redirected their focus to developing more ecologically sustainable forests capable of meeting a broad array of objectives including an increasing emphasis on the development of late-seral and old-growth characteristics. Thinning has been identified as a promising method for promoting late-seral characteristics in managed stands. Recent long-term studies have shown that thinning stands does indeed accelerate the development of at least some late-seral structure characteristics, particularly when varying levels of thinning intensity and non-uniform retention patterns are incorporated into silvicultural prescriptions. Likewise, thinning has also shown some ability to increase the abundance of late-seral associated plant species in the understory. The impacts of thinning on vegetation dynamics are complicated by external factors such as natural disturbance events and the influence of pre-treatment vegetation on post-treatment communities. Within the context of managing for late-seral attributes, thinning is used to imitate natural disturbance processes. However, this does not preclude the occurrence of natural disturbances, which may either disrupt or compound treatment effects. Initial site conditions create another potential complication for the development of late-seral attributes by limiting the potential for change in understory communities. While some studies have shown that thinning improves late-seral plant abundance, others have found that the legacy of pre-treatment vegetation has a stronger impact on post-treatment communities. This study focused on the impacts of ice storm disturbance and pre-treatment vegetation on the understory of mature Douglas-fir forests using the ongoing Mature Forest Study (MFS), a long-term silvicultural experiment evaluating the effects of thinning and understory vegetation management treatments, as a framework. The first study examined the impact of an ice storm (glaze disturbance) on planted understory trees. Specifically, I looked at the effect of understory tree species, tree size, and overstory neighborhood environment on the type (bending, crown loss), source (ice loading, falling debris), and severity of damage experienced by planted understory trees at one of the MFS sites. Tree species, size, and overstory environment all affected the amount of understory glaze damage. Frequency and severity of damage both varied among underplanted tree species. In general, smaller trees were more prone to being bent, while larger trees were more susceptible to crown loss. The Douglas-fir component of the overstory provided enough additional sheltering that the increased risk to understory trees from falling debris was balanced by a corresponding decrease in the odds of damage by ice loading. This was not the case for the hardwood component; increasing risk of damage to understory trees from falling debris with increasing hardwood basal area drove an overall increase in the risk of understory damage as hardwood basal area increased. This study suggests that species, tree size, and overstory environment all need to be considered by managers hoping to reduce glaze damage risk to younger cohorts in multi-aged stands. The second study investigated the impacts of thinning intensity and herbicide application on the long-term (20-year post-treatment) development of understory vegetation communities on both of the MFS sites. Trends were examined with a focus on the ability of herbicide application, in concert with thinning treatments, to reduce the legacy of common pre-treatment species and promote the abundance of late-seral associates. Results indicated that both thinning intensity and herbicide application affected 20-year changes in understory plant community composition. Herbicide application was associated with a decrease in the abundance of common pre-treatment species, suggesting that it did reduce the legacy effect. However, this was not associated with any change in the abundance of late-seral species. While light thinning showed some ability to mitigate decline in late-seral species relative to higher intensity thinnings, there was no evidence of treatment interaction with herbicide application. These results suggest that while managers may be able to reduce the influence of initial site conditions on post-treatment vegetation communities, the use of herbicides offers little control over the successional trajectory of the understory. Light thinning appears to be the most effective means of increasing late-seral species abundance, although the use of herbicides to meet other management objectives is not contraindicated by the results of this study. Overall, these results suggest that the best options available to managers to both reduce glaze disturbance impacts to understory trees and hasten the development of late-seral plant communities are heavy thinning with unmanaged leave patches to provide late-seral refugia, or light thinning with gaps to provide growing space for better tree regeneration. Title: Silvicultural Treatment Impacts on Understory Trees and 20-Year Understory Vegetation Dynamics in Mature Douglas-Fir Forests Author: Priebe, Jim E. Policy regarding the management of public forests has undergone a drastic shift over the past couple of decades due to the loss old-growth forests in the Pacific Northwest (PNW) of the United States. For much of the 20th century, forest management on public lands emphasized timber production through the use of even-aged management practices. There has been increasing recognition, however, that traditional even-aged management approaches are unable to support species that rely on the complex, heterogeneous structures provided by old-growth forests. In response, public forest managers have redirected their focus to developing more ecologically sustainable forests capable of meeting a broad array of objectives including an increasing emphasis on the development of late-seral and old-growth characteristics. Thinning has been identified as a promising method for promoting late-seral characteristics in managed stands. Recent long-term studies have shown that thinning stands does indeed accelerate the development of at least some late-seral structure characteristics, particularly when varying levels of thinning intensity and non-uniform retention patterns are incorporated into silvicultural prescriptions. Likewise, thinning has also shown some ability to increase the abundance of late-seral associated plant species in the understory. The impacts of thinning on vegetation dynamics are complicated by external factors such as natural disturbance events and the influence of pre-treatment vegetation on post-treatment communities. Within the context of managing for late-seral attributes, thinning is used to imitate natural disturbance processes. However, this does not preclude the occurrence of natural disturbances, which may either disrupt or compound treatment effects. Initial site conditions create another potential complication for the development of late-seral attributes by limiting the potential for change in understory communities. While some studies have shown that thinning improves late-seral plant abundance, others have found that the legacy of pre-treatment vegetation has a stronger impact on post-treatment communities. This study focused on the impacts of ice storm disturbance and pre-treatment vegetation on the understory of mature Douglas-fir forests using the ongoing Mature Forest Study (MFS), a long-term silvicultural experiment evaluating the effects of thinning and understory vegetation management treatments, as a framework. The first study examined the impact of an ice storm (glaze disturbance) on planted understory trees. Specifically, I looked at the effect of understory tree species, tree size, and overstory neighborhood environment on the type (bending, crown loss), source (ice loading, falling debris), and severity of damage experienced by planted understory trees at one of the MFS sites. Tree species, size, and overstory environment all affected the amount of understory glaze damage. Frequency and severity of damage both varied among underplanted tree species. In general, smaller trees were more prone to being bent, while larger trees were more susceptible to crown loss. The Douglas-fir component of the overstory provided enough additional sheltering that the increased risk to understory trees from falling debris was balanced by a corresponding decrease in the odds of damage by ice loading. This was not the case for the hardwood component; increasing risk of damage to understory trees from falling debris with increasing hardwood basal area drove an overall increase in the risk of understory damage as hardwood basal area increased. This study suggests that species, tree size, and overstory environment all need to be considered by managers hoping to reduce glaze damage risk to younger cohorts in multi-aged stands. The second study investigated the impacts of thinning intensity and herbicide application on the long-term (20-year post-treatment) development of understory vegetation communities on both of the MFS sites. Trends were examined with a focus on the ability of herbicide application, in concert with thinning treatments, to reduce the legacy of common pre-treatment species and promote the abundance of late-seral associates. Results indicated that both thinning intensity and herbicide application affected 20-year changes in understory plant community composition. Herbicide application was associated with a decrease in the abundance of common pre-treatment species, suggesting that it did reduce the legacy effect. However, this was not associated with any change in the abundance of late-seral species. While light thinning showed some ability to mitigate decline in late-seral species relative to higher intensity thinnings, there was no evidence of treatment interaction with herbicide application. These results suggest that while managers may be able to reduce the influence of initial site conditions on post-treatment vegetation communities, the use of herbicides offers little control over the successional trajectory of the understory. Light thinning appears to be the most effective means of increasing late-seral species abundance, although the use of herbicides to meet other management objectives is not contraindicated by the results of this study. Overall, these results suggest that the best options available to managers to both reduce glaze disturbance impacts to understory trees and hasten the development of late-seral plant communities are heavy thinning with unmanaged leave patches to provide late-seral refugia, or light thinning with gaps to provide growing space for better tree regeneration. Close

• 5. [Article] Establishment and growth of natural and planted conifers ten years after overstory thinning and vegetation control in 50-year-old Douglas-fir stands

  Forest managers of public lands in western Oregon and Washington have become increasingly interested in creating additional conifer cohorts in young, even-aged, second-growth Douglas-fir stands. The purpose ...

  Citation × Citation Citation Abstract Copy Title: Establishment and growth of natural and planted conifers ten years after overstory thinning and vegetation control in 50-year-old Douglas-fir stands Author: Nabel, Mark R. Forest managers of public lands in western Oregon and Washington have become increasingly interested in creating additional conifer cohorts in young, even-aged, second-growth Douglas-fir stands. The purpose of our research was to assess the establishment, survival, and growth of naturally-regenerated and underplanted conifers 10-13 years after overstory thinning and understory vegetation control in 50-year-old Douglas-fir stands. Two sites within the Oregon Coast Range were used in our study. One site was relatively dry (~130 cm of annual precipitation) and contained a mostly pure Douglas-fir overstory. The second site was moister (~175 cm of annual precipitation) and contained a Douglas-fir/western hemlock mixed-species overstory. At each site, stands were thinned to basal areas ranging from approximately 18 to 32 m2/ha using either a uniform or gappy thinning pattern. In the gappy treatment, 20% of the total area was comprised of 0.06- and 0.10-ha gaps. Understory vegetation was controlled across a portion of each thinning plot using a broadcast application of herbicides prior to thinning. After thinning, the following species were underplanted: Douglas-fir (Pseudotsuga menziesii), western hemlock (Tsuga heterophylla), western redcedar (Thuja plicata), and grand fir (Abies grandis, dry site only). Establishment of natural regeneration and survival of underplanted conifers was greater for all species under lower overstory densities at the drier site, but on the moister site, where western hemlock regenerated prolifically in the understory, no differences were detected. At both sites, underplanted and dominant naturally-regenerated conifers growing under lower overstory retention levels were generally taller and had larger diameters after ten years. Thinning pattern had no effect on the establishment, survival, or growth of understory conifers at the overall stand level, but the tallest individuals were located within gaps. Controlling competing vegetation increased the rate of establishment for naturally-regenerated Douglas-fir and the rate of survival for planted Douglas-fir and western hemlock. Vegetation control generally resulted in larger underplanted seedlings of all species after ten years, but the size of dominant naturally-regenerated conifers was not affected. Shade-tolerant species generally outperformed Douglas-fir in understories except where western redcedar suffered heavy browsing damage. Natural regeneration of western hemlock may contribute to the development of an understory conifer layer when a seed source is present in the overstory, but in mostly pure Douglas-fir stands, underplanting would likely be required to supplement sporadic and slow-growing natural regeneration. In either case, future thinnings of rapidly closing overstory canopies would be required to maintain the long-term development of an understory conifer layer. Title: Establishment and growth of natural and planted conifers ten years after overstory thinning and vegetation control in 50-year-old Douglas-fir stands Author: Nabel, Mark R. Forest managers of public lands in western Oregon and Washington have become increasingly interested in creating additional conifer cohorts in young, even-aged, second-growth Douglas-fir stands. The purpose of our research was to assess the establishment, survival, and growth of naturally-regenerated and underplanted conifers 10-13 years after overstory thinning and understory vegetation control in 50-year-old Douglas-fir stands. Two sites within the Oregon Coast Range were used in our study. One site was relatively dry (~130 cm of annual precipitation) and contained a mostly pure Douglas-fir overstory. The second site was moister (~175 cm of annual precipitation) and contained a Douglas-fir/western hemlock mixed-species overstory. At each site, stands were thinned to basal areas ranging from approximately 18 to 32 m2/ha using either a uniform or gappy thinning pattern. In the gappy treatment, 20% of the total area was comprised of 0.06- and 0.10-ha gaps. Understory vegetation was controlled across a portion of each thinning plot using a broadcast application of herbicides prior to thinning. After thinning, the following species were underplanted: Douglas-fir (Pseudotsuga menziesii), western hemlock (Tsuga heterophylla), western redcedar (Thuja plicata), and grand fir (Abies grandis, dry site only). Establishment of natural regeneration and survival of underplanted conifers was greater for all species under lower overstory densities at the drier site, but on the moister site, where western hemlock regenerated prolifically in the understory, no differences were detected. At both sites, underplanted and dominant naturally-regenerated conifers growing under lower overstory retention levels were generally taller and had larger diameters after ten years. Thinning pattern had no effect on the establishment, survival, or growth of understory conifers at the overall stand level, but the tallest individuals were located within gaps. Controlling competing vegetation increased the rate of establishment for naturally-regenerated Douglas-fir and the rate of survival for planted Douglas-fir and western hemlock. Vegetation control generally resulted in larger underplanted seedlings of all species after ten years, but the size of dominant naturally-regenerated conifers was not affected. Shade-tolerant species generally outperformed Douglas-fir in understories except where western redcedar suffered heavy browsing damage. Natural regeneration of western hemlock may contribute to the development of an understory conifer layer when a seed source is present in the overstory, but in mostly pure Douglas-fir stands, underplanting would likely be required to supplement sporadic and slow-growing natural regeneration. In either case, future thinnings of rapidly closing overstory canopies would be required to maintain the long-term development of an understory conifer layer. Close

• 6. [Article] Tree Vigor Response and Competitive Zone Density in Mature Ponderosa Pine

  This study explored the relationship of mature ponderosa pine (Pinus ponderosa Dougl. ex Laws) tree vigor and competitive zone density following thinning and fuel reduction treatments. Competitive zone ...

  Citation × Citation Citation Abstract Copy Title: Tree Vigor Response and Competitive Zone Density in Mature Ponderosa Pine Author: Shaw, Randi C. This study explored the relationship of mature ponderosa pine (Pinus ponderosa Dougl. ex Laws) tree vigor and competitive zone density following thinning and fuel reduction treatments. Competitive zone density (CZD) was defined as the total tree basal area (m²/ha) surrounding a given ponderosa pine tree within a 10m fixed radius plot. The vigor of 60 trees were observed at two separate mature ponderosa pine stands in south-central Oregon, Lookout Mountain (LM), and Sycan Marsh (SY). Tree vigor response was measured using three physiological metrics: xylem sap flow (kg/day), bole water potential (MPa), and resin exudation (ml/day). Dominant and co-dominant pines were chosen across the available range of CZD at each site. Data were collected over the course of one growing season in 2015, from late June to early October. Sap flow was recorded at four 6-day periods over the season to incorporate the influence of seasonal moisture decline on tree physiological response. Water potential was observed twice, at mid- and late-season. Resin exudation was observed in August to correspond generally with flight periods of various bark beetle species. To account for environmental influences on tree physiology, soil volumetric water content (SWC, m3/m3) was measured approximately every 3-4 weeks and daily potential evapotranspiration (PET) data were obtained from the Western Regional Climate Center’s Remote Access Weather Stations (WRCC RAWS) archives. Data were analyzed using a linear mixed model including both environmental and competitive variables. A quadratic relationship between post-thinning CZD and sap flow was evident, indicating a positive but highly variable association up to a threshold of roughly 40 m²/ha. Above this density threshold sap flow declined markedly, suggesting that tree competition becomes an important factor in constraining water resource access in these sites at higher densities. Bole water potential in mature ponderosa pine was also related to CZD at the LM site. Resin exudation showed disparate relationships to CZD by site and may be more directly influenced by factors other than CZD. Overall tree vigor was higher at LM despite comparatively lower SWC, potentially due to the greater size or age of the study trees. The positive correlation of vigor and CZD up to the middle density range implies that there may be facilitative effects of adjacent trees to be considered for mature ponderosa pine, especially when reducing drought vulnerability is a management goal. Title: Tree Vigor Response and Competitive Zone Density in Mature Ponderosa Pine Author: Shaw, Randi C. This study explored the relationship of mature ponderosa pine (Pinus ponderosa Dougl. ex Laws) tree vigor and competitive zone density following thinning and fuel reduction treatments. Competitive zone density (CZD) was defined as the total tree basal area (m²/ha) surrounding a given ponderosa pine tree within a 10m fixed radius plot. The vigor of 60 trees were observed at two separate mature ponderosa pine stands in south-central Oregon, Lookout Mountain (LM), and Sycan Marsh (SY). Tree vigor response was measured using three physiological metrics: xylem sap flow (kg/day), bole water potential (MPa), and resin exudation (ml/day). Dominant and co-dominant pines were chosen across the available range of CZD at each site. Data were collected over the course of one growing season in 2015, from late June to early October. Sap flow was recorded at four 6-day periods over the season to incorporate the influence of seasonal moisture decline on tree physiological response. Water potential was observed twice, at mid- and late-season. Resin exudation was observed in August to correspond generally with flight periods of various bark beetle species. To account for environmental influences on tree physiology, soil volumetric water content (SWC, m3/m3) was measured approximately every 3-4 weeks and daily potential evapotranspiration (PET) data were obtained from the Western Regional Climate Center’s Remote Access Weather Stations (WRCC RAWS) archives. Data were analyzed using a linear mixed model including both environmental and competitive variables. A quadratic relationship between post-thinning CZD and sap flow was evident, indicating a positive but highly variable association up to a threshold of roughly 40 m²/ha. Above this density threshold sap flow declined markedly, suggesting that tree competition becomes an important factor in constraining water resource access in these sites at higher densities. Bole water potential in mature ponderosa pine was also related to CZD at the LM site. Resin exudation showed disparate relationships to CZD by site and may be more directly influenced by factors other than CZD. Overall tree vigor was higher at LM despite comparatively lower SWC, potentially due to the greater size or age of the study trees. The positive correlation of vigor and CZD up to the middle density range implies that there may be facilitative effects of adjacent trees to be considered for mature ponderosa pine, especially when reducing drought vulnerability is a management goal. Close